Somaticus

***Somaticus*** er en slekt av skyggebiller der de fleste av artene er knyttet til tørre miljøer.

## Utseende
Middelsstore til ganske store, gjerne brede og høyt hvelvede, svært harde, ofte matt gråaktige til svarte skyggebiller. Antennene er trådformede og forholdsvis korte. Dekkvingene er gjerne høyt hvelvede med punktstriper. De fleste artene mangler flygevinger. Gruppen omfatter de største av alle skyggebiller, opptil 80 millimeter lange.

## Levevis
Nesten alle artene er nattaktive. Kroppsoverflaten viser noen tilpasninger til å ta opp fuktighet fra omgivelsene. Larvene til noen arter kan gjøre noe skade på dyrkede planter.

## Utbredelse
De fleste av artene lever i tørre områder i Afrika, mest i den sørlige delen.

## Systematisk inndeling

* Ordenen biller, Coleoptera
** Underordenen Polyphaga
*** Gruppen (infraorden) Cucujiformia
**** Overfamilien heteromerer, Tenebrionoidea Latreille, 1802
***** Familien **[[skyggebiller]]**, Tenebrionidae Latreille, 1802
****** Underfamilien **[[Pimeliinae]]** 
******* Stammen **[[Sepidiini]]** Eschscholtz, 1829
******** Understammen Trachynotina Koch, 1955
********* Slekten ***Somaticus*** Hope, 1840
********** Underslekten *Acromaticus* Koch, 1955
*********** *[[Somaticus adventitus]]* (Péringuey, 1899)
*********** *[[Somaticus albanyensis]]* Koch, 1955
*********** *[[Somaticus algoensis]]* Koch, 1955
*********** *[[Somaticus bisinuatus]]* Koch, 1955
*********** *[[Somaticus caviventris]]* Koch, 1955
*********** *[[Somaticus cohaerens]]* Koch, 1955
*********** *[[Somaticus corallipes]]* Koch, 1955
*********** *[[Somaticus dimorphus]]* Koch, 1955
*********** *[[Somaticus georgensis]]* Koch, 1955
*********** *[[Somaticus hirundo]]* Koch, 1955
*********** *[[Somaticus karrooensis]]* Koch, 1955
*********** *[[Somaticus licinoides]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus malaisei]]* Koch, 1955
*********** *[[Somaticus marlothi]]* Koch, 1955
*********** *[[Somaticus moerens]]* (Haag-Rutenberg, 1879)
*********** *[[Somaticus namaquensis]]* Koch, 1955
*********** *[[Somaticus nitens]]* (Péringuey, 1899)
*********** *[[Somaticus nollothensis]]* Koch, 1955
*********** *[[Somaticus praephallatus]]* Koch, 1955
*********** *[[Somaticus punctiger]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus purcelli]]* Koch, 1955
*********** *[[Somaticus saxicola]]* Koch, 1955
*********** *[[Somaticus scaber]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus striatus]]* (Thunberg, 1787)
*********** *[[Somaticus suturalifer]]* Koch, 1955
*********** *[[Somaticus transmontanus]]* Koch, 1955
*********** *[[Somaticus vansonianus]]* Koch, 1955
********** Underslekten *Bechuanitis* Koch, 1955
*********** *[[Somaticus bohemani]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus brucki]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus cinctus]]* Haag-Rutenberg, 1873
*********** *[[Somaticus geniculatus]]* Haag-Rutenberg, 1873
*********** *[[Somaticus hereroensis]]* Koch, 1955
*********** *[[Somaticus rugulosicollis]]* (Hesse, 1935)
********** Underslekten *Ceromelaephus* Koch, 1955
*********** *[[Somaticus badeni]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus seminitidus]]* Koch, 1955
*********** *[[Somaticus strangulatus]]* (Gebien, 1920)
*********** *[[Somaticus wahlbergi]]* Haag-Rutenberg, 1873
********** Underslekten *Clinocranion* Solier, 1844
*********** *[[Somaticus planatus]]* (Solier, 1843)
*********** *[[Somaticus spinosus]]* Solier, 1843
********** Underslekten *Diacis* Koch, 1955
*********** *[[Somaticus distinctus]]* Peringuey, 1892
*********** *[[Somaticus regalis]]* (Haag-Rutenberg, 1875)
********** Underslekten *Somaticus* (i snever forstand)
*********** *[[Somaticus aeneus]]* Solier, 1843
*********** *[[Somaticus bisbicostatus]]* (Gebien, 1920)
*********** *[[Somaticus decoratipes]]* Koch, 1955
*********** *[[Somaticus glabriventris]]* Koch, 1955
*********** *[[Somaticus rugosus]]* (Fabricius, 