Question:
Hugh Trumble (12 May 1867 - 14 August 1938) was an Australian cricketer who played 32 Test matches as a bowling all-rounder between 1890 and 1904. He captained the Australian team in two Tests, winning both. Trumble took 141 wickets in Test cricket--a world record at the time of his retirement--at an average of 21.78 runs per wicket. He is one of only four bowlers to twice take a hat-trick in Test cricket.
Early in the 1889-90 Australian season, Trumble endured a period where he was not able to take wickets consistently. With selection of the Australian team to tour England in 1890 due at this time, Trumble was anxious about this poor run of form. Noting his anxiety while playing, a friend offered him a beer during the lunch break to revive his spirits. Previously a teetotaler, Trumble enjoyed his first taste and ordered another before re-entering the field of play. Feeling relaxed, although wondering about his steadiness of step, Trumble took a succession of wickets to ensure his selection in the Australian team. Trumble finished the season with 27 wickets at an average of 14.20 per wicket. The 1890 Australian team touring England was relatively inexperienced. The team missed the all-round ability of George Giffen, who had refused to join the squad, thinking it unlikely the tour would be a sporting or financial success. The Australians won 13 matches on tour, losing 16 and drawing 9. Trumble made his Test cricket debut in the First Test against the English team at Lord's Cricket Ground. He took only one wicket, dismissing Bobby Peel caught and bowled for 1. Batting at number eleven in the first innings he made 1 not out and in the second, 5 runs batting at number ten. Despite this lack of success, he retained his spot in the team for the Second Test at The Oval where he failed to take a wicket. He was selected for the Third Test at Old Trafford but continuous rain saw the match abandoned without a ball being bowled. Trumble played 28 first-class matches during the tour, scoring 288 runs at an average of 8.47 and took 52 wickets at an average of 21.75. Wisden Cricketers' Almanack wrote, "Reports from Australia had led us to expect a great deal of ... Trumble" but his "straightness and regular length [were] insufficient to compensate for an obvious lack of 'devil' and variety". Trumble was not selected for the Australian team to play Lord Sheffield's touring English team in 1891-92. He did not return to the Australian team until his selection in the squad to tour England in 1893. Before the Test matches he took 14 wickets for 116 runs (14/116) against the Players followed by 12/84 against Kent at Gravesend. He played in all three Test matches in 1893, taking 6 wickets at an average of 39.00. Trumble scored 58 runs in the Tests with a highest score of 35 but had more success in the other matches, scoring 774 runs, including one century in all first-class matches on tour. Wisden noted that "An immense improvement on his form of three years before was shown by Hugh Trumble, who bowled consistently well all through the tour" and "... the reports of Hugh Trumble's improvement in batting were amply borne out, his hitting in many matches being remarkably fine". When Andrew Stoddart's English team visited Australia in 1894-95, Trumble played only one Test, the Second at the Melbourne Cricket Ground. In the first innings, England scored 75 runs with Trumble taking 3 wickets. England fought back in their second innings, scoring 475 runs to win the Test by 94 runs; Trumble failed to take a wicket.
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Feeling relaxed, although wondering about his steadiness of step, Trumble took a succession of wickets to ensure his selection in the Australian team.

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Richard Charles "Dick" Lewontin (born March 29, 1929) is an American evolutionary biologist, mathematician, geneticist, and social commentator. A leader in developing the mathematical basis of population genetics and evolutionary theory, he pioneered the application of techniques from molecular biology, such as gel electrophoresis, to questions of genetic variation and evolution. In a pair of seminal 1966 papers co-authored with J.L. Hubby in the journal Genetics, Lewontin helped set the stage for the modern field of molecular evolution. In 1979 he and Stephen Jay Gould introduced the term "spandrel" into evolutionary theory.
Lewontin has worked in both theoretical and experimental population genetics. A hallmark of his work has been an interest in new technology. He was the first person to do a computer simulation of the behavior of a single gene locus (previous simulation work having been of models with multiple loci). In 1960 he and Ken-Ichi Kojima were the first population geneticists to give the equations for change of haplotype frequencies with interacting natural selection at two loci. This set off a wave of theoretical work on two-locus selection in the 1960s and 1970s. Their paper gave a theoretical derivation of the equilibria expected, and also investigated the dynamics of the model by computer iteration. Lewontin later introduced the D' measure of linkage disequilibrium. (He also introduced the term "linkage disequilibrium", about which many population geneticists have been unenthusiastic.) In 1966, he and Jack Hubby published a paper that revolutionized population genetics. They used protein gel electrophoresis to survey dozens of loci in the fruit fly Drosophila pseudoobscura, and reported that a large fraction of the loci were polymorphic, and that at the average locus there was about a 15% chance that the individual was heterozygous. (Harry Harris reported similar results for humans at about the same time.) Previous work with gel electrophoresis had been reports of variation in single loci and did not give any sense of how common variation was. Lewontin and Hubby's paper also discussed the possible explanation of the high levels of variability by either balancing selection or neutral mutation. Although they did not commit themselves to advocating neutrality, this was the first clear statement of the neutral theory for levels of variability within species. Lewontin and Hubby's paper had great impact--the discovery of high levels of molecular variability gave population geneticists ample material to work on, and gave them access to variation at single loci. The possible theoretical explanations of this rampant polymorphism became the focus of most population genetics work thereafter. Martin Kreitman was later to do a pioneering survey of population-level variability in DNA sequences while a Ph.D. student in Lewontin's lab.

What explanation did the paper talk about?
Their paper gave a theoretical derivation of the equilibria expected, and also investigated the dynamics of the model by computer iteration.