Ptomaphagus

***Ptomaphagus*** er en slekt av mycelbiller som særlig finnes knyttet til sopp og åtsler.

## Utseende
Små  (2-4 millimeter), halvmatte, mørkbrunlige mycelbiller, kroppsformen nærmest dråpeformet med lange bein. Antennene er litt kølleformede og 11-leddete. De fem ytterste leddene danner en avlang klubbe, men det åttende leddet er mye mindre enn de andre og er vanligvis forkortet til en liten skive. Hodet er oftest kraftig innskåret bak fasettøynene, med en smal hals, men dette kan være vanskelig å se da billene gjerne trekker hodet inn under brystskjoldet. Hannen har utvidete føtter på forbeina.

## Levevis
Artene kan finnes på små åtsler, mellom råtnende plantemateriale og av og til i musereir.

## Systematisk inndeling

* Ordenen biller, **Coleoptera**
** Underordenen Polyphaga
*** Overfamilien åtsel- og rovbiller, **Staphylinoidea**
**** Familien **[[mycelbiller]], [[Leiodidae]]** Fleming, 1821
***** Underfamilien kadaverbiller, **[[Cholevinae]]**
****** Stammen Ptomaphagini
******* Slekten ***Ptomaphagus*** Hellwig, 1795  – tre arter i Norge
******** *[[Ptomaphagus acuminatus]]* Salgado, 2011
******** *[[Ptomaphagus acutus]]* Peck and Gnaspini, 1997
******** *[[Ptomaphagus agilis]]* (P.H.Lucas, 1847)
******** *[[Ptomaphagus alleghenyensis]]* (Peck, 1979)
******** *[[Ptomaphagus altus]]* Peck, 1973
******** *[[Ptomaphagus appalachianus]]* (Peck, 1979)
******** *[[Ptomaphagus argenticornis]]* Peck, 1977
******** *[[Ptomaphagus aritzensis]]* Jeannel, 1934
******** *[[Ptomaphagus barbarae]]* Peck, 1973
******** *[[Ptomaphagus barri]]* Peck, 1973
******** *[[Ptomaphagus bedfordensis]]* (Peck, 1979)
******** *[[Ptomaphagus biharicus]]* Jeannel, 1934
******** *[[Ptomaphagus boliviensis]]* Salgado, 2014
******** *[[Ptomaphagus bordoni]]* (Jeannel, 1965)
******** *[[Ptomaphagus brevior]]* Jeannel, 1949
******** *[[Ptomaphagus californicus]]* (LeConte, 1854)
******** *[[Ptomaphagus caucasicus]]* Jeannel, 1934
******** *[[Ptomaphagus cavernicola]]* Schwarz, 1898
******** *[[Ptomaphagus championi]]* Jeannel, 1936
******** *[[Ptomaphagus chendae]]* J.Müller, 1921
******** *[[Ptomaphagus chenggongi]]* Wang, Cheng-Bin, Nishikawa, Perreau, Růžička & Y.Hayashi, 2016
******** *[[Ptomaphagus chihuahuensis]]* (Peck, 1976)
******** *[[Ptomaphagus chromolithus]]* Peck, 1984
******** *[[Ptomaphagus circassicus]]* (Reitter, 1888)
******** *[[Ptomaphagus clavalis]]* (Reitter, 1884)
******** *[[Ptomaphagus cocytus]]* Peck, 1973
******** *[[Ptomaphagus colima]]* Peck, 1977
******** *[[Ptomaphagus conejera]]* Peck, 1977
******** *[[Ptomaphagus consobrinus]]* (LeConte, 1853)
******** *[[Ptomaphagus costanama]]* Peck, 2003
******** *[[Ptomaphagus costarica]]* Peck, 2003
******** *[[Ptomaphagus cubensis]]* Salgado, 2012
******** *[[Ptomaphagus cumberlandus]]* (Peck, 1979)
******** *[[Ptomaphagus dacicus]]* Jeannel, 1934
******** *[[Ptomaphagus davisii]]* Stephens, 1830
******** *[[Ptomaphagus delsur]]* Peck, 1977
******** *[[Ptomaphagus densus]]* Hatch, 1957
******** *[[Ptomaphagus distinctus]]* Peck, 1977
******** *[[Ptomaphagus divaricatus]]* Jeannel, 1934
******** *[[Ptomaphagus elabra]]* Peck, 1971
******** *[[Ptomaphagus episcopus]]* Peck, 1973
******** *[[Ptomaphagus fecundus]]* (Barr, 1963)
******** *[[Ptomaphagus fiskei]]* Peck, 1973
******** *[[Ptomaphagus fisus]]* Horn, 1885
******** *[[Ptomaphagus flabellatus]]* Szymczakowski, 1961
******** *[[Ptomaphagus forticornis]]* A.Matthews, 1888
******** *[[Ptomaphagus fumosus]]* (Peck, 1979)
******** *[[Ptomaphagus funiu]]* Wang, Cheng-Bin, Perreau, Růžička & Nishikawa, 2017
******** *[[Ptomaphagus geomysi]]* Peck and Skelley, 2002
******** *[[Ptomaphagus germari]]* Schlechtendahl, 1888
******** *[[Ptomaphagus giaquintoi]]* Jeannel, 1936
******** *[[Ptomaphagus gypsum]]* Peck, 1973
******** *[[Ptomaphagus haba]]* Wang, Cheng-Bin, Perreau, Růžička & Nishikawa, 2017
******** *[[Ptomaphagus hastatellus]]* Szymczakowski, 1976
******** *[[Ptomaphagus hatchi]]* Jeannel, 1933
******** *[[Ptomaphagus hazelae]]* Peck, 1973
******** *[[Ptomaphagus hayashii]]* Wang, 2016
******** *[[Ptomaphagus hirtus]]* (Tellkampf, 1844)
******** *[[Ptomaphagus hispaniolensis]]* Peck & Cook, 2014
******** *[[Ptomaphagus hubrichti]]* Barr, 1958
******** *[[Ptomaphagus inbio]]* Peck, 2003
******** *[[Ptomaphagus inyoensis]]* Peck and Gnaspini, 1997
******** *[[Ptomaphagus izabal]]* Perreau, 2013
******** *[[Ptomaphagus jalisco]]* Peck, 1977
******** *[[Ptomaphagus jamesi]]* Peck, 1973
******** *[[Ptomaphagus joanna]]* (Peck, 1979)
******** *[[Ptomaphagus jonesi]]* (Peck, 1979)
******** *[[Ptomaphagus julius]]* Peck, 1973
******** *[[Ptomaphagus kuntzeni]]* Sokolowski, 1957
******** *[[Ptomaphagus laselva]]* Peck, 1977
******** *[[Ptomaphagus laticornis]]* Jeannel, 1949
******** *[[Ptomaphagus leo]]* Peck, 1973
******** *[[Ptomaphagus lincolnensis]]* Peck, 1978
******** *[[Ptomaphagus liquidambar]]* Peck, 1977
******** *[[Ptomaphagus loedingi]]* Hatch, 1933
******** *[[Ptomaphagus longicornis]]* Jeannel, 1949
******** *[[Ptomaphagus manzano]]* Peck, 1978
******** *[[Ptomaphagus masumotoi]]* Nishikawa, 2011
******** *[[Ptomaphagus mckenziei]]* Peck, 1977
******** *[[Ptomaphagus medius]]* (Rey, 1889)  - finnes i Norge
******** *[[Ptomaphagus merritti]]* Tishechkin, 2007
******** *[[Ptomaphagus meximontanus]]* Peck, 1973
******** *[[Ptomaphagus microps]]* Peck, 1978
******** *[[Ptomaphagus mitchellensis]]* (Hatch, 1933)
******** *[[Ptomaphagus nashvillensis]]* (Peck, 1979)
******** *[[Ptomaphagus neojonesi]]* Peck & Newton, 2017
******** *[[Ptomaphagus nepalensis]]* Perreau, 1988
******** *[[Ptomaphagus nevadicus]]* Horn, 1880
******** *[[Ptomaphagus newtoni]]* Peck, 1973
******** *[[Ptomaphagus nubloso]]* Peck, 2003
******** *[[Ptomaphagus oaxaca]]* Peck, 1973
******** *[[Ptomaphagus orichalcum]]* (Peck, 1979)
******** *[[Ptomaphagus oriental]]* Peck, 1977
******** *[[Ptomaphagus parashant]]* Peck and Wynne, 2013
******** *[[Ptomaphagus pecki]]* Gnaspini, 1996
******** *[[Ptomaphagus piccoloi]]* Wang, Cheng-Bin, Růžička, Nishikawa, Perreau & Y.Hayashi, 2016
******** *[[Ptomaphagus piperi]]* Hatch, 