1781)
*********** *[[Somaticus stali]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus straminicornis]]* Koch, 1955
*********** *[[Somaticus stridulatus]]* Koch, 1955
*********** *[[Somaticus welwitschi]]* Koch, 1955
********** Underslekten *Tracheloeum* Hope, 1840
*********** *[[Somaticus carinatus]]* (Solier, 1843)
*********** *[[Somaticus contractus]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus dilatatus]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus fahraeusi]]* Koch, 1955
*********** *[[Somaticus giganteus]]* Koch, 1955
*********** *[[Somaticus hoffmanni]]* (Haag-Rutenberg, 1878)
*********** *[[Somaticus impressicollis]]* (Péringuey, 1885)
*********** *[[Somaticus intermedius]]* (Haag-Rutenberg, 1878)
*********** *[[Somaticus laticollis]]*  (Hope, 1840)
*********** *[[Somaticus maculosus]]* (Fåhraeus, 1870)
*********** *[[Somaticus marginatum]]* (Thunberg, 1787)
*********** *[[Somaticus pretorianus]]* Koch, 1955
*********** *[[Somaticus silphoides]]* (Fåhraeus, 1870)
*********** *[[Somaticus similis]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus transvaalensis]]* Koch, 1955
*********** *[[Somaticus vittiger]]* (Haag-Rutenberg, 1873)
********** Underslekten *Trachyderes* Koch, 1955
*********** *[[Somaticus albomaculatus]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus barnardi]]* Koch, 1955
*********** *[[Somaticus bipunctatus]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus braunsi]]* Koch, 1955
*********** *[[Somaticus cordipennis]]* Koch, 1955
*********** *[[Somaticus dubius]]* (Péringuey, 1885)
*********** *[[Somaticus dutoiti]]* Koch, 1955
*********** *[[Somaticus eremicola]]* Koch, 1955
*********** *[[Somaticus fitzsimonsi]]* Koch, 1955
*********** *[[Somaticus goryi]]* (Solier, 1843)
*********** *[[Somaticus gracilipes]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus gunvorae]]* (Koch, 1953)
*********** *[[Somaticus haagi]]* Peringuey, 1886
*********** *[[Somaticus incostatus]]* (Gebien, 1920)
*********** *[[Somaticus kungorum]]* Koch, 1955
*********** *[[Somaticus plutus]]* Koch, 1955
*********** *[[Somaticus pygmaeus]]* (Fåhraeus, 1870)
*********** *[[Somaticus ratus]]* Koch, 1955
*********** *[[Somaticus tentyrioides]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus tibialis]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus zinni]]* Koch, 1955
********** Underslekten *Trichotrachys* Koch, 1955
*********** *[[Somaticus angulatus]]* (Fåhraeus, 1873)
*********** *[[Somaticus darlingtoni]]* Koch, 1955
*********** *[[Somaticus funestus]]* (Fåhraeus, 1870)
*********** *[[Somaticus griseus]]* (Fåhraeus, 1870)
*********** *[[Somaticus hispidus]]* Hesse, 1935
*********** *[[Somaticus histrio]]* Koch, 1955
*********** *[[Somaticus lutulentus]]* (Péringuey, 1910)
*********** *[[Somaticus metropolis]]* Koch, 1955
*********** *[[Somaticus newtoni]]* Koch, 1955
*********** *[[Somaticus obscurus]]* Koch, 1955
*********** *[[Somaticus schalkwykae]]* Koch, 1955
*********** *[[Somaticus sinuatus]]* Koch, 1955
*********** *[[Somaticus sordidus]]* Gerstäcker, 1854
*********** *[[Somaticus terricola]]* (Fåhraeus, 1873)
*********** *[[Somaticus testudo]]* Koch, 1955
*********** *[[Somaticus varicollis]]* Koch, 1955
*********** *[[Somaticus vestitus]]* (Haag, 1910)
*********** *[[Somaticus zumpti]]* Koch, 1955
********** Underslekt *Trichotrichus* Koch, 1955
*********** *[[Somaticus crinitus]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus kraatzi]]* (Haag-Rutenberg, 1873)
*********** *[[Somaticus serratus]]* (Péringuey, 1885)
********** Underslekt *Tropitrachys* Koch, 1955
*********** *[[Somaticus peregrinator]]* (Koch, 1953)
*********** *[[Somaticus tropicalis]]* Koch, 1955
********** Usikker plassering
*********** *[[Somaticus angustus]]* (Péringuey, 1886)
*********** *[[Somaticus damarinus]]* (Peringuey, 1904)
*********** *[[Somaticus scutelliformis]]* (Laporte, 1840)

## Referanser




## Eksterne lenker
*