Glyphonyx

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***Glyphonyx*** er en slekt av biller som hører til familien smellere (Elateridae).

## Utseende
Små (gjerne 3 - 4 millimeter), smale, blanke, brunlige smellere. Antennene er tynne, pronotum hvelvet med rundede sider. Oversiden er fint hårete.

**Larvene** er sylindriske, glatte med en voksaktige overflate og spredte hår, gjerne rød- eller gulaktige. Hodet er noe flatt med fremoverstrakte kjever. Det bakerste leddet har vanligvis et par kraftige utvekster av varierende form. Beina er ganske korte.

## Levevis
Storsmellernes larver lever for det meste i morken, død ved. Hva de spiser kan variere – noen arter synes å være hovedsakelig planteetere, andre er rovdyr. Mange arter bruker flere år på larveutviklingen. De voksne billene flyr godt og treffes ofte flygende, på blomster eller i vegetasjonen. I likhet med mange andre arter knyttet til død ved er mange av storsmellerne sjeldne og truede arter.
Gruppen Agriotini har larver som lever i jorda og noen kan gjøre betydelig skade på planterøtter.

## Systematisk inndeling

 

* Ordenen biller, Coleoptera
** Underordenen Polyphaga
*** Overfamilien smellere, Elateroidea
**** Familien smellere, Elateridae
***** Underfamilien storsmellere, Elaterinae Leach, 1815
****** Stammen Adrastini Candéze, 1863 - noen ganger regnet til underfamilien Melanotinae
******* Slekten ***Glyphonyx*** Candéze, 1863 
******** *[[Glyphonyx aberdarensis]]* Fleutiaux
******** *[[Glyphonyx aberrans]]* Candèze
******** *[[Glyphonyx accedens]]* Candèze
******** *[[Glyphonyx acristatus]]* Vats & Chauhan, 1992
******** *[[Glyphonyx acuminatus]]* Fleutiaux, 1942
******** *[[Glyphonyx africanus]]* Schwarz, 1896
******** *[[Glyphonyx ami]]* Kishii, 1991
******** *[[Glyphonyx angulatus]]* Champion, 1896
******** *[[Glyphonyx antennalis]]* Fleutiaux, 1918
******** *[[Glyphonyx antiquus]]* Candèze, 1881
******** *[[Glyphonyx apayao]]* Kishii, 1991
******** *[[Glyphonyx arisanus]]* Miwa, 1931
******** *[[Glyphonyx atayal]]* Kishii, 1991
******** *[[Glyphonyx attonitus]]* Candèze, 1897
******** *[[Glyphonyx bauensis]]* Ôhira, 1973
******** *[[Glyphonyx bicolor]]* Candèze, 1895
******** *[[Glyphonyx bimarginatus]]* Schaeffer, 1916
******** *[[Glyphonyx birmanicus]]* Fleutiaux, 1942
******** *[[Glyphonyx bivittatus]]* Candèze, 1863
******** *[[Glyphonyx brevicollis]]* Champion, 1896
******** *[[Glyphonyx brevis]]* Fleutiaux, 1940
******** *[[Glyphonyx brevistylus]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx brunneipennis]]* Miwa, 1931
******** *[[Glyphonyx cadierei]]* Fleutiaux, 1940
******** *[[Glyphonyx carinifrons]]* Candèze, 1893
******** *[[Glyphonyx castaneus]]* Miwa, 1931
******** *[[Glyphonyx championi]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx chiapasensis]]* Zaragoza Caballero, S, 1990
******** *[[Glyphonyx chipenensis]]* Kishii, 1994
******** *[[Glyphonyx cinctus]]* (Champion, 1896)
******** *[[Glyphonyx cordieri]]* Fleutiaux, 1940
******** *[[Glyphonyx cruciellus]]* Erichson, 1842
******** *[[Glyphonyx cuneatus]]* Champion, 1896
******** *[[Glyphonyx dalopioides]]* Nakane, 1959
******** *[[Glyphonyx depremerus]]* Vats & Chauhan, 1992
******** *[[Glyphonyx depressus]]* Champion, 1896
******** *[[Glyphonyx dimidiatus]]* Schwarz, 1902
******** *[[Glyphonyx dirrhagoides]]* Fleutiaux, 1940
******** *[[Glyphonyx dissimilis]]* Candèze, 1897
******** *[[Glyphonyx dorsalis]]* Candèze
******** *[[Glyphonyx