Appendix 1 in Integrative taxonomy reveals four new species in the superdiverse genus Macrobiotus (Tardigrada: Eutardigrada: Macrobiotidae)
Authors: Stec, Daniel
Type: Dataset
Published: 2026-05-18
Keywords: Biodiversity, Taxonomy

<p class="zenodo-description-adequate-caption"><b>Appendix 1</b> (continued on next five pages). Sequences used for phylogenetic analyses. Sequences obtained in this study are indicated in bold, and asterisks denotes <i>Macrobiotus</i> C.A.S. Schultze, 1834 sequences that was misidentified by previous authors, as noted by (Stec 2024). Shaded rows at the end of the table indicate outgroup taxa.</p><table><tbody><tr><th><b>Species</b></th><th><b>18S</b></th><th><b>28S</b></th><th><b>ITS-2</b></th><th><b>COI</b></th><th><b>Source</b></th></tr></tbody><tbody><tr><th><b><i>Macrobiotus witalinskii</i> MG. 001.01</b></th><td><b>PX789912</b></td><td><b>PX789920</b></td><td><b>PX789928</b></td><td><b>PX794742</b></td><td><b>This study</b></td></tr><tr><th><b><i>Macrobiotus witalinskii</i> MG. 001.02</b></th><td><b>PX789913</b></td><td><b>PX789921</b></td><td><b>PX789929</b></td><td><b>PX794743</b></td><td><b>This study</b></td></tr><tr><th><b><i>Macrobiotus hoianicus</i> VN. 004.01</b></th><td><b>PX789914</b></td><td><b>PX789922</b></td><td><b>PX789930</b></td><td><b>PX794744</b></td><td><b>This study</b></td></tr><tr><th><b><i>Macrobiotus hoianicus</i> VN. 004.02</b></th><td><b>PX789915</b></td><td><b>PX789923</b></td><td><b>PX789931</b></td><td><b>PX794745</b></td><td><b>This study</b></td></tr><tr><th><b><i>Macrobiotus surmaczi</i> VN. 003.01</b></th><td><b>PX789916</b></td><td><b>PX789925</b></td><td><b>PX789932</b></td><td><b>PX794746</b></td><td><b>This study</b></td></tr><tr><th><b><i>Macrobiotus surmaczi</i> VN. 003.02</b></th><td><b>PX789917</b></td><td><b>PX789924</b></td><td><b>PX789933</b></td><td><b>PX794747</b></td><td><b>This study</b></td></tr><tr><th><b><i>Macrobiotus dalaticus</i> VN. 002.01</b></th><td><b>PX789918</b></td><td><b>PX789927</b></td><td><b>PX789934</b></td><td><b>PX794748</b></td><td><b>This study</b></td></tr><tr><th><b><i>Macrobiotus dalaticus</i> VN. 002.02</b></th><td><b>PX789919</b></td><td><b>PX789926</b></td><td><b>PX789935</b></td><td><b>PX794749</b></td><td><b>This study</b></td></tr><tr><th><i>Macrobiotus vattenrikense</i></th><td>PX093663</td><td></td><td>PX093644</td><td>PX093655</td><td>Hulterström <i>et al.</i> 2025</td></tr><tr><th><i>Macrobiotus anshui</i></th><td>OQ525805</td><td>OQ526003</td><td></td><td>OQ525803</td><td>Kayastha <i>et al.</i> 2025</td></tr><tr><th><i>Macrobiotus sharapovi</i></th><td>PV283176</td><td>PV283177</td><td>PV283179</td><td>PV282532</td><td>Polishchuk <i>et al.</i> 2025</td></tr><tr><th><i>Macrobiotus olgae</i> 1</th><td></td><td></td><td>PQ461834</td><td>PQ276069</td><td>Rocha <i>et al.</i> 2025</td></tr><tr><th><i>Macrobiotus olgae</i> 2</th><td></td><td></td><td>PQ461835</td><td>PQ276070</td><td>Rocha <i>et al.</i> 2025</td></tr><tr><th><i>Macrobiotus kathyae</i> 1</th><td>PP391285</td><td>PP391287</td><td>PP391289</td><td>PP386934</td><td>Massa & Vecchi 2024</td></tr><tr><th><i>Macrobiotus kathyae</i> 2</th><td>PP391286</td><td>PP391288</td><td>PP391290</td><td>PP386935</td><td>Massa & Vecchi 2024</td></tr><tr><th><i>Macrobiotus ovovittatus</i> GL. 001.01</th><td>OR543310</td><td>OR543318</td><td>OR543314</td><td>OR544395</td><td>Stec 2024</td></tr><tr><th><i>Macrobiotus ovovittatus</i> GL. 001.02</th><td>OR543311</td><td>OR543319</td><td>OR543315</td><td>OR544396</td><td>Stec 2024</td></tr><tr><th><i>Macrobiotus mileri</i> IL. 001.01</th><td>OR543312</td><td>OR543320</td><td>OR543316</td><td>OR544397</td><td>Stec 