Hypothenemus

***Hypothenemus*** er en slekt av biller som tilhører gruppen barkbiller (Scolytinae) i den artsrike familien snutebiller (Curculionidae).

## Utseende
Ganske små, sylindriske, fint hårete barkbiller. 

## Levevis
Den best kjente arten i slekten er *Hypothenemus hampei*, som er et alvorlig skadedyr på kaffe over alt der denne blir dyrket. Hunnene legger egg på kaffibær, larvene utvikler seg i kaffibønnene i ødelegger disse med sine tunneler. Hannene i den neste generasjonen parrer seg med hunnene (vanligvis sine egne søstre) inne i kaffibæret og forlater det aldri, mens hunnene flyr ut og sprer seg til andre bær. Slike livssykler med stor grad av innavl er ikke uvanlig hos barkbiller.

## Utbredelse
Slekten er utbredt i varme deler av den Gamle verden, men *H. hampei* er spredt til nesten over alt der det blir dyrket kaffe.

## Systematisk inndeling

Etter en ny klassifikasjon  blir denne gruppen flyttet ut fra Cryphalini til sin egen stamme.

* Vingede insekter 
** Orden biller (Coleoptera Linnaeus, 1758)
*** Underorden Polyphaga Emery, 1886
*** Gruppe Cucujiformia Lameere, 1938
**** Overfamilie snute- og barkbiller (Curculionoidea Latreille, 1802)
***** Familien snutebiller (Curculionidae Latreille, 1802)
****** Underfamilie barkbiller (Scolytinae Latreille, 1804)
******* Stamme **[[Trypophloeini]]** Nüsslin, 1911
******** Slekten ***Hypothenemus*** Westwood, 1834 
### Liste over artene

* *[[Hypothenemus aberrans]]* Browne, 1973
* *[[Hypothenemus abhorrens]]* Wood, 2007
* *[[Hypothenemus abruptus]]* (Schedl, 1961)
* *[[Hypothenemus acaciae]]* (Eggers, 1920)
* *[[Hypothenemus adscitus]]* (Schedl, 1950)
* *[[Hypothenemus adustus]]* Bright, 2019
* *[[Hypothenemus aethiops]]* (Schedl, 1965)
* *[[Hypothenemus africanus]]* (Hopkins, 1915)
* *[[Hypothenemus agnatus]]* (Eggers, 1924)
* *[[Hypothenemus alternatus]]* (Eggers, 1943)
* *[[Hypothenemus amakusanus]]* (Murayama, 1934)
* *[[Hypothenemus amplissimus]]* Bright & Torres, 2006
* *[[Hypothenemus apicalis]]* Wood, 1974
* *[[Hypothenemus areccae]]* (Hornung, 1842)
* *[[Hypothenemus artocarpi]]* Browne, 1978
* *[[Hypothenemus arundinis]]* (Eichhoff, 1878)
* *[[Hypothenemus ascitus]]* Wood, 1971
* *[[Hypothenemus ater]]* Eggers, 1932
* *[[Hypothenemus aterrimulus]]* Wood, 1989
* *[[Hypothenemus aterrimus]]* (Schedl, 1951)
* *[[Hypothenemus atomus]]* Hopkins, 1915
* *[[Hypothenemus atratus]]* (Schedl, 1964)
* *[[Hypothenemus attenuatus]]* (Eggers, 1935)
* *[[Hypothenemus aulmanni]]* Hagedorn, 1912
* *[[Hypothenemus avitus]]* Bright & Poinar, 1994
* *[[Hypothenemus balachowskyi]]* Menier, 1971
* *[[Hypothenemus baloghi]]* (Schedl, 1967)
* *[[Hypothenemus bambusae]]* Browne, 1980
* *[[Hypothenemus barinensis]]* Wood, 2007
* *[[Hypothenemus bauhaniae]]* (Schedl, 1950)
* *[[Hypothenemus bezaziani]]* Peyerimhoff, 1935
* *[[Hypothenemus bicinctus]]* Schedl, 1959
* *[[Hypothenemus bidens]]* Browne, 1973
* *[[Hypothenemus bifurcatus]]* Bright, 2019
* *[[Hypothenemus birmanus]]* (Eichhoff, 1878)
* *[[Hypothenemus biseriatus]]* (Eggers, 1919)
* *[[Hypothenemus bolivianus]]* (Eggers, 1931)
* *[[Hypothenemus brevicollis]]* (Eggers, 1927)
* *[[Hypothenemus brevis]]* Eggers, 1932
* *[[Hypothenemus californicus]]* Hopkins, 1915
* *[[Hypothenemus camerunus]]* (Eggers, 1922)
* *[[Hypothenemus carbonarius]]* Eggers, 1943
* *[[Hypothenemus carinafrons]]* Bright, 2019
* *[[Hypothenemus colae]]* (Schedl, 1957)
* *[[Hypothenemus collinus]]* Bright, 2019
* *[[Hypothenemus columbi]]* Hopkins, 1915
* *[[Hypothenemus concolor]]* Hagedorn, 1909
* *[[Hypothenemus costatus]]* (Eichhoff, 1878)
* *[[Hypothenemus crinatus]]* Bright, 2019
* *[[Hypothenemus criticus]]* (Schedl, 1937)
* *[[Hypothenemus crudiae]]* (Panzer, 1791)
* *[[Hypothenemus cryphaloides]]* (Eichhoff, 1878)
* *[[Hypothenemus cuneolus]]* (Schedl, 1936)
* *[[Hypothenemus curtipennis]]* (Schedl, 1950)
* *[[Hypothenemus cylindraceus]]* Schedl, 1972
* *[[Hypothenemus cynometrae]]* Schedl, 1957
* *[[Hypothenemus delicatus]]* Schedl, 1964
* *[[Hypothenemus deprecator]]* (Schedl, 1941)
* *[[Hypothenemus dexter]]* (Sampson, 1922)
* *[[Hypothenemus dimorphus]]* (Schedl, 1959)
* *[[Hypothenemus dipterocarpi]]* Hopkins, 1915
* *[[Hypothenemus discordis]]* Bright, 2019
* *[[Hypothenemus dissimilis]]* (Zimmermann, 1868)
* *[[Hypothenemus distinctus]]* Wood, 1954
* *[[Hypothenemus dolichocola]]* Hopkins, 1915
* *[[Hypothenemus donisi]]* (Schedl, 1957)
* *[[Hypothenemus dorsosignatus]]* (Schedl, 1950)
* *[[Hypothenemus dubitalis]]* Bright, 2019
* *[[Hypothenemus ebenus]]* Wood, 2007
* *[[Hypothenemus elephas]]* (Eichhoff, 1872)
* *[[Hypothenemus emmi]]* (Hagedorn, 1913)
* *[[Hypothenemus erectus]]* LeConte, 1876
* *[[Hypothenemus eruditus]]* (Westwood, 1834)
* *[[Hypothenemus euphorbiae]]* (Schedl, 1961)
* *[[Hypothenemus exceptus]]* Bright, 2019
