Zygadenia

***Zygadenia*** er en slekt av utdødde biller som trolig levde i døde eller dødende trær eller deres røtter. Slekten levde i mesozoikum. Det blir nå plassert ca. 70 fossile arter i slekten, noe som gjør den til ens vært artrsik fossil-slekt.

## Utseende
Ommatidae er generelt mørke, slanke, flate biller, dekkvingene har opphøyde ribber og nettmønster. Kjevene (*mandiblene*) er fremoverrettede og ganske store. Antennene er mer eller mindre sylindriske eller perlekjede-aktige.

## Historie
Ommatidae er en gammel gruppe som åpenbart må ha vært mye mer artsrik i tidligere tidsperioder. Arten plassert i denne slekten er kjent fra mye av jordens mellomalder, fra trias, jura og kritt, mellom 232-94 millioner år før nåtid, fra det næværende England, Tyskland, Spania, Russland, Kasakhstan, Mongolia, Kina, Australia og Argentina.

## Systematisk inndeling
Disse artene har tidligere vært plassert i en rekke ulike slekter, se liste i taksoboksen.

* Ordenen biller, Coleoptera
** Underordenen Archostemata
*** Familien Ommatidae Sharp & Muir, 1912
**** Gruppen Notocupedini Ponomarenko, 1966
***** Slekten ***Zygadenia*** Handlirsch, 1906 
****** †*[[Zygadenia aliena]]* (Tan and Ren 2006) Yixian Formation, China, Aptian
****** †*[[Zygadenia angliae]]* (Giebel 1856) Purbeck Group, United Kingdom, Berriasian, Weald Clay, United Kingdom, Barremian
****** †*[[Zygadenia brachycephalus]]* (Ponomarenko, 1994) Bakhar Formation, Mongolia, Bajocian, Shar Teeg, Mongolia, Tithonian
****** †*[[Zygadenia caducus]]* (Ponomarenko, 1969) Kzyl-Zhar, Kazakhstan, Turonian
****** †*[[Zygadenia caudatus]]* (Ponomarenko, 1966) Zaza Formation, Russia, Aptian
****** †*[[Zygadenia cellulosus]]* (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
****** †*[[Zygadenia crassus]]* (Ponomarenko, 1969) Sulyukta Formation, Kyrgyzstan, Toarcian
****** †*[[Zygadenia diazromerali]]* Soriano & Delclòs, 2006 Las Hoyas, Spain, Barremian
****** †*[[Zygadenia dischides]]* (Zhang 1986) Jiulongshan Formation, China, Callovian
****** †*[[Zygadenia dundulaensis]]* (Ponomarenko, 1994) Dzun-Bain Formation, Mongolia, Aptian
****** †*[[Zygadenia elegans]]* (Ponomarenko, 1994) Dzun-Bain Formation, Mongolia, Aptian
****** †*[[Zygadenia epicharis]]* (Tan et al. 2005) Yixian Formation, China, Aptian
****** †*[[Zygadenia eumeura]]* (Tan et al. 2006) Yixian Formation, China, Aptian
****** †*[[Zygadenia excellens]]* (Ponomarenko, 1966)  Zaza Formation, Russia, Aptian
****** †*[[Zygadenia exiguus]]* (Ponomarenko, 1994) Bakhar Formation, Mongolia, Bajocian, Shar Teeg, Mongolia, Tithonian
****** †*[[Zygadenia floodpagei]]* Jarzembowski et al. 2015 Weald Clay, United Kingdom, Hauterivian-Barremian
****** †*[[Zygadenia foersteri]]* (Ponomarenko, 1971) Karabastau Formation, Kazakhstan, Oxfordian
****** †*[[Zygadenia giebeli]]* Ponomarenko 2014 Shar Teeg, Mongolia, Tithonian
****** †*[[Zygadenia handlirschi]]* Ponomarenko 2014 Shar Teeg, Mongolia, Tithonian
****** †*[[Zygadenia homorus]]* (Lin, 1986) Guanyintan Formation, China, Hettangian
****** †*[[Zygadenia issykkulensis]]* (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
****** †*[[Zygadenia jurassica]]* Hong 1983 Jiulongshan Formation, China, Callovian
****** †*[[Zygadenia kezuoensis]]* Hong 1987 Shahai Formation, China, Aptian
****** †*[[Zygadenia khasurtyiensis]]* (Strelnikova 2019) Khasurty, Russia, Aptian
****** †*[[Zygadenia khetanensis]]* (Ponomarenko in Cromov et al., 1993) Emanra Formation, Russia, Turonian
****** †*[[Zygadenia kirghizicus]]* (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
****** †*[[Zygadenia laetus]]* (Lin, 1976)  Jianshangou Formation, China, Barremian
****** †*[[Zygadenia laiyangensis]]* (Hong, 1990) Laiyang Formation, China, Aptian
****** †*[[Zygadenia lapidarius]]* (Ponomarenko, 1968) Karabastau Formation, Kazakhstan, Oxfordian
****** †*[[Zygadenia