2024</td></tr><tr><th><i>Macrobiotus mileri</i> IL. 001.02</th><td>OR543313</td><td>OR543321</td><td>OR543317</td><td>OR544398</td><td>Stec 2024</td></tr><tr><th><i>Macrobiotus mileri</i> IL. 001.03</th><td></td><td></td><td></td><td>OR544399</td><td>Stec 2024</td></tr><tr><th><i>Macrobiotus mileri</i> IL. 001.04</th><td></td><td></td><td></td><td>OR544400</td><td>Stec 2024</td></tr><tr><th><i>Macrobiotus kosmali</i> M8.1</th><td>OP142472</td><td>OP143765</td><td>OP153786</td><td>OP141639</td><td>Kayastha <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus kosmali</i> M8.2</th><td>OP142473</td><td>OP143766</td><td></td><td>OP141640</td><td>Kayastha <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus hupingensis</i> *</th><td>MW183923</td><td>MZ470349</td><td>MZ474842</td><td>MW186952</td><td>Yuan <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus dolosus</i> C3209_2</th><td>OP596290</td><td></td><td></td><td>OP561772</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus dolosus</i> C3209_US2</th><td></td><td></td><td></td><td>OP561773</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus dolosus</i> C3209_4</th><td></td><td></td><td></td><td>OP561774</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus dolosus</i> C3581_V6</th><td>OP596292</td><td></td><td></td><td>OP561775</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus dolosus</i> C3209_1</th><td>OP596289</td><td></td><td></td><td></td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus dolosus</i> C3581_V5</th><td>OP596291</td><td></td><td></td><td></td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus sidereus</i> C2796_2</th><td>OP596293</td><td></td><td></td><td>OP561776</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus sidereus</i> C3282_UFK</th><td></td><td></td><td></td><td>OP561777</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus sidereus</i> C3282_ UFL</th><td></td><td></td><td></td><td>OP561778</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus sidereus</i> C3282_ UFM</th><td></td><td></td><td></td><td>OP561779</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus sidereus</i> C3282_ UFN</th><td>OP596294</td><td></td><td></td><td>OP561780</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus sidereus</i> C3282_ UFO</th><td></td><td></td><td></td><td>OP561781</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus sidereus</i> C3282_ UFP</th><td></td><td></td><td></td><td>OP561782</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus fonturai</i> C2861_5</th><td>OP596295</td><td></td><td></td><td>OP561783</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus fonturai</i> C2861_ US1</th><td>OP596296</td><td></td><td></td><td>OP561784</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus muralis</i> C2861_FG</th><td>OP596297</td><td></td><td></td><td></td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus muralis</i> C2861_6</th><td>OP596298</td><td></td><td></td><td></td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>muralis</i> C3251_2</th><td>OP596299</td><td></td><td></td><td>OP561785</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>muralis</i> C3251_3</th><td>OP596300</td><td></td><td></td><td>OP561786</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>muralis</i> C3251_FA</th><td>OP596301</td><td></td><td></td><td>OP561787</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>muralis</i> C3251_4</th><td></td><td></td><td></td><td>OP561788</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>muralis</i> C3251_5</th><td></td><td></td><td></td><td>OP561789</td><td>Bertolani <i>et al.</i> 2023</td></tr><tr><th><i>Macrobiotus hufelandi</i> C2953_ A02</th><td>OP596302</td><td></td><td></td><td>HQ876586</td><td>Bertolani <i>et al.