Brower, A.V.Z. og Warren, A. (2007) på Burns, J.M. & Janzen, D.H. (2005) Pan-Neotropical genus (Hesperiidae; Pyrginae) is not monotypic: four new species occur on one volcano in the Area de Conservación Guanacaste, Costa Rica. 59: 19-34. Markku Savela's Lepidoptera and some other life forms: Sjekkliste over artene. Hentet 31.5. 2012.