Read the bio below and try to give details on charles roy henderson 's: - doctoral students - known for - awards - name Bio: charles roy henderson -lrb- -- -rrb- was an american statistician and a pioneer in animal breeding -- the application of quantitative methods for the genetic evaluation of domestic livestock . he developed mixed model equations to obtain best linear unbiased predictions of breeding values and , in general , any random effect . he invented three methods for the estimation of variance components in unbalanced settings of mixed models , and invented a method for constructing the inverse of wright 's numerator relationship matrix based on a simple list of pedigree information . he , with his ph.d. student shayle r. searle , greatly extended the use of matrix notation in statistics . his methods are widely used by the domestic livestock industry throughout the world and are a cornerstone of linear model theory . henderson obtained his b.sc. , m.sc . -lrb- nutrition -rrb- and ph.d. -lrb- breeding -rrb- degrees at iowa state university , where he was a student of professor l. n. hazel . henderson joined the faculty of the department of animal science at cornell university in 1948 , and headed the animal breeding division for nearly 30 years until he retired in 1976 . after retiring from cornell , he was a visiting professor at the university of guelph and university of illinois until his death . he completed his book in 1984 at the university of guelph .
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- doctoral students is shayle r. searle- known for is best variance component estimation linear unbiased prediction -lrb- blup -rrb- mixed models- awards is elected to the united states national academy of sciences- name is charles roy henderson