drapetoides]]* Fleutiaux, 1940
******** *[[Glyphonyx dubius]]* Schaeffer, 1916
******** *[[Glyphonyx dugesi]]* Candèze, 1893
******** *[[Glyphonyx ebeninus]]* Champion, 1896
******** *[[Glyphonyx elegans]]* Fleutiaux, 1940
******** *[[Glyphonyx erraticus]]* Candèze, 1875
******** *[[Glyphonyx esakii]]* Miwa, 1934
******** *[[Glyphonyx exilis]]* Kishii, 1991
******** *[[Glyphonyx falsus]]* (Candèze, 1897)
******** *[[Glyphonyx fasciatus]]* Champion, 1896
******** *[[Glyphonyx feneus]]* Candèze, 1897
******** *[[Glyphonyx ferrugineus]]* Schwarz
******** *[[Glyphonyx ferruginosus]]* Schaeffer, 1916
******** *[[Glyphonyx flavicollis]]* Kishii, 1991
******** *[[Glyphonyx fleutiauxi]]* Ôhira, 1973
******** *[[Glyphonyx flohri]]* Champion, 1896
******** *[[Glyphonyx formosanus]]* Ôhira, 1972
******** *[[Glyphonyx frequens]]* Fleutiaux, 1942
******** *[[Glyphonyx frontalis]]* Candèze
******** *[[Glyphonyx fuscicollis]]* Kishii, 1991
******** *[[Glyphonyx fusculus]]* Erichson, 1842
******** *[[Glyphonyx gagates]]* Fleutiaux, 1940
******** *[[Glyphonyx gestroi]]* Candèze
******** *[[Glyphonyx godavariensis]]* Ôhira & Becker, 1972
******** *[[Glyphonyx gracilis]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx gregarius]]* Schwarz
******** *[[Glyphonyx grossus]]* Kishii, 1994
******** *[[Glyphonyx gundlachii]]* Candèze, 1863
******** *[[Glyphonyx gunonganus]]* Ôhira, 1973
******** *[[Glyphonyx haterumarum]]* Ôhira, 1968
******** *[[Glyphonyx helix]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx housaii]]* Kishii, 1991
******** *[[Glyphonyx ihai]]* Ôhira, 1968
******** *[[Glyphonyx illepidus]]* Candèze, 1873
******** *[[Glyphonyx inconsultus]]* Candèze, 1893
******** *[[Glyphonyx inquinatus]]* (Say, 1834)
******** *[[Glyphonyx intermedius]]* Fleutiaux, 1940
******** *[[Glyphonyx iriomotensis]]* Miwa, 1934
******** *[[Glyphonyx kankaui]]* Miwa, 1931
******** *[[Glyphonyx kintaroui]]* Kishii, 1991
******** *[[Glyphonyx kishiii]]* Ôhira, 1968
******** *[[Glyphonyx knulli]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx koshikiensis]]* Ôhira, 2003
******** *[[Glyphonyx kosnipata]]* Johnson, 2018
******** *[[Glyphonyx kulashi]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx kuni]]* Platia & Schimmel, 2007
******** *[[Glyphonyx kurosawai]]* Nakane & Kishii, 1958
******** *[[Glyphonyx laszlorum]]* Platia & Schimmel, 2007
******** *[[Glyphonyx laticollis]]* Champion, 1896
******** *[[Glyphonyx latiusculus]]* Schwarz
******** *[[Glyphonyx littoralis]]* Fleutiaux, 1902
******** *[[Glyphonyx liukuiensis]]* Kishii, 1989
******** *[[Glyphonyx longicornis]]* Kishii, 1989
******** *[[Glyphonyx longipennis]]* Ôhira, 1966
******** *[[Glyphonyx longulus]]* Miwa, 1930
******** *[[Glyphonyx maai]]* Ôhira, 1973
******** *[[Glyphonyx makiharai]]* Ôhira, 1973
******** *[[Glyphonyx malaisei]]* Fleutiaux, 1942
******** *[[Glyphonyx marginalis]]* Kishii, 1994
******** *[[Glyphonyx matobai]]* Kishii, 1974
******** *[[Glyphonyx mimeticus]]* Horn, 1874
******** *[[Glyphonyx minicollis]]* Ôhira, 1973
******** *[[Glyphonyx minimus]]* Champion, 1896
******** *[[Glyphonyx montanus]]* Fleutiaux, 1902
******** *[[Glyphonyx montivagus]]* Fleutiaux, 1942
******** *[[Glyphonyx multus]]* Fleutiaux, 1940
******** *[[Glyphonyx muneaka]]* Kishii, 1991
******** *[[Glyphonyx nanus]]* Fleutiaux, 1940
******** *[[Glyphonyx nanus]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx nepalensis]]* Ôhira & Becker, 1972
******** *[[Glyphonyx nigritus]]* Candèze, 1878
******** *[[Glyphonyx nigrus]]* Steinheil, 1875
******** *[[Glyphonyx nitidicollis]]* Kishii, 1991
******** *[[Glyphonyx nomurai]]* Ôhira, 1968
******** *[[Glyphonyx occidentalis]]* Németh & Platia, 2014
******** *[[Glyphonyx oblongus]]* Fleutiaux, 1940
******** *[[Glyphonyx oiwakensis]]* Kishii, 1991
******** *[[Glyphonyx okinawanus]]* Chûjô, 1959
******** *[[Glyphonyx paiwan]]* Kishii, 1991
******** *[[Glyphonyx pallens]]* Fleutiaux, 1940
******** *[[Glyphonyx pallidipes]]* Miwa, 1934
******** *[[Glyphonyx pallidulus]]* Candèze, 1863
******** *[[Glyphonyx pallipes]]* Fleutiaux, 1940
******** *[[Glyphonyx parallelaris]]* Kishii, 1991
******** *[[Glyphonyx parallelus]]* Champion, 1896
******** *[[Glyphonyx perroti]]* Fleutiaux, 1940
******** *[[Glyphonyx posticus]]* Candèze, 1875
******** *[[Glyphonyx praecox]]* Erichson, 1842
******** *[[Glyphonyx praevius]]* Erichson, 1842
******** *[[Glyphonyx propinquus]]* Champion, 1896
******** *[[Glyphonyx punctatus]]* Becker, 1963
******** *[[Glyphonyx quadraticollis]]* Champion, 1896
******** *[[Glyphonyx quadricollis]]* Fleutiaux, 1940
******** *[[Glyphonyx quadrinotatus]]* Champion, 1896
******** *[[Glyphonyx quietus]]* (Say, 1839)
******** *[[Glyphonyx ranauensis]]* Ôhira, 1973
******** *[[Glyphonyx rasariti]]* Ôhira & Becker, 1972
******** *[[Glyphonyx recticollis]]* Say, 1823
******** *[[Glyphonyx recticollis]]* (Say, 1823)
******** *[[Glyphonyx rhopalacanthus]]* Smith & Balsbaugh, 1984
******** *[[Glyphonyx rubiginosus]]* Kishii, 1991
******** *[[Glyphonyx rubricollis]]* Miwa, 1928
******** *[[Glyphonyx rubricus]]* Candèze
******** *[[Glyphonyx ruficaudis]]* Candèze
******** *[[Glyphonyx rufithorax]]* Kishii, 1991
******** *[[Glyphonyx rugipennis]]* Champion, 1896
******** *[[Glyphonyx sarikeiensis]]* Ôhira, 1973
******** *[[Glyphonyx sauteri]]* Miwa, 1931
******** *[[Glyphonyx scabiusculus]]* Chevrolat, 1867
******** *[[Glyphonyx scutellecrista]]* Vats & Chauhan, 1992
******** *[[Glyphonyx shibatai]]* Ôhira, 1968
******** *[[Glyphonyx shirouzuanus]]* Kishii, 1991
******** *[[Glyphonyx similis]]* Fleutiaux, 1940
******** *[[Glyphonyx striatus]]* Schwarz, 1902
******** *[[Glyphonyx subopacus]]* Candèze
******** *[[Glyphonyx superbus]]* (Zayas, 1988)
******** *[[Glyphonyx suturalis]]* Candèze, 1902
******** *[[Glyphonyx taiwan]]* Kishii, 1991
******** *[[Glyphonyx taiwanus]]* Kishii, 1991
******** *[[Glyphonyx takasago]]* Kishii, 1989
******** *[[Glyphonyx tamurai]]* Kishii, 1985
******** *[[Glyphonyx teapensis]]* Champion, 1896
******** *[[Glyphonyx tenompokensis]]* Ôhira, 1973
******** *[[Glyphonyx tenuis]]* Champion, 1896
******** *[[Glyphonyx testaceus]]* (Melsheimer, 1845)
******** *[[Glyphonyx thoracicus]]* Fleutiaux, 1918
******** *[[Glyphonyx tonkinensis]]* Fleutiaux, 1940
******** *[[Glyphonyx truncatus]]* Fleutiaux, 1942
******** *[[Glyphonyx uedai]]* Kishii, 1977
******** *[[Glyphonyx variatus]]* Candèze, 1894
******** *[[Glyphonyx vittatus]]* Champion, 1896
******** *[[Glyphonyx vunun]]* Kishii, 1991
******** *[[Glyphonyx yona]]* Kishii, 1996
******** *[[Glyphonyx yonaguni]]* Kishii, 1974
******** *[[Glyphonyx yoshimotoi]]* Ôhira, 1971
******** *[[Glyphonyx yuwancola]]* Ôhira, 1971
******** *[[Glyphonyx zonatus]]* Candèze, 1878

## Eksterne lenker
* bugguide.net - bildedatabase over nordamerikanske insekter. Besøkt 15.11. 2019
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