Tachys

***Tachys*** er slekt av små løpebiller, gjerne rødbrune på farge.

## Utseende
Små (gjerne 2-3 millimeter) løpebiller, kroppsformen forholdsvis slank og oftest litt flat, overflaten oftest blank. Mange er mer eller mindre rødbrune, men det finnes også mange blanksvarte eller metalliske arter. **Hodet** er ofte forholdsvis lite og smalt med store, fremstrakte overkjever (mandibler). Fasettøynene er middels store, litt utstående. Antennene er vanligvis lange og tynne. Det ytterste leddet på maksillarpalpene (munnfølere) er redusert til en nålefin tapp. **Brystskjoldet** (pronotum) er rundet, vanligvis med markerte bakhjørner. **Dekkvingene** er jevnbrede uten framtredende "skuldre", gjerne uten tydelige punktrader med med noen oppstående børster. **Beina** er nokså lange og slanke.

## Levevis
Skyggeløperne kan finnes i mange ulike miljøer på bakken, og er ofte svært tallrike. De er rovdyr og angriper ulike små byttedyr. De fleste er raske løpere, men en god del arter har mistet flygeevnen.

## Systematisk inndeling

* Ordenen biller,** Coleoptera** Linnaeus, 1758
** Underordenen Adephaga 
*** Overfamilien løpebiller og vannkalver, Caraboidea
**** Familien løpebiller, Carabidae Latreille, 1802
***** Underfamilien **[[skyggeløpere]], [[Trechinae]]** Bonelli, 1810
****** Stammen Bembidiini Stephens, 1827  
******* Gruppen Tachyina Motschulsky, 1862
******** Slekten ***Tachys*** Dejean, 1821
### Liste over artene
* *[[Tachys abruptus]]* Darlington, 1934 c g
* *[[Tachys aeneipennis]]* Motschulsky, 1862 i c g
* *[[Tachys aequinoctialis]]* (Motschulsky, 1855) c g
* *[[Tachys albipes]]* Leconte, 1863
* *[[Tachys algiricus]]* (Lucas, 1846) c g
* *[[Tachys ambulatus]]* Darlington, 1962 c g
* *[[Tachys androyanus]]* (Jeannel, 1946) c g
* *[[Tachys angustulus]]* Reitter, 1899 c g
* *[[Tachys apex]]* Darlington, 1962 c g
* *[[Tachys arcanicola]]* Blackburn, 1878 i c g
* *[[Tachys arculus]]* Fauvel, 1882 c g
* *[[Tachys argentinicus]]* Csiki, 1928 c g
* *[[Tachys artensis]]* Montrouzier, 1860
* *[[Tachys asemus]]* (Basilewsky, 1968) c g
* *[[Tachys assamicus]]* (Jedlicka, 1964) c g
* *[[Tachys atamuradovi]]* Kryzhanovskij & Mikhailov, 1987 c g
* *[[Tachys atomarius]]* Wollaston, 1867 c g
* *[[Tachys atomus]]* Blackburn, 1878 i c g
* *[[Tachys atratus]]* A.Costa, 1883 c g
* *[[Tachys atridermis]]* Sloane, 1921 c g
* *[[Tachys austinicus]]* Casey, 1918
* *[[Tachys australicus]]* Sloane, 1896 c g
* *[[Tachys avius]]* Darlington, 1962 c g
* *[[Tachys baldiensis]]* Blackburn, 1891 c g
* *[[Tachys barbieri]]* Straneo, 1953 c g
* *[[Tachys bathyglyptus]]* Andrewes, 1925 c g
* *[[Tachys beatus]]* Darlington, 1962 c g
* *[[Tachys beaumonti]]* Casey, 1918 c g
* *[[Tachys bicoloratus]]* Burgeon, 1935 c g
* *[[Tachys bistriatus]]* (Duftschmid, 1812) c g
* *[[Tachys blackburni]]* Sloane, 1921 c g
* *[[Tachys blemoides]]* (Jeannel, 1946) c g
* *[[Tachys bloetei]]* Andrewes, 1930 c g
* *[[Tachys bolellus]]* Darlington, 1962 c g
* *[[Tachys bolus]]* Darlington, 1962 c g
* *[[Tachys bonariensis]]* Steinheil, 1869 c g
* *[[Tachys borkuensis]]* (Bruneau de Miré, 1952) c g
* *[[Tachys bouchardi]]* Andrewes, 1931 c g
* *[[Tachys brachys]]* Andrewes, 1925 c g
* *[[Tachys bradycellinus]]* Hayward, 1900 i c g
* *[[Tachys bredoi]]* (Basilewsky, 1953) c g
* *[[Tachys brevicornis]]* Chaudoir, 1846
* *[[Tachys brunneus]]* Andrewes, 1925 c g
* *[[Tachys brunniceps]]* Andrewes, 1930 c g
* *[[Tachys bryanti]]* Lindroth, 1966 i c g
* *[[Tachys caffer]]* Péringuey, 1896 c g
* *[[Tachys caheni]]* (Basilewsky, 1972) c g
* *[[Tachys caledonicus]]* Csiki, 1928 c g
* *[[Tachys callipygus]]* (Boheman, 1858) c g
* *[[Tachys cameroni]]* Andrewes, 1925 c g
* *[[Tachys captus]]* Blackburn, 1888 c g
* *[[Tachys cardioderus]]* Chaudoir, 1850 c g
* *[[Tachys cardoni]]* Andrewes, 1925 c g
* *[[Tachys carib]]* Darlington, 1936 c g
* *[[Tachys carinulatus]]* Sloane, 1921 c g
* *[[Tachys caspius]]* Kolenati, 1845 c g
* *[[Tachys castaneicolor]]* Bates, 1882 c g
* *[[Tachys centralis]]* J.sahlberg, 1900
* *[[Tachys centriustatus]]* Reitter, 1894
* *[[Tachys centromaculatus]]* Wollaston, 1864 c g
* *[[Tachys chiriquinus]]* Bates, 1882 c g
* *[[Tachys cinctus]]* Putzeys, 1875 c g
* *[[Tachys collardi]]* (Basilewsky, 1955) c g
* *[[Tachys colonicus]]* Casey, 1918 c g
* *[[Tachys columbiensis]]* Hayward, 1900 i c g
* *[[Tachys corax]]* Leconte, 1852
* *[[Tachys coriaceus]]* Broun, 1908 c g
* *[[Tachys crypticola]]* (Britton, 1960) c g
* *[[Tachys cubax]]* Darlington, 1934 c g
* *[[Tachys curtulus]]* (Basilewsky, 1953) c g
* *[[Tachys curvans]]* Bates, 1882
* *[[Tachys cycloderus]]* Bates, 1871 c g
* *[[Tachys cyprius]]* Coulon, 2004 c g
* *[[Tachys delamarei]]* (Jeannel, 1962) c g
* *[[Tachys delicatus]]* Andrewes, 1925 c g
* *[[Tachys dentatus]]* Andrewes, 1925 c g
* *[[Tachys descarpentriesi]]* (Bruneau de Miré, 1965) c g
* *[[Tachys devroeyi]]* Burgeon
* *[[Tachys diaphanus]]* Casey, 1918
* *[[Tachys dimediatus]]* Motschulsky, 1849 c g
* *[[Tachys dimidiatus]]* Motschulsky, 1849 g
* *[[Tachys diminutus]]* Bates, 1871 c g
* *[[Tachys diploharpinus]]* Bates, 1878 c g
* *[[Tachys discipennis]]* Fauvel, 1903
* *[[Tachys dives]]* Péringuey, 1926 c
* *[[Tachys dominicanus]]* Darlington, 1934 c g
* *[[Tachys dorsalis]]* Motschulsky, 1851 c g
* *[[Tachys dostali]]* Baehr, 2014 c g
* *[[Tachys dromioides]]* Bates, 1871 c g
* *[[Tachys dzosonicus]]* Pawlowski, 1974 c g
* *[[Tachys ectromioides]]* Sloane, 1896 c g
* *[[Tachys edax]]* Leconte, 1852
* *[[Tachys ellenbergeri]]* (Bruneau de Miré, 1964) c g
* *[[Tachys elongatulus]]* (Dejean, 1831) c g
* *[[Tachys enormis]]* (Dostal, 2015) c g
* *[[Tachys erwini]]* Reichardt, 1976 c g
* *[[Tachys euryodes]]* Bates, 1892 c g
* *[[Tachys excisicollis]]* Baehr, 2012 c g
* *[[Tachys exochrias]]* Darlington, 1962 c g
* *[[Tachys fasciatus]]* (Motschulsky, 1851) c g
* *[[Tachys federicae]]* Baehr, 2016 c g
* *[[Tachys filax]]* Darlington, 1934 c g
* *[[Tachys flavax]]* Darlington, 1962 c g
* *[[Tachys flavicollis]]* Motschulsky, 1862 c g
* *[[Tachys fortestriatus]]* Baehr, 2003 c g
* *[[Tachys fortunatus]]* Machado, 1989 c g
* *[[Tachys frischi]]* Coulon & Felix, 2009
* *[[Tachys fulvicollis]]* (Dejean, 1831) c g
* *[[Tachys fumax]]* Darlington, 1962 c g
* *[[Tachys gilvus]]* Schaum, 1863 c g
* *[[Tachys gracchus]]* Andrewes, 1935 c g
* *[[Tachys gratus]]* Péringuey, 1896 c
* *[[Tachys guangxicus]]* (Deuve & Tian, 2010) c g
* *[[Tachys gyotokuensis]]* Tanaka, 1956 c g
* *[[Tachys halophilus]]* Lindroth, 1966 i c g
* *[[Tachys haywardi]]* Wickham, 1913 c g
* *[[Tachys holmi]]* (Basilewsky, 1963) c g
* *[[Tachys hyalinus]]* Casey, 1918 i c g
* *[[Tachys hydrophilus]]* (Germain, 1906) c g
* *[[Tachys impictus]]* Andrewes, 1925 c g
* *[[Tachys impressipennis]]* Motschulsky, 1859 c g
* *[[Tachys impressus]]* Motschulsky, 1851 c g
* *[[Tachys incertus]]* Andrewes, 1925 c g
* *[[Tachys infuscatus]]* Blackburn, 1888 c g
* *[[Tachys intermedius]]* Baehr, 2012 c g
* *[[Tachys iridipennis]]* Chaudoir, 1876 c g
* *[[Tachys jeannei]]* Coulon, 2004 c g
* *[[Tachys joannae]]* (Basilewsky, 1962) c g
* *[[Tachys jucundulus]]* Péringuey, 1908 c g
* *[[Tachys kabylianus]]* Puel, 1935 c g
* *[[Tachys kahuzianus]]* (Basilewsky, 1953) c g
* *[[Tachys kaorutanakai]]* Habu, 1977 c g
* *[[Tachys koizumii]]* Habu, 1961 c g
* *[[Tachys koreanorum]]* Pawlowski, 1974 c g
* *[[Tachys laevigatus]]* (Boheman, 1858) c g
* *[[Tachys laevus]]* (Say, 1823) c g
* *[[Tachys lamottei]]* (Bruneau de Miré, 1964) c g
* *[[Tachys languidus]]* Andrewes, 1925 c g
* *[[Tachys latalatus]]* Csiki, 1928 c g
* *[[Tachys leleupi]]* (Basilewsky, 1953) c g
* *[[Tachys lenkoranus]]* Csiki, 1928 c g
* *[[Tachys leptocerus]]* Chaudoir, 1876 c g
* *[[Tachys leytensis]]* (Baehr, 2016) c g
* *[[Tachys limbatellus]]* Bates, 1884 c g
* *[[Tachys lindi]]* Blackburn, 1888 c g
* *[[Tachys lissonotus]]* Andrewes, 1925 c g
* *[[Tachys litoralis]]* Casey, 1884 i c g
* *[[Tachys lividus]]* (Bates, 1871) c g
* *[[Tachys longulus]]* Andrewes, 1925 c g
* *[[Tachys lugubris]]* Motschulsky, 1862 c g
* *[[Tachys lusciosus]]* Antoine, 1943
* *[[Tachys luscus]]* Darlington, 1962 c g
* *[[Tachys macleayi]]* Sloane, 1896 c g
* *[[Tachys macrops]]* Baehr, 2012 c g
* *[[Tachys mameti]]* Alluaud, 1933 c g
* *[[Tachys marggii]]* Kirschenhofer, 1986 c g
* *[[Tachys marri]]* Baehr, 1987 c g
* *[[Tachys masculus]]* Darlington, 1962 c g
* *[[Tachys massauxi]]* (Basilewsky, 1955) c g
* *[[Tachys mastersi]]* Sloane, 1921 c g
* *[[Tachys micros]]* (Fischer von Waldheim, 1828) c g
* *[[Tachys microscopicus]]* (Bates, 1873) c g
* *[[Tachys minusculus]]* (Motschulsky, 1862)
* *[[Tachys minutissimus]]* (R.F.Sahlberg, 1847) c
* *[[Tachys mirandus]]* Coulon & Wrase, 2009
* *[[Tachys mirei]]* (Basilewsky, 1968) c g
* *[[Tachys misellus]]* Laferte-Senectere, 1841
* *[[Tachys mitchelli]]* Sloane, 1895 c g
* *[[Tachys moira]]* (Erwin, 1984)
* *[[Tachys monostictus]]* Bates, 1871 c g
* *[[Tachys mordax]]* Leconte, 1852
* *[[Tachys mulwalensis]]* Sloane, 1900 c g
* *[[Tachys mus]]* Andrewes, 1925 c g
* *[[Tachys myrmecophilus]]* (Jeannel, 1946) c g
* *[[Tachys nanniscus]]* Péringuey, 1896 c g
* *[[Tachys naraensis]]* Ueno, 1953 c g
* *[[Tachys nephelodes]]* Andrewes, 1930 c g
* *[[Tachys niger]]* (Jeannel, 1953) c g
* *[[Tachys nitmiluki]]* (Giachino, 2003) c g
* *[[Tachys nympha]]* (Erwin, 1984)
* *[[Tachys oahuensis]]* Blackburn, 1878 i c g
* *[[Tachys oblitus]]* Casey, 1918 i c g b
* *[[Tachys obtusiusculus]]* (Jeannel, 1941) c g
* *[[Tachys ochrias]]* Andrewes, 1925 c g
* *[[Tachys ochrioides]]* Darlington, 1962 c g
* *[[Tachys olemartini]]* Kirschenhofer, 1986 c g
* *[[Tachys opalescens]]* Andrewes, 1925 c g
* *[[Tachys orphninus]]* Andrewes, 1925 c g
* *[[Tachys otini]]* (Antoine, 1944)
* *[[Tachys pallidus]]* Chaudoir, 1868 i c g b
* *[[Tachys pallorus]]* Kopecky in Löbl & Smetana, 2003 c g
* *[[Tachys palustris]]* Reitter, 1894
* *[[Tachys panamensis]]* Casey, 1918 c g
* *[[Tachys particula]]* Andrewes, 1936 c g
* *[[Tachys paulax]]* Darlington, 1934 c g
* *[[Tachys pharao]]* Schatzmayr & Koch, 1934 c g
* *[[Tachys piceolus]]* Laferte-Senectere, 1841
* *[[Tachys plagiatus]]* Putzeys, 1875 c g
* *[[Tachys pliginskii]]* Solodovnikov, 2001 c g
* *[[Tachys politissimus]]* Baehr, 2014 c g
* *[[Tachys potomaca]]* (Erwin, 1981) i b
* *[[Tachys potomacus]]* (Erwin, 1981) c g
* *[[Tachys prionotus]]* Andrewes, 1936 c g
* *[[Tachys privus]]* Darlington, 1962 c g
* *[[Tachys prolixus]]* Bates, 1892 c g
* *[[Tachys proximus]]* Say, 1823
* *[[Tachys pseudolusciosus]]* Coulon, 2004 c g
* *[[Tachys pseudosericeus]]* Kirschenhofer, 1986 c g
* *[[Tachys pujoli]]* (Bruneau de Miré, 1964) c g
* *[[Tachys pulchellus]]* Laferte-Senectere, 1841
* *[[Tachys pumilus]]* (Dejean, 1831) i c g b
* *[[Tachys pusillimus]]* Péringuey, 1896 c g
* *[[Tachys quadrillum]]* Schaum, 1860
* *[[Tachys queenslandicus]]* Sloane, 1903 c g
* *[[Tachys rambai]]* Jedlicka, 1966 c g
* *[[Tachys rapoporti]]* (Jeannel, 1962)
* *[[Tachys rectangulus]]* Notman, 1919 i c g
* *[[Tachys rhodeanus]]* Casey, 1918 i c g b
* *[[Tachys ridiculus]]* (L.Schaufuss, 1879)
* *[[Tachys riedeli]]* Baehr, 2014 c g
* *[[Tachys rufotestaceus]]* Hayward, 1900
* *[[Tachys sagax]]* Casey, 1918 i c g
* *[[Tachys schuberti]]* Jedlicka, 1968 c g
* *[[Tachys scitulus]]* Leconte, 1848 i c g b
* *[[Tachys scutellaris]]* Stephens, 1828 c g
* *[[Tachys sellatus]]* Fairmaire, 1892 c g
* *[[Tachys sequax]]* Leconte, 1848
* *[[Tachys sericans]]* Bates, 1873 c g
* *[[Tachys sericeus]]* Motschulsky, 1851 c g
* *[[Tachys serra]]* Darlington, 1962 c g