</i> 2011, 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>hufelandi</i> C2959_A01</th><td>OP596303</td><td></td><td></td><td>HQ876590</td><td>Bertolani <i>et al.</i> 2011, 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>hufelandi</i> C2959_A02</th><td>OP596304</td><td></td><td></td><td>HQ876591</td><td>Bertolani <i>et al.</i> 2011, 2023</td></tr><tr><th><i>Macrobiotus vladimiri</i> C2688_ A02</th><td>OP596305</td><td></td><td></td><td>HM136932</td><td>Bertolani <i>et al.</i> 2011, 2023</td></tr><tr><th><i>Macrobiotus terminalis</i> C2868_ N02</th><td>OP596308</td><td></td><td></td><td>JN673959</td><td>Bertolani <i>et al.</i> 2011, 2023</td></tr><tr><th><i>Macrobiotus</i> cf. <i>nelsonae</i> 1</th><td>HQ604965</td><td></td><td></td><td></td><td>Bertolani <i>et al.</i> 2014</td></tr><tr><th><i>Macrobiotus</i> cf. <i>nelsonae</i> 2</th><td>HQ604966</td><td></td><td></td><td></td><td>Bertolani <i>et al.</i> 2014</td></tr><tr><th><i>Macrobiotus rebecchii</i> 1</th><td>OP479887</td><td></td><td></td><td>OP477442</td><td>Stec 2022a</td></tr><tr><th><i>Macrobiotus rebecchii</i> 2</th><td>OP479888</td><td></td><td></td><td>OP477443</td><td>Stec 2022a</td></tr><tr><th><i>Macrobiotus kyoukenus</i> v3</th><td>ON818312</td><td></td><td>ON818300</td><td>ON809461</td><td>Cesari <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus kyoukenus</i> us3</th><td>ON818314</td><td></td><td>ON818301</td><td>ON809462</td><td>Cesari <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus kyoukenus</i> us4</th><td>ON818315</td><td></td><td>ON818302</td><td>ON809463</td><td>Cesari <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus kyoukenus</i> us5</th><td>ON818316</td><td></td><td>ON818303</td><td>ON809464</td><td>Cesari <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus scoticus</i> DK. 056.01</th><td>OK663218</td><td>OK663207</td><td>OK662989</td><td>OK663228</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus scoticus</i> DK. 056.02</th><td>OK663217</td><td>OK663206</td><td>OK662988</td><td>OK663229</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus naginae</i> S226_1</th><td>OK663219</td><td>OK663230</td><td>OK663209</td><td>OK662990</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus naginae</i> S226_2</th><td>OK663220</td><td>OK663231</td><td>OK663208</td><td>OK662991</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus naginae</i> S605_1</th><td>OK663221</td><td>OK663232</td><td>OK663210</td><td>OK662992</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus naginae</i> S605_2</th><td>OK663222</td><td>OK663233</td><td>OK663211</td><td>OK662993</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus sandrae</i> S859_1</th><td>OK663223</td><td>OK663234</td><td>OK663212</td><td>OK662994</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus</i> cf. <i>sapiens</i></th><td>OK663226</td><td>OK663237</td><td>OK663215</td><td>OK662997</td><td>Vecchi <i>et al.</i> 2022a</td></tr><tr><th><i>Macrobiotus sandrae</i> 1</th><td>MW695445</td><td></td><td></td><td>HQ876573</td><td>Bertolani <i>et al.</i> 2011; Marnissi <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus sandrae</i> 2</th><td>MW695446</td><td></td><td></td><td>HQ876577</td><td>Bertolani <i>et al.</i> 2011; Marnissi <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus azzunae</i> 1</th><td>MW695447</td><td></td><td>MW695454</td><td>MW698697</td><td>Marnissi <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus azzunae</i> 2</th><td>MW695448</td><td></td><td>MW695455</td><td>MW698698</td><td>Marnissi <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus birendrai</i></th><td>MW680641</td><td>MW680644</td><td>MW680418</td><td>MW656266</td><td>Kayastha <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus</i> a. <i>groenlandicus</i> 1</th><td>MZ463664</td><td>MZ463679</td><td>MZ463654</td><td>MZ461006</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus</i> a. <i>groenlandicus</i> 2</th><td>MZ463663</td><td>MZ463678</td><td>MZ463655</td><td>MZ461007</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus</i> a. <i>groenlandicus</i> 3</th><td>MZ463662</td><td>MZ463677</td><td>MZ463653</td><td>MZ461005</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus kirghizicus</i> 1</th><td>MZ463666</td><td>MZ463672</td><td>MZ463659</td><td>MZ461002</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus kirghizicus</i> 2</th><td>MZ463665</td><td>MZ463671</td><td>MZ463660</td><td>MZ461003</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus kirghizicus</i> 3</th><td>MZ463667</td><td>MZ463673</td><td>MZ463661</td><td>MZ461004</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus</i> a. <i>ariekammensis</i> 1</th><td>MZ463668</td><td>MZ463674</td><td>MZ463656</td><td>MZ460999</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus</i> a. <i>ariekammensis</i> 2</th><td>MZ463669</td><td>MZ463675</td><td>MZ463657</td><td>MZ461000</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus</i> a. <i>ariekammensis</i> 3</th><td>MZ463670</td><td>MZ463676</td><td>MZ463658</td><td>MZ461001</td><td>Stec <i>et al.</i> 2022</td></tr><tr><th><i>Macrobiotus</i> aff. <i>pseudohufelandi</i> PL</th><td>MN888373</td><td>MN888358</td><td>MN888345</td><td>MN888325</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus</i> aff. <i>pseudohufelandi</i> ZA</th><td>MN888374</td><td>MN888359</td><td>MN888345</td><td>MN888326</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus annewintersae</i> 1</th><td>MW588024</td><td>MW588030</td><td>MW588019</td><td>MW593927</td><td>Vecchi & Stec 2021</td></tr><tr><th><i>Macrobiotus annewintersae</i> 2</th><td>MW588025</td><td>MW588031</td><td>MW588018</td><td>MW593928</td><td>Vecchi & Stec 2021</td></tr><tr><th><i>Macrobiotus basiatus</i></th><td>MT498094</td><td>MT488397</td><td>MT505165</td><td>MT502116</td><td>Nelson <i>et al.</i> 2020</td></tr><tr><th><i>Macrobiotus caelestis</i></th><td>MK737073</td><td>MK737071</td><td>MK737072</td><td>MK737922</td><td>Coughlan <i>et al.</i> 2019</td></tr><tr><th><i>Macrobiotus canaricus</i> 1</th><td>MH063925</td><td>MH063934</td><td>MH063928</td><td>MH057765</td><td>Stec <i>et al.</i> 2018d</td></tr><tr><th><i>Macrobiotus canaricus</i> 2</th><td></td><td></td><td>MH063929</td><td>MH057766</td><td>Stec <i>et al.</i> 2018d</td></tr><tr><th><i>Macrobiotus</i> cf. <i>polonicus</i> 1</th><td>MW588026</td><td>MW588032</td><td>MW588021</td><td>MW593929</td><td>Vecchi & Stec 2021</td></tr><tr><th><i>Macrobiotus</i> cf. <i>polonicus</i> 2</th><td>MW588027</td><td>MW588033</td><td>MW588020</td><td>MW593930</td><td>Vecchi & Stec 2021</td></tr><tr><th><i>Macrobiotus</i> cf. <i>recens</i> 1</th><td>MH063927</td><td>MH063936</td><td>MH063932</td><td>MH057768</td><td>Stec <i>et al.</i> 2018d</td></tr><tr><th><i>Macrobiotus</i> cf. <i>recens</i> 2</th><td></td><td></td><td>MH063933</td><td>MH057769</td><td>Stec <i>et al.</i> 2018d</td></tr><tr><th><i>Macrobiotus crustulus</i></th><td>MT261912</td><td>MT261903</td><td>MT261907</td><td>MT260371</td><td>Stec <i>et al.</i> 2020d</td></tr><tr><th><i>Macrobiotus engbergi</i> 1</th><td>MN443039</td><td>MN443034</td><td>MN443036</td><td>MN444824</td><td>Stec <i>et al.</i> 2020c</td></tr><tr><th><i>Macrobiotus engbergi</i> 2</th><td></td><td></td><td>MN443037</td><td>MN444825</td><td>Stec <i>et al.</i> 2020c</td></tr><tr><th><i>Macrobiotus engbergi</i> 3</th><td></td><td></td><td></td><td>MN444826</td><td>Stec <i>et al.</i> 2020c</td></tr><tr><th><i>Macrobiotus glebkai</i></th><td>MW247177</td><td>MW247176</td><td>MW247180</td><td>MW246134</td><td>Kiosya <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus hannae</i></th><td>MH063922</td><td>MH063924</td><td>MH063923</td><td>MH057764</td><td>Nowak & Stec 2018</td></tr><tr><th><i>Macrobiotus kamilae</i> 1</th><td>MK737070</td><td>MK737064</td><td>MK737067</td><td>MK737920</td><td>Coughlan & Stec 2019</td></tr><tr><th><i>Macrobiotus kamilae</i> 2</th><td></td><td></td><td></td><td>MK737921</td><td>Coughlan & Stec 2019</td></tr><tr><th><i>Macrobiotus kristenseni</i></th><td>KC193577</td><td></td><td></td><td>KC193573</td><td>Guidetti <i>et al.</i> 2013</td></tr><tr><th><i>Macrobiotus macrocalix</i></th><td>MH063926</td><td>MH063935</td><td>MH063931</td><td>MH057767</td><td>Stec <i>et al.</i> 2018d</td></tr><tr><th><i>Macrobiotus noongaris</i> 1</th><td>MK737069</td><td>MK737063</td><td>MK737065</td><td>MK737919</td><td>Coughlan & Stec 2019</td></tr><tr><th><i>Macrobiotus noongaris</i> 2</th><td></td><td></td><td>MK737066</td><td></td><td>Coughlan & Stec 2019</td></tr><tr><th><i>Macrobiotus papei</i></th><td>MH063881</td><td>MH063880</td><td>MH063921</td><td>MH057763</td><td>Stec <i>et al.</i> 2018a</td></tr><tr><th><i>Macrobiotus paulinae</i></th><td>KT935502</td><td>KT935501</td><td>KT935500</td><td>KT951668</td><td>Stec <i>et al.</i> 2015</td></tr><tr><th><i>Macrobiotus polonicus</i> AT 1</th><td>MN888369</td><td>MN888355</td><td>MN888337</td><td>MN888317</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus polonicus</i> AT 2</th><td></td><td></td><td>MN888338</td><td>MN888318</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus polonicus</i> AT 3</th><td></td><td></td><td></td><td>MN888319</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus polonicus</i> SK 1</th><td>MN888370</td><td>MN888356</td><td>MN888332</td><td>MN888320</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus polonicus</i> SK 2</th><td></td><td></td><td>MN888333</td><td>MN888321</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus polonicus</i> SK 3</th><td></td><td></td><td>MN888334</td><td></td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus porifini</i> 1</th><td>MT241900</td><td>MT241897</td><td></td><td>MT246659</td><td>Kuzdrowska <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus porifini</i> 2</th><td></td><td>MT241898</td><td></td><td>MT246660</td><td>Kuzdrowska <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus porifini</i> 3</th><td></td><td>MT241899</td><td></td><td>MT246661</td><td>Kuzdrowska <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus polypiformis</i> 1</th><td>KX810008</td><td>KX810009</td><td>KX810010</td><td>KX810011</td><td>Roszkowska <i>et al.</i> 2017</td></tr><tr><th><i>Macrobiotus polypiformis</i> 2</th><td></td><td></td><td></td><td>KX810012</td><td>Roszkowska <i>et al.</i> 2017</td></tr><tr><th><i>Macrobiotus pseudohufelandi</i> 1</th><td>HQ604989</td><td></td><td></td><td>AY598776</td><td>Bertolani <i>et al.</i> 2014</td></tr><tr><th><i>Macrobiotus pseudohufelandi</i> 2</th><td>HQ604990</td><td></td><td></td><td>AY598777</td><td>Bertolani <i>et al.</i> 2014</td></tr><tr><th><i>Macrobiotus rybaki</i> 1</th><td>MW588029</td><td>MW588034</td><td>MW588022</td><td>MW593931</td><td>Vecchi & Stec 2021</td></tr><tr><th><i>Macrobiotus rybaki</i> 2</th><td>MW588028</td><td>MW588035</td><td>MW588023</td><td>MW593932</td><td>Vecchi & Stec 2021</td></tr><tr><th><i>Macrobiotus scoticus</i></th><td>KY797265</td><td>KY797266</td><td>KY797268</td><td>KY797267</td><td>Stec <i>et al.</i> 2017b</td></tr><tr><th><i>Macrobiotus shonaicus</i> 1</th><td>MG757132</td><td>MG757133</td><td>MG757134</td><td>MG757136</td><td>Stec <i>et al.</i> 2018b</td></tr><tr><th><i>Macrobiotus shonaicus</i> 2</th><td></td><td></td><td>MG757135</td><td>MG757137</td><td>Stec <i>et al.</i> 2018b</td></tr><tr><th><i>Macrobiotus sottilei</i></th><td>MW247178</td><td>MW247175</td><td>MW247179</td><td>MW246133</td><td>Kiosya <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus vladimiri</i> FI</th><td>MN888375</td><td>MN888360</td><td>MN888347</td><td>MN888327</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Macrobiotus wandae</i></th><td>MN435112</td><td>MN435116</td><td>MN435120</td><td>MN482684</td><td>Kayastha <i>et al.</i> 2020</td></tr><tr><th><i>Macrobiotus pallarii</i> 1</th><td>MT809069</td><td>MT809081</td><td>MT809094</td><td>MT807924</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus pallarii</i> 2</th><td>MT809070</td><td>MT809082</td><td>MT809095</td><td>MT807925</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus pallarii</i> 3</th><td>MT809071</td><td>MT809083</td><td>MT809096</td><td>MT807926</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus pseudopallarii</i> 1</th><td>MT809067</td><td>MT809079</td><td>MT809091</td><td>MT807921</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus pseudopallarii</i> 2</th><td>MT809068</td><td>MT809080</td><td>MT809092</td><td>MT807922</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus ripperi</i> FI</th><td>MT809076</td><td>MT809089</td><td>MT809103</td><td>MT807930</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus ripperi</i> PL</th><td>MT809074</td><td>MT809086</td><td>MT809100</td><td>MT807933</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus margoae</i> US</th><td>MT809072</td><td>MT809084</td><td>MT809098</td><td>MT807927</td><td>Stec <i>et al.</i> 2021b</td></tr><tr><th><i>Macrobiotus gretae</i> 1</th><td>MW588434</td><td></td><td>MW588431</td><td>MW581665</td><td>Massa <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus gretae</i> 2</th><td>MW588437</td><td></td><td>MW588433</td><td>MW581668</td><td>Massa <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus gretae</i> 3</th><td>MW588435</td><td></td><td>MW588432</td><td>MW581666</td><td>Massa <i>et al.</i> 2021</td></tr><tr><th><i>Macrobiotus gretae</i> 4</th><td>MW588436</td><td></td><td></td><td>MW581667</td><td>Massa <i>et al.</i> 2021</td></tr><tr><th><i>Sisubiotus hakaiensis</i> 1</th><td>OM523054</td><td>OM523059</td><td>OM523057</td><td>OM523181</td><td>Vecchi <i>et al.</i> 2022b</td></tr><tr><th><i>Sisubiotus hakaiensis</i> 2</th><td>OM523055</td><td>OM523060</td><td>OM523058</td><td>OM523182</td><td>Vecchi <i>et al.</i> 2022b</td></tr><tr><th><i>Sisubiotus spectabilis</i> FI 1</th><td>MN888371</td><td>MN888357</td><td>MN888331</td><td>MN888322</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Sisubiotus spectabilis</i> FI 2</th><td></td><td></td><td></td><td>MN888323</td><td>Stec <i>et al.</i> 2021a</td></tr><tr><th><i>Sisubiotus spectabilis</i> NO</th><td>MN888372</td><td>MN888364</td><td>MN888344</td><td>MN888324</td><td>Ste <i>et al.</i> 2021a</td></tr><tr><th><i>Sisubiotus splendidus</i></th><td>PV744336</td><td>PV744341</td><td>PV746802</td><td></td><td>Atherton 2025</td></tr><tr><th><i>Mesobiotus efa</i> 1</th><td>OR804457</td><td>OR805135</td><td>OR805169</td><td>OR803035</td><td>Tumanov <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus efa</i> 2</th><td>OR804458</td><td>OR805136</td><td>OR805170</td><td>OR803036</td><td>Tumanov <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus efa</i> 3</th><td>OR804459</td><td>OR805137</td><td>OR805171</td><td>OR803037</td><td>Tumanov <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus vulpinus</i> 1</th><td>OR804461</td><td>OR805140</td><td>OR805172</td><td>OR803040</td><td>Tumanov <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus vulpinus</i> 2</th><td>OR804462</td><td>OR805141</td><td>OR805173</td><td>OR803041</td><td>Tumanov <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus occultatus</i> DT 97</th><td>OR794157</td><td>OR794158</td><td>OR803042</td><td>OR805249</td><td>Tumanov <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus aradasi</i></th><td>OQ933000</td><td>OQ932999</td><td>OQ940183</td><td>OQ928739</td><td>Vecchi <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus huecoensis</i> S2027.1</th><td>OQ756249</td><td></td><td></td><td>OQ756246</td><td>Vecchi <i>et al.</i> 2023</td></tr><tr><th><i>Mesobiotus huecoensis</i> S2027.4</th><td>OQ756248</td><td></td><td></td><td>OQ756247</td><td>Vecchi <i>et al.</i> 2023</td></tr><tr><th><i>Mesobiotus mandalori</i> 1</th><td>OP829143</td><td>OP829056</td><td>OP829058</td><td>OP825090</td><td>Erdmann <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus mandalori</i> 2</th><td>OP829144</td><td>OP829057</td><td></td><td>OP825091</td><td>Erdmann <i>et al.</i> 2024</td></tr><tr><th><i>Mesobiotus anastasiae</i></th><td>MT903468</td><td>MT903612</td><td>MT903470</td><td>MT904513</td><td>Tumanov 2020</td></tr><tr><th><i>Mesobiotus hilariae</i></th><td>KT226070</td><td></td><td></td><td>KT226108</td><td>Vecchi <i>et al.</i> 2016</td></tr><tr><th><i>Mesobiotus</i> cf. <i>barabanovi</i></th><td>MN310392</td><td>MN310388</td><td>MN310390</td><td>MN313170</td><td>Kaczmarek <i>et al.</i> 2020</td></tr><tr><th><i>Mesobiotus datanlanicus</i></th><td>MK584659</td><td>MK584658</td><td>MK584657</td><td>MK578905</td><td>Stec 2019</td></tr><tr><th><i>Mesobiotus dilimanensis</i></th><td>MN257048</td><td>MN257049</td><td>MN257050</td><td>MN257047</td><td>Itang <i>et al.</i> 2020</td></tr><tr><th><i>Mesobiotus ethiopicus</i></th><td>MF678793</td><td>MF678792</td><td>MN122776</td><td>MF678794</td><td>Stec & Kristensen 2017</td></tr><tr><th><i>Mesobiotus fiedleri</i></th><td>MH681585</td><td>MH681693</td><td>MH681724</td><td>MH676056</td><td>Kaczmarek <i>et al.</i> 2020</td></tr><tr><th><i>Mesobiotus harmsworthi</i></th><td>MH197146</td><td>MH197264</td><td>MH197154</td><td>MH195150</td><td>Kaczmarek <i>et al.</i> 2018b</td></tr><tr><th><i>Mesobiotus insanis</i></th><td>MF441488</td><td>MF441489</td><td>MF441490</td><td>MF441491</td><td>Mapalo <i>et al.</i> 2017</td></tr><tr><th><i>Mesobiotus occultatus</i></th><td>MH197147</td><td></td><td>MH197155</td><td>MH195152</td><td>Kaczmarek <i>et al.</i> 2018b</td></tr><tr><th><i>Mesobiotus philippinicus</i></th><td>KX129793</td><td>KX129794</td><td>KX129795</td><td>KX129796</td><td>Mapalo <i>et al.</i> 2016</td></tr><tr><th><i>Mesobiotus radiatus</i> 1</th><td>MH197153</td><td>MH197152</td><td>MH197267</td><td>MH195147</td><td>Stec <i>et al.</i> 2018e</td></tr><tr><th><i>Mesobiotus radiatus</i> 2</th><td></td><td></td><td>MH197268</td><td>MH195148</td><td>Stec <i>et al.</i> 2018e</td></tr><tr><th><i>Mesobiotus romani</i></th><td>MH197158</td><td>MH197151</td><td>MH197150</td><td>MH195149</td><td>Roszkowska <i>et al.</i> 2018</td></tr><tr><th><i>Mesobiotus imperialis</i> 1</th><td>OL257854</td><td>OL257866</td><td></td><td>OL311514</td><td>Stec 2021</td></tr><tr><th><i>Mesobiotus imperialis</i> 2</th><td>OL257855</td><td>OL257867</td><td></td><td>OL311515</td><td>Stec 2021</td></tr><tr><th><i>Mesobiotus marmoreus</i> 1</th><td>OL257856</td><td>OL257868</td><td>OL257861</td><td>OL311516</td><td>Stec 2021</td></tr><tr><th><i>Mesobiotus marmoreus</i> 2</th><td>OL257857</td><td>OL257869</td><td>OL257862</td><td>OL311517</td><td>Stec 2021</td></tr><tr><th><i>Mesobiotus marmoreus</i> 3</th><td>OL257858</td><td>OL257870</td><td>OL257863</td><td>OL311518</td><td>Stec 2021</td></tr><tr><th><i>Mesobiotus skorackii</i></th><td></td><td>MW680636</td><td></td><td>MW656257</td><td>Kayastha <i>et al.