* *[[Hypothenemus exiguus]]* (Wood, 1986)
* *[[Hypothenemus eximius]]* Schedl, 1951
* *[[Hypothenemus externedentatus]]* Schedl, 1959
* *[[Hypothenemus flavus]]* Hopkins, 1915
* *[[Hypothenemus fuscicollis]]* (Eichhoff, 1878)
* *[[Hypothenemus georgiae]]* (Hopkins, 1915)
* *[[Hypothenemus glabratulus]]* (Schedl, 1957)
* *[[Hypothenemus glabripennis]]* (Hopkins, 1915)
* *[[Hypothenemus gossypii]]* (Hopkins, 1915)
* *[[Hypothenemus grandis]]* Schedl, 1939
* *[[Hypothenemus granulatus]]* Bright, 2019
* *[[Hypothenemus hampei]]* (Ferrari, 1867) - er funnet innført til Norge
* *[[Hypothenemus hirsutus]]* (Wood, 1954)
* *[[Hypothenemus hystrix]]* (Eggers, 1919)
* *[[Hypothenemus ignotus]]* Bright, 2019
* *[[Hypothenemus improvidus]]* Bright, 2019
* *[[Hypothenemus incognitus]]* (Schedl, 1967)
* *[[Hypothenemus indigens]]* Wood, 1974
* *[[Hypothenemus indigenus]]* Bright & Peck, 1998
* *[[Hypothenemus indistinctus]]* Bright, 2019
* *[[Hypothenemus ingens]]* Schedl, 1942
* *[[Hypothenemus inordinatus]]* Bright, 2019
* *[[Hypothenemus insulanus]]* Bright, 2002
* *[[Hypothenemus interstitialis]]* (Hopkins, 1915)
* *[[Hypothenemus intricatus]]* (Schedl, 1950)
* *[[Hypothenemus japonicus]]* (Niisima, 1910)
* *[[Hypothenemus javanus]]* (Eggers, 1908)
* *[[Hypothenemus kamathi]]* Beaver, 1995
* *[[Hypothenemus kraunhiae]]* Murayama, 1950
* *[[Hypothenemus lefevrei]]* (Schedl, 1952)
* *[[Hypothenemus leprieuri]]* (Perris, 1866)
* *[[Hypothenemus leptosquamus]]* Bright, 2019
* *[[Hypothenemus liberiensis]]* (Hopkins, 1915)
* *[[Hypothenemus liliputianus]]* Bright, 2019
* *[[Hypothenemus lineatus]]* (Eggers, 1927)
* *[[Hypothenemus longipennis]]* (Eggers, 1935)
* *[[Hypothenemus longipilis]]* Schedl, 1952
* *[[Hypothenemus loranthus]]* (Schedl, 1942)
* *[[Hypothenemus macrolobii]]* (Eggers, 1940)
* *[[Hypothenemus madagascariensis]]* Schedl, 1953
* *[[Hypothenemus magnus]]* (Eggers, 1924)
* *[[Hypothenemus major]]* Browne, 1970
* *[[Hypothenemus malayensis]]* (Schedl, 1977)
* *[[Hypothenemus mallyi]]* (Hopkins, 1915)
* *[[Hypothenemus malus]]* (Schedl, 1957)
* *[[Hypothenemus mangovorus]]* Schedl, 1961
* *[[Hypothenemus marginatus]]* Johnson, 2020
* *[[Hypothenemus marshalli]]* (Eggers, 1936)
* *[[Hypothenemus mateui]]* (Schedl, 1965)
* *[[Hypothenemus melanarius]]* (Schedl, 1953)
* *[[Hypothenemus melasomus]]* (Lea, 1910)
* *[[Hypothenemus meridensis]]* Wood, 2007
* *[[Hypothenemus miles]]* (LeConte, 1878)
* *[[Hypothenemus minor]]* (Eggers, 1927)
* *[[Hypothenemus modestus]]* (Murayama, 1940)
* *[[Hypothenemus morigerus]]* (Schedl, 1957)
* *[[Hypothenemus morio]]* (Eggers, 1940)
* *[[Hypothenemus morosus]]* Schedl, 1965
* *[[Hypothenemus mozambiquensis]]* Eggers, 1943
* *[[Hypothenemus mulongensis]]* (Eggers, 1940)
* *[[Hypothenemus multidentatulus]]* (Schedl, 1962)
* *[[Hypothenemus multidentatus]]* (Hopkins, 1915)
* *[[Hypothenemus multipunctatus]]* (Schedl, 1939)
* *[[Hypothenemus muticus]]* (Schedl, 1961)
* *[[Hypothenemus namosianus]]* Browne, 1983
* *[[Hypothenemus nanellus]]* Wood, 1971
* *[[Hypothenemus nanoparvus]]* Bright, 2019
* *[[Hypothenemus natalensis]]* (Schedl, 1941)
* *[[Hypothenemus nesiotus]]* Bright, 2019
* *[[Hypothenemus nigropiceus]]* (Schedl, 1951)
* *[[Hypothenemus novateutonicus]]* (Schedl, 1951)
* *[[Hypothenemus obscurifrons]]* Bright, 2019
* *[[Hypothenemus obscurus]]* (Fabricius, 1801)
* *[[Hypothenemus opacus]]* (Eichhoff, 1872)
* *[[Hypothenemus paradoxus]]* Schedl, 1964
* *[[Hypothenemus parallelus]]* (Hopkins, 1915)
* *[[Hypothenemus parasquamosus]]* Bright, 2019
* *[[Hypothenemus parvulosus]]* Bright, 2019
* *[[Hypothenemus parvulus]]* Browne, 1984
* *[[Hypothenemus paulus]]* Bright, 2019
* *[[Hypothenemus perappositus]]* (Schedl, 1934)
* *[[Hypothenemus perexiguus]]* Bright, 2019
* *[[Hypothenemus perhispidus]]* (Eggers, 1927)
* *[[Hypothenemus perpunctatus]]* (Eggers, 1940)
* *[[Hypothenemus piaparolinae]]* Johnson, Atkinson & Hulcr, 2016
* *[[Hypothenemus pilosus]]* Hopkins, 1915
* *[[Hypothenemus ponticus]]* Bright, 2019
* *[[Hypothenemus praecellens]]* (Schedl, 1972)
* *[[Hypothenemus pubescens]]* Hopkins, 1915
* *[[Hypothenemus pubipennis]]* (Eggers, 1935)
* *[[Hypothenemus puertoricensis]]* (Bright & Torres, 2006)
* *[[Hypothenemus pullus]]* (Wood, 1971)
* *[[Hypothenemus pygmaeomorphus]]* Bright, 2019
* *[[Hypothenemus rotundicollis]]* (Eichhoff, 1878)
* *[[Hypothenemus rubrithorax]]* Bright, 2019
* *[[Hypothenemus ruficeps]]* Perkins, 1900
* *[[Hypothenemus rugifer]]* (Schedl, 1965)
* *[[Hypothenemus ruginosus]]* Wood, 1989
* *[[Hypothenemus rugosipes]]* Wood, 2007
* *[[Hypothenemus sambesianus]]* Eggers, 1943
* *[[Hypothenemus sapporoensis]]* (Niisima, 1910)
* *[[Hypothenemus sassaensis]]* (Eggers, 1924)
* *[[Hypothenemus schedli]]* Browne, 1963
* *[[Hypothenemus scutiae]]* (Schedl, 1959)
* *[[Hypothenemus seoulensis]]* Choo & Woo, 1989
* *[[Hypothenemus seriatus]]* (Eichhoff, 1872)
* *[[Hypothenemus setiferous]]* Bright, 2019
* *[[Hypothenemus setosus]]* (Eichhoff, 1868)
* *[[Hypothenemus simoni]]* (Reitter, 1887)
* *[[Hypothenemus sobrinus]]* (Schedl, 1965)
* *[[Hypothenemus socialis]]* (Schedl, 1957)
* *[[Hypothenemus solitarius]]* (Schedl, 1950)
* *[[Hypothenemus solivagus]]* Bright, 2019
* *[[Hypothenemus solocis]]* Wood, 1974
* *[[Hypothenemus sparsedentatus]]* (Schedl, 1942)
* *[[Hypothenemus sparsus]]* Hopkins, 1915
* *[[Hypothenemus spinatus]]* (Schedl, 1977)
* *[[Hypothenemus spinicollis]]* (Schedl, 1965)
* *[[Hypothenemus spinosus]]* (Schedl, 1979)
* *[[Hypothenemus squamosulus]]* Johnson, 2020
* *[[Hypothenemus squamosus]]* (Hopkins, 1915)
* *[[Hypothenemus stigmosus]]* (Schedl, 1951)
* *[[Hypothenemus striatus]]* (Atkinson, 1993)
* *[[Hypothenemus styrax]]* (Schedl, 1942)
* *[[Hypothenemus subacuminatus]]* (Schedl, 1942)
* *[[Hypothenemus subterrestris]]* Johnson, Atkinson & Hulcr, 2016
* *[[Hypothenemus suspectus]]* Wood, 1974
* *[[Hypothenemus tectus]]* Bright, 2019
* *[[Hypothenemus teretis]]* Wood, 1971
* *[[Hypothenemus teteforti]]* (Menier, 1971)
* *[[Hypothenemus tredli]]* (Reitter, 1908)
* *[[Hypothenemus tristis]]* (Eichhoff, 1876)
* *[[Hypothenemus trivialis]]* Wood, 1974
* *[[Hypothenemus tuberosus]]* (Schedl, 1942)
* *[[Hypothenemus turnbowi]]* Bright, 2019
* *[[Hypothenemus ustulatus]]* Bright, 2019
* *[[Hypothenemus vernaculus]]* Bright, 2019
* *[[Hypothenemus versicolor]]* Bright, 2019
* *[[Hypothenemus vesculus]]* Wood, 1974
* *[[Hypothenemus villosus]]* Bright, 2019
* *[[Hypothenemus virolae]]* Wood, 2007
* *[[Hypothenemus vitis]]* Browne, 1970
* *[[Hypothenemus winkleri]]* (Reitter, 1907)
* *[[Hypothenemus woodi]]* Bright, 2019
* *[[Hypothenemus xanthophloeae]]* (Schedl, 1957)

## Referanser




## Eksterne lenker
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