IN: Nash was born on July 9, 1959, to a devout Christian family, in southwest Detroit, Michigan. Nash's father, Robert, died of a heart attack on April 4, 1968, aged 36, when Nash was eight years old. On December 27, 1994, Nash's mother, Wanda, died after a four-year struggle against breast cancer. He attended Aquinas High School, followed by the University of Tennessee, where he majored in psychology and minored in educational philosophy.

Nash debuted in Total Nonstop Action Wrestling alongside the returning Scott Hall on November 7, 2004 at the inaugural monthly TNA pay-per-view, Victory Road, with the duo helping NWA World Heavyweight Champion Jeff Jarrett retain his title in a ladder match with Jeff Hardy. In subsequent weeks, the trio identified themselves as the Kings of Wrestling and began feuding with Hardy and A.J. Styles. At Turning Point on December 5, the Kings of Wrestling were defeated by Hardy, Styles, and Randy Savage. Hall left TNA in early 2005, and Nash and Jarrett separated after Nash made clear his desire to win the NWA World Heavyweight Championship. Nash received a title shot against Jarrett on February 13, 2005 at Against All Odds, but lost following interference from the debuting Outlaw. Following the defeat, Nash joined forces with Sean Waltman and began feuding with the newly formed Planet Jarrett. At Destination X on March 13, Nash lost to The Outlaw in a First Blood match following interference from Jarrett, who struck Nash with his title belt. The rivalry between Planet Jarrett and Nash and his allies culminated in a scheduled Lethal Lockdown match at Lockdown on April 24 pitting Nash, Waltman, and Diamond Dallas Page against Jarrett, The Outlaw, and "The Alpha Male" Monty Brown. Nash, however, was removed from the card and replaced with B.G. James after contracting a staph infection, which left him sidelined for much of 2005. Nash returned to TNA on October 1 for the first episode of Impact! on Spike TV, attacking and powerbombing Jarrett. Nash went on to challenge Jarrett to a match for the NWA World Heavyweight Championship at Bound for Glory on October 23. In the weeks preceding the event, Nash and Jarrett had several heated confrontations, on one occasion brawling with one another and with guest referee Tito Ortiz. On October 22, one day before Bound for Glory, Nash was hospitalized with chest pains. At Bound for Glory, a battle royal was held to determine the number one contender; Rhino won and then defeated Jarrett for the NWA World Heavyweight Championship. Nash was later discharged from the hospital, having suffered a mild cardiac episode. He made a partial return to the ring in December 2005, wrestling several matches on a tour of South Africa.
QUESTION: What was this
IN: Richard Charles "Dick" Lewontin (born March 29, 1929) is an American evolutionary biologist, mathematician, geneticist, and social commentator. A leader in developing the mathematical basis of population genetics and evolutionary theory, he pioneered the application of techniques from molecular biology, such as gel electrophoresis, to questions of genetic variation and evolution. In a pair of seminal 1966 papers co-authored with J.L. Hubby in the journal Genetics, Lewontin helped set the stage for the modern field of molecular evolution. In 1979 he and Stephen Jay Gould introduced the term "spandrel" into evolutionary theory.

Lewontin has worked in both theoretical and experimental population genetics. A hallmark of his work has been an interest in new technology. He was the first person to do a computer simulation of the behavior of a single gene locus (previous simulation work having been of models with multiple loci). In 1960 he and Ken-Ichi Kojima were the first population geneticists to give the equations for change of haplotype frequencies with interacting natural selection at two loci. This set off a wave of theoretical work on two-locus selection in the 1960s and 1970s. Their paper gave a theoretical derivation of the equilibria expected, and also investigated the dynamics of the model by computer iteration. Lewontin later introduced the D' measure of linkage disequilibrium. (He also introduced the term "linkage disequilibrium", about which many population geneticists have been unenthusiastic.) In 1966, he and Jack Hubby published a paper that revolutionized population genetics. They used protein gel electrophoresis to survey dozens of loci in the fruit fly Drosophila pseudoobscura, and reported that a large fraction of the loci were polymorphic, and that at the average locus there was about a 15% chance that the individual was heterozygous. (Harry Harris reported similar results for humans at about the same time.) Previous work with gel electrophoresis had been reports of variation in single loci and did not give any sense of how common variation was. Lewontin and Hubby's paper also discussed the possible explanation of the high levels of variability by either balancing selection or neutral mutation. Although they did not commit themselves to advocating neutrality, this was the first clear statement of the neutral theory for levels of variability within species. Lewontin and Hubby's paper had great impact--the discovery of high levels of molecular variability gave population geneticists ample material to work on, and gave them access to variation at single loci. The possible theoretical explanations of this rampant polymorphism became the focus of most population genetics work thereafter. Martin Kreitman was later to do a pioneering survey of population-level variability in DNA sequences while a Ph.D. student in Lewontin's lab.
QUESTION:
What explanation did the paper talk about?