</i> 2021</td></tr><tr><th><i>Mesobiotus</i> sp. Macro07_042</th><td>MW751942</td><td></td><td></td><td>MW727957</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> Macro 06_296</th><td>MW751936</td><td></td><td></td><td>MW727958</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> Macro 06_310</th><td>MW751937</td><td></td><td></td><td>MW727961</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> Macro 06_313</th><td>MW751939</td><td></td><td></td><td>MW727960</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> CC _MF_4</th><td>MW751949</td><td></td><td></td><td>MW727933</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> ABDC _MF_3</th><td>MW751944</td><td></td><td></td><td>MW727932</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> KPRI _MF_1</th><td>MW751962</td><td></td><td></td><td>MW727934</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> HMI _MF_1</th><td>MW751957</td><td></td><td></td><td>MW727941</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> EBNI _MF_2</th><td>MW751952</td><td></td><td></td><td>MW727937</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> EBNI _MF_4</th><td>MW751954</td><td></td><td></td><td>MW727938</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> PSAI _MF_2</th><td>MW751967</td><td></td><td></td><td>MW727939</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> Macro 06_162</th><td>MW751934</td><td></td><td></td><td>MW727955</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> Macro 06_171</th><td>MW751935</td><td></td><td></td><td>MW727956</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> JN 07_MF_1</th><td>MW751959</td><td></td><td></td><td>MW727951</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> JN 07_MF_4</th><td>MW751960</td><td></td><td></td><td>MW727953</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> JN 07_MF_8</th><td>MW751961</td><td></td><td></td><td>MW727947</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus</i> cf. <i>furciger</i> FN 01_MF_6</th><td>MW751955</td><td></td><td></td><td>MW727945</td><td>Short <i>et al.</i> 2022</td></tr><tr><th><i>Mesobiotus diegoi</i> 1</th><td>OP142527</td><td>OP142520</td><td>OP142514</td><td>OP143858</td><td>Stec 2022b</td></tr><tr><th><i>Mesobiotus diegoi</i> 2</th><td>OP142526</td><td>OP142521</td><td>OP142515</td><td>OP143857</td><td>Stec 2022b</td></tr><tr><th><i>Mesobiotus maklowiczi</i> 1</th><td>OP142525</td><td>OP142518</td><td></td><td>OP143855</td><td>Stec 2022b</td></tr><tr><th><i>Mesobiotus maklowiczi</i> 2</th><td>OP142524</td><td>OP142519</td><td></td><td>OP143856</td><td>Stec 2022b</td></tr><tr><th><i>Mesobiotus peterseni</i> 1</th><td>OP142528</td><td>OP142522</td><td>OP142516</td><td>OP143859</td><td>Stec 2022b</td></tr><tr><th><i>Mesobiotus peterseni</i> 2</th><td>OP142529</td><td>OP142523</td><td>OP142517</td><td>OP143860</td><td>Stec 2022b</td></tr><tr><th><i>Minibiotus ioculator</i></th><td>MT023998</td><td>MT024041</td><td>MT024000</td><td>MT023412</td><td>Stec <i>et al.</i> 2020b</td></tr><tr><th><i>Paramacrobiotus areolatus</i></th><td>MH664931</td><td>MH664948</td><td>MH666080</td><td>MH675998</td><td>Stec <i>et al.</i> 2020b</td></tr><tr><th><i>Tenuibiotus zandrae</i></th><td>MN443040</td><td>MN443035</td><td>MN443038</td><td>MN444827</td><td>Stec <i>et al.</i> 2020c</td></tr></tbody></table>