Cryptocephalus

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***Cryptocephalus*** er en slekt med biller som tilhører familiegruppen bladbiller (Chrysomelidae).  

## Utseende
*Cryptocephalus* er vanligvis mellom 2-10&nbsp;mm lange. De er korte, og butte både foran og bak. Det siste leddet av bakkroppen er synlig bak dekkvingene. Undersiden er noe «oppsvulmet». Antennene er uvanlig lange i forhold til hva som er vanlig for andre bladbiller. På farge er de mer eller mindre metallisk fargede, svarte, gulaktige eller rødlige. Pronotum er rundet, gjerne ikke mye kortere enn dekkvingene.

Mange av artene er svært like hverandre og for noen kan sikker artsbestemmelse bare skje ved undersøkelse av genitaliene. Opplysninger om næringsplanten kