Cambrian explosion

* made a big contribution to interest in the Cambrian explosion. ]]
* specimen shows how clear and detailed the fossils are from the Burgess Shale lagerstätte.]]
.]]
*, an Ediacaran animal with a quilted appearance.]]
.  This specimen of *Olenoides serratus'', from the Burgess shale, preserves 'soft parts'&nbsp;– the antennae and legs. ]]

The **Cambrian explosion** is when many animal phyla first appeared in the fossil record. It happened 541 million years ago (mya). Probably most had evolved before then, but that was their first appearance as fossils.

Before about 580 mya it seems that most organisms were simple. They were made of individual cells, occasionally organized into colonies. Over the following 70 or 80 million years the rate of evolution seemed to accelerate. By the end of the Cambrian most phyla we know today existed.

The Cambrian explosion has caused much scientific debate. The seemingly rapid appearance of fossils in the 'primordial strata' was noted as early as the mid 19th century,  and Charles Darwin saw it as one of the main objections that could be made against his theory of evolution by natural selection.

## Key issues 
The long-running puzzlement about the appearance of the Cambrian fauna, seemingly abruptly and from nowhere, centres on three key points: 
#whether it actually happened. Was there really a mass diversification of complex organisms over a relatively short period of time during the early Cambrian?
#what might have caused such rapid change.
#what it might say about the origin and evolution of animals.
Understanding it is difficult because the supply of evidence is limited. The evidence is an incomplete fossil record, and chemical signatures left in Cambrian rocks.

### Large-scale evolution 
The Cambrian explosion is the classic example of megaevolution. By that is not meant a different kind of evolution; rather, it means evolution which produces a tremendous effect.

Precambrian marine diversity was dominated by small fossils known as acritarchs. This term describes almost any small organic walled fossilfrom the egg cases of small metazoans to resting cysts of many different kinds of green algae. After appearing around 2000 mya, acritarchs had a boom around 1000 mya, increasing in number, diversity, size, complexity of shape and especially size and number of spines. Their increasingly spiny forms in the last billion years may indicate an increased need for defence against predation. Other groups of small organisms from the Neoproterozoic era also show signs of anti-predator defences. seems to show an increase in predation around this time.
However, in general, the rate of evolution in the Precambrian was very slow, with many cyanobacterial species lasting unchanged for billions of years. Of course, bacteria are defined mainly by their biochemistry, especially their genomes. Changes in their biochemistry would usually leave no trace in the fossil record.

If the predatory organisms which grazed on bacteria and acritarchs really were metazoans, this means that Cambrian animals did not appear "from nowhere" at the base of the Cambrian; their ancestors had existed for hundreds of millions of years.

## Related pages 
* Life timeline

## Other sources 
*Gould S.J. 1989. *Wonderful life: the Burgess Shale and the nature of history*. W.W. Norton & Company.
*Conway Morris S. 1997. *The crucible of creation: the Burgess Shale and the rise of animals*. Oxford University Press. .
* 

## References