This effectively means that populations of organisms must have reached a certain measurable level of difference to be recognised as subspecies. Dean Amadon proposed in 1949 that subspecies would be defined according to the seventy-five percent rule which means that 75% of a population must lie outside 99% of the range of other populations for a given defining morphological character or a set of characters. The seventy-five percent rule still has defenders but other scholars argue that it should be replaced with ninety or ninety-five percent rule.
If it is possible to answer this question, answer it for me (else, reply "unanswerable"): What must organisms have a measurable level of to be seen as a subspecies?
Ah, so.. difference

Most organisms studied to date utilize a rewardpunishment mechanism: for instance, worms and insects can alter their behavior to seek food sources or to avoid dangers. In vertebrates, the reward-punishment system is implemented by a specific set of brain structures, at the heart of which lie the basal ganglia, a set of interconnected areas at the base of the forebrain. There is substantial evidence that the basal ganglia are the central site at which decisions are made: the basal ganglia exert a sustained inhibitory control over most of the motor systems in the brain; when this inhibition is released, a motor system is permitted to execute the action it is programmed to carry out. Rewards and punishments function by altering the relationship between the inputs that the basal ganglia receive and the decision-signals that are emitted. The reward mechanism is better understood than the punishment mechanism, because its role in drug abuse has caused it to be studied very intensively. Research has shown that the neurotransmitter dopamine plays a central role: addictive drugs such as cocaine, amphetamine, and nicotine either cause dopamine levels to rise or cause the effects of dopamine inside the brain to be enhanced.
If it is possible to answer this question, answer it for me (else, reply "unanswerable"): Which of the two systems, reward or punishment is better understood?
Ah, so.. The reward mechanism

The Kingdom of Prussia became the leading state of the German Empire after its creation in 1871. However, the Treaty of Versailles following World War I granted West Prussia to Poland and made East Prussia an exclave of Weimar Germany (the new Polish Corridor separating East Prussia from the rest of Germany), while the Memel Territory was detached and was annexed by Lithuania in 1923. Following Nazi Germany's defeat in World War II in 1945, war-torn East Prussia was divided at Joseph Stalin's insistence between the Soviet Union (the Kaliningrad Oblast in the Russian SFSR and the constituent counties of the Klaipeda Region in the Lithuanian SSR) and the People's Republic of Poland (the Warmian-Masurian Voivodeship). The capital city Konigsberg was renamed Kaliningrad in 1946. The German population of the province was largely evacuated during the war or expelled shortly thereafter in the expulsion of Germans after World War II. An estimated 300,000 (around one fifth of the population) died either in war time bombings raids or in the battles to defend the province.[citation needed]
If it is possible to answer this question, answer it for me (else, reply "unanswerable"): What year did the Kingdom of Prussia become the leading state of the German Empire?
Ah, so..
1871