Given the task definition and input, reply with output. In this task you will be given a passage and a yes/no question based on the passage. You should answer the question using the information from the passage.

passage: Unlike oxidative phosphorylation, oxidation and phosphorylation are not coupled in the process of substrate-level phosphorylation, and reactive intermediates are most often gained in the course of oxidation processes in catabolism. Most ATP is generated by oxidative phosphorylation in aerobic or anaerobic respiration while substrate-level phosphorylation provides a quicker, less efficient source of ATP, independent of external electron acceptors. This is the case in human erythrocytes, which have no mitochondria, and in oxygen-depleted muscle.
question: can substrate level phosphorylation occur in the absence of oxygen?
Yes