What type of details about alan grafen can be gathered from the following bio? Bio: alan grafen is a scottish ethologist and evolutionary biologist . he currently teaches and undertakes research at st john 's college , oxford . along with regular contributions to scientific journals , grafen is known publicly for his work as co-editor -lrb- with mark ridley -rrb- of the 2006 festschrift '' '' , honouring the achievements of his colleague and former academic advisor . he has worked extensively in the field of biological game theory , and , in 1990 , devised a model showing that zahavi 's well-known handicap principle could theoretically exist in natural populations . he was elected a fellow of the royal society in 2011
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