1933
******** *[[Ptomaphagus pisgahensis]]* (Peck, 1979)
******** *[[Ptomaphagus pius]]* (Seidlitz, 1887)
******** *[[Ptomaphagus pruina]]* Peck, 1977
******** *[[Ptomaphagus pruinosus]]* (Reitter, 1881)
******** *[[Ptomaphagus puertoricensis]]* Peck, 1972
******** *[[Ptomaphagus pyrenaeus]]* Jeannel, 1934
******** *[[Ptomaphagus quercus]]* Peck, 1977
******** *[[Ptomaphagus reddelli]]* Peck, 1973
******** *[[Ptomaphagus rhagianicus]]* Sbordoni, 1967
******** *[[Ptomaphagus richlandensis]]* (Peck, 1979)
******** *[[Ptomaphagus ruficollis]]* Portevin, 1903
******** *[[Ptomaphagus sardus]]* (Seidlitz, 1887)
******** *[[Ptomaphagus schwarzi]]* Hatch, 1933
******** *[[Ptomaphagus sciakyi]]* (Zoia, 1993)
******** *[[Ptomaphagus scottsboroensis]]* (Peck, 1979)
******** *[[Ptomaphagus sericatus]]* (Chaudoir, 1845) - finnes i Norge
******** *[[Ptomaphagus setiger]]* (Horn, 1885)
******** *[[Ptomaphagus shapardi]]* Sanderson, 1939
******** *[[Ptomaphagus sibiricus]]* Jeannel, 1934
******** *[[Ptomaphagus solanum]]* Peck, 1973
******** *[[Ptomaphagus sonorensis]]* (Peck, 1976)
******** *[[Ptomaphagus spelaeus]]* (Bilimek, 1867)
******** *[[Ptomaphagus steevesi]]* (Peck, 1979)
******** *[[Ptomaphagus subtruncatus]]* Mäklin, 1881
******** *[[Ptomaphagus subvillosus]]* (Goeze, 1777) - finnes i Norge
******** *[[Ptomaphagus suteri]]* (Peck, 1979)
******** *[[Ptomaphagus szymczakowskii]]* Peck, 1977
******** *[[Ptomaphagus tabascensis]]* Sbordoni, 1974
******** *[[Ptomaphagus talamanca]]* Peck, 1973
******** *[[Ptomaphagus tauricus]]* Jeannel, 1934
******** *[[Ptomaphagus tenuicornis]]* (Rosenhauer, 1856)
******** *[[Ptomaphagus texanus]]* Melander, 1902
******** *[[Ptomaphagus thebeatles]]* Schilthuizen *et al*., 2020
******** *[[Ptomaphagus thomomysi]]* Hatch, 1957
******** *[[Ptomaphagus tierrabaja]]* Peck, 1977
******** *[[Ptomaphagus tingtingae]]* Wang, Cheng-Bin, Nishikawa, Perreau, Růžička & Y.Hayashi, 2016
******** *[[Ptomaphagus torodei]]* Peck, 1984
******** *[[Ptomaphagus trezzii]]* Perreau, 2009
******** *[[Ptomaphagus troglodytes]]* Blas & Vives, 1983
******** *[[Ptomaphagus troglomexicanus]]* Peck, 1968
******** *[[Ptomaphagus tuza]]* Peck, 1977
******** *[[Ptomaphagus ulkei]]* Horn, 1885
******** *[[Ptomaphagus valentinei]]* Jeannel, 1933
******** *[[Ptomaphagus validus]]* (Kraatz, 1852)
******** *[[Ptomaphagus vallombrosae]]* (Seidlitz, 1887)
******** *[[Ptomaphagus varicornis]]* (rosenhauer, 1847) - finnes i Norge
******** *[[Ptomaphagus volcanica]]* Peck, 1977
******** *[[Ptomaphagus walteri]]* Peck, 1973
******** *[[Ptomaphagus whiteselli]]* Barr, 1963
******** *[[Ptomaphagus yasutoshii]]* Nishikawa, 1993
******** *[[Ptomaphagus yuvila]]* Peck, 1977

## Referanser




## Kilder
* Leiodidae i Tree of life

* Silfverberg, H. 1992. *Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae* (liste over Nordens biller). Helsinki.

## Eksterne lenker
* Artsdatabanken - *Ptomaphagus* https://artsdatabanken.no/ScientificName/Ptomaphagus/11149
* *Ptomapahgus* på GBIF https://www.gbif.org/species/3261754/treatments
*