Input: Read this: Genomes are more than the sum of an organism's genes and have traits that may be measured and studied without reference to the details of any particular genes and their products. Researchers compare traits such as chromosome number (karyotype), genome size, gene order, codon usage bias, and GC-content to determine what mechanisms could have produced the great variety of genomes that exist today (for recent overviews, see Brown 2002; Saccone and Pesole 2003; Benfey and Protopapas 2004; Gibson and Muse 2004; Reese 2004; Gregory 2005).
Question: What were other traits besides karyotype studied by Brown in 2002?

Output: unanswerable

Input: Read this: The doctrine of the immaculate conception (Mary being conceived free from original sin) is not to be confused with her virginal conception of her son Jesus. This misunderstanding of the term immaculate conception is frequently met in the mass media. Catholics believe that Mary was not the product of a virginal conception herself but was the daughter of a human father and mother, traditionally known by the names of Saint Joachim and Saint Anne. In 1677, the Holy See condemned the belief that Mary was virginally conceived, which had been a belief surfacing occasionally since the 4th century. The Church celebrates the Feast of the Immaculate Conception (when Mary was conceived free from original sin) on 8 December, exactly nine months before celebrating the Nativity of Mary. The feast of the Annunciation (which commemorates the virginal conception and the Incarnation of Jesus) is celebrated on 25 March, nine months before Christmas Day.
Question: When were contradictory views to the belief of Mary's parentage struck down ?

Output: 1677, the Holy See condemned the belief that Mary was virginally conceived,

Input: Read this: Rex Stout's fictional detective Nero Wolfe is described as fond of solving the London Times' crossword puzzle at his New York home, in preference to those of American papers.
Question: In what part of the world did the fictional detective, Nero Wolfe, live in?

Output: New York

Input: Read this: During the Middle Ages, Thuringia was situated at the border between Germanic and Slavic territories, marked by the Saale river. The Ostsiedlung movement led to the assimilation of Slavic people between the 11th and the 13th century under German rule. The population growth increased during the 18th century and stayed high until World War I, before it slowed within the 20th century and changed to a decline since 1990. Since the beginning of Urbanisation around 1840, the Thuringian cities have higher growth rates resp. smaller rates of decline than rural areas (many villages lost half of their population since 1950, whereas the biggest cities (Erfurt and Jena) keep growing).
Question: Where was Thuringia in the Middle Ages?

Output:
the border between Germanic and Slavic territories