laticella]]* (Ponomarenko, 1969) Madygen Formation, Kyrgyzstan, Carnian
****** †*[[Zygadenia latus]]* (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
****** †*[[Zygadenia lentus]]* (Ren, 1995) Dabeigou Formation, China, Hauterivian
****** †*[[Zygadenia lini]]* (Ponomarenko et al. 2012) Guanyintan Formation, China Toarcian/Aalenian
****** †*[[Zygadenia liui]]* Jarzembowski et al. 2015 Yixian Formation, China, Aptian
****** †*[[Zygadenia longicollis]]* (Ponomarenko, 1994) Bakhar Formation, Mongolia, Bajocian
****** †*[[Zygadenia longicoxa]]* Soriano & Delclòs, 2006  Las Hoyas, Spain, Barremian
****** †*[[Zygadenia ludongensis]]* (Wang and Liu 1996) Laiyang Formation, China, Aptian
****** †*[[Zygadenia martinae]]* Jarzembowski et al. 2015 Cattamarra Coal Measures, Australia, Toarcian - bør kanskje plasseres i en egen slekt
****** †*[[Zygadenia martinclosas]]* Soriano & Delclòs, 2006 La Pedrera de Rúbies Formation, Las Hoyas, Spain, Barremian
****** †*[[Zygadenia miniscula]]* Tan et al. 2006, Yixian Formation, China, Aptian
****** †*[[Zygadenia mongolicus]]* (Ponomarenko, 1994) Ulan-Argalant Formation, Mongolia, Barremian
****** †*[[Zygadenia nigromonticola]]* (Ponomarenko, 1968) Karabastau Formation, Kazakhstan, Oxfordian
****** †*[[Zygadenia oculata]]* Soriano & Delclòs, 2006 La Pedrera de Rúbies Formation, Spain, Barremian
****** †*[[Zygadenia oxypygus]]* (Ponomarenko, 1969) Koldzat Formation, Kazakhstan, Carnian
****** †*[[Zygadenia patulus]]* Ponomarenko, 1985 Tyumen Formation, Russian, Callovian
****** †*[[Zygadenia picturatus]]* (Ponomarenko, 1964) Karabastau Formation
****** †*[[Zygadenia pingi]]* (Ponomarenko and Ren 2010) Daohugou, China, Callovian
****** †*[[Zygadenia porrecta]]* Tan et al. 2006 Yixian Formation, China, Aptian
****** †*[[Zygadenia premeris]]* (Lee, Nam & Li, 2022) Jinju Formation, Korea, Albian
****** †*[[Zygadenia protensa]]* Tan et al. 2006 Yixian Formation, China, Aptian
****** †*[[Zygadenia psilata]]* Tan et al. 2005 Yixian Formation, China, Aptian
****** †*[[Zygadenia pulcher]]* (Ponomarenko, 1968) Karabastau Formation, Kazakhstan, Oxfordian
****** †*[[Zygadenia reticulatus]]* (Oppenheim, 1888) Solnhofen, Germany, Tithonian
****** †*[[Zygadenia rostratus]]* (Ponomarenko, 1969) Madygen Formation, Kyrgyzstan, Carnian
****** †*[[Zygadenia rudis]]* Tan et al. 2005 Yixian Formation, China, Aptian
****** †*[[Zygadenia semen]]* (Ponomarenko, 2000) Argun Formation, Russia, Aptian
****** †*[[Zygadenia shiluoensis]]* Hong 1984 Tongchuan Formation, China, Ladinian
****** †*[[Zygadenia sibirica]]* Ponomarenko, 2000 Argun Formation, Russia, Aptian
****** †*[[Zygadenia simpsoni]]* Jarzembowski et al. 2015 Weald Clay, United Kingdom, Hauterivian-Barremian
****** †*[[Zygadenia siniestri]]* Soriano & Delclòs, 2006 Las Hoyas, Spain, Barremian
****** †*[[Zygadenia sinitzae]]* Ponomarenko, 2000 Argun Formation, Russia, Aptian
****** †*[[Zygadenia sogutensis]]* (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
****** †*[[Zygadenia stabilis]]* Tan et al. 2005 Yixian Formation, China, Aptian
****** †*[[Zygadenia tenuis]]* (Ponomarenko, 1969) Madygen Formation, Kyrgyzstan, Carnian
****** †*[[Zygadenia tripartitus]]* (Oppenheim, 1888) Solnhofen, Germany, Tithonian
****** †*[[Zygadenia tuanwangensis]]* (Hong, 1990) Laiyang Formation, China, Aptian
****** †*[[Zygadenia tuberculatus]]* (Giebel, 1856) Purbeck Group, United Kingdom, Berriasian
****** †*[[Zygadenia valida]]* Lin 1976 Jehol Group, China, Aptian
****** †*[[Zygadenia viridis]]* Soriano & Delclòs, 2006 Las Hoyas, Spain, Barremian
****** †*[[Zygadenia vitimensis]]* (Ponomarenko, 1966) Zaza Formation, Russia, Aptian
****** †*[[Zygadenia westraliensis]]* Riek 1968 Cattamarra Coal Measures, Australia, Toarcian - bør kanskje plasseres i en egen slekt

## Referanser


## Eksterne lenker
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