* *[[Tachys serratus]]* Andrewes, 1925 c g
* *[[Tachys serrula]]* Darlington, 1962 c g
* *[[Tachys sexguttatus]]* Fairmaire, 1849
* *[[Tachys shahinei]]* Schatzmayr & Koch, 1934 c g
* *[[Tachys shirazi]]* Jedlicka, 1968 c g
* *[[Tachys sibling]]* Darlington, 1962 c g
* *[[Tachys similis]]* Blackburn, 1888 c g
* *[[Tachys simulator]]* Coulon, 2004 c g
* *[[Tachys spadix]]* Casey, 1918 i c g
* *[[Tachys striax]]* Darlington, 1934 c g
* *[[Tachys subangulatus]]* Bates, 1871 c g
* *[[Tachys subbrunneus]]* Darlington, 1962 c g
* *[[Tachys sublobatus]]* Darlington, 1962 c g
* *[[Tachys suboculatus]]* (Basilewsky, 1958) c g
* *[[Tachys sudanicus]]* (Basilewsky, 1948) c g
* *[[Tachys suensoni]]* Kirschenhofer, 1986 c g
* *[[Tachys sundaicus]]* Andrewes, 1925 c g
* *[[Tachys suturifer]]* Reitter, 1884 c g
* *[[Tachys sydneyensis]]* Sloane, 1923 c g
* *[[Tachys szekessyi]]* Jedlicka, 1968 c g
* *[[Tachys tantillus]]* Motschulsky, 1862
* *[[Tachys tenuiserra]]* Darlington, 1962 c g
* *[[Tachys terra]]* (Erwin, 1984)
* *[[Tachys testaceus]]* (Basilewsky, 1953) c
* *[[Tachys tetraphacus]]* Bedel, 1896 c g
* *[[Tachys tienmushaniensis]]* Kirschenhofer, 1986 c g
* *[[Tachys torretassoi]]* Schatzmayr & Koch, 1934 c g
* *[[Tachys transcaucasicus]]* Coulon, 2004 c g
* *[[Tachys translucens]]* Darlington, 1937 i c g
* *[[Tachys transumbratus]]* Bates, 1892 c g
* *[[Tachys transversicollis]]* (W.J.Macleay, 1871)
* *[[Tachys trechoderus]]* Coulon, 2004 c g
* *[[Tachys trechulus]]* Darlington, 1936 c g
* *[[Tachys troglocola]]* (Deuve & Tian, 2010) c g
* *[[Tachys troglophilus]]* Ueno, 1953 c g
* *[[Tachys tropicalis]]* Baehr, 2012 c g
* *[[Tachys tropicus]]* (Nietner, 1858) c g
* *[[Tachys truncatus]]* (Nietner, 1858) c g
* *[[Tachys turkestanicus]]* Csiki, 1928
* *[[Tachys ucayali]]* (Erwin, 1984)
* *[[Tachys uelensis]]* Burgeon, 1935 c g
* *[[Tachys uenoianus]]* Habu, 1974 c g
* *[[Tachys umbripennis]]* Chaudoir, 1868 i c g
* *[[Tachys uniformis]]* Blackburn, 1888 c g
* *[[Tachys vandeli]]* (Mateu & Colas, 1954) c g
* *[[Tachys vandepolli]]* (Bruneau de Miré, 1964) c g
* *[[Tachys varsavianorum]]* Pawlowski, 1974 c g
* *[[Tachys ventricosus]]* Leconte, 1863
* *[[Tachys venustus]]* Andrewes, 1936 c g
* *[[Tachys vernilis]]* Casey, 1918 i c g
* *[[Tachys vibex]]* Kopecky in Löbl & Smetana, 2003 c g
* *[[Tachys vietnami]]* Jedlicka, 1968 c g
* *[[Tachys vilis]]* Andrewes, 1925 c g
* *[[Tachys virgo]]* Leconte, 1852
* *[[Tachys vittiger]]* Leconte, 1852
* *[[Tachys vorax]]* Leconte, 1852
* *[[Tachys windsorensis]]* Baehr, 1991 c g
* *[[Tachys xanthochrous]]* Chaudoir, 1876 c g
* *[[Tachys yeboensis]]* Burgeon, 1935 c g
* *[[Tachys yodai]]* Jedlicka, 1964 c g
* *[[Tachys yorkensis]]* Baehr, 2012 c g
* *[[Tachys yunchengensis]]* Kirschenhofer, 1986 c g
* *[[Tachys zonatus]]* Andrewes, 1925 c g
* *[[Tachys zulficari]]* Schatzmayr & Koch, 1934 c g

## Litteratur

## Eksterne lenker
* 
*Bilder: