reapxdev/zenodo-scraper huggingface dataset:
Protoma harrisoni Kiel & Hadi & Ataabadi & Xi 2026, comb. nov.
Authors: Kiel, Steffen, Hadi, Mehdi, Ataabadi, Majid Mirzaie, Xi, Dangpeng
Type: Taxonomic treatment
Published: 2026-07-07
Keywords: Biodiversity, Taxonomy, Animalia, Mollusca, Gastropoda, Turritellidae, Protoma, Protoma harrisoni

<p><b><i>Protoma harrisoni</i> (Cox, 1936) comb. nov.</b></p><p>Fig. 2A–D</p><p><i>Turritella harrisoni</i> Cox, 1936: 39, pl. 8 figs 25–26, 28.</p><p><i>Turritella</i> sp. – Hadi <i>et al</i>. 2024: 4, fig. 4.1a–b.</p><p>Non <i>Turritella harrisoni</i> Cox, 1936: pl. 8 figs 27, 29.</p><p>Non <i>Turritella angulata</i> Sowerby, 1840 – Crame 1984: pl. 1 fig. 1.</p><p><b>Material examined</b></p><p>IRAN – <b>Sistan and Baluchestan</b> • 10 shells; Dar Pahn Unit;, GDZU-0001 to GDZU-0010.</p><p><b>Remarks</b></p><p>Cox (1936) introduced <i>Turritella harrisoni</i> for specimens from the Pliocene ‘Mekran beds’ near Bandar-e Jask on the western end of the Makran margin in southern Iran. He described the whorls as “convex”, and among the five primary spirals, he noted “the three more posterior ones [are] weaker than the other two”. This description matches only some of the specimens he illustrated, including the one he selected as holotype (Cox 1936: pl. 8 fig. 26). Two specimens (Cox 1936: pl. 8 figs 27, 29), however, show a nearly flat whorl profile and spirals of almost equal strength. These specimens are now assigned to <i>Protoma kayalensis</i> Dey, 1961 (see below).</p><p>A specimen illustrated as <i>Turritella angulata</i> from the Talar Sandstone, between Garr Koh and Chatti at the Makran margin in southwestern Pakistan (Crame 1984) also belongs to <i>P. harrisoni</i>. The actual <i>Turritella angulata</i> is an Early Miocene species from Kutch in India, and is much more angular due to a prominent keel near the base of the whorls (see Sowerby 1840: pl. 26 fig. 7, or Kulkarni <i>et al.</i> 2010: fig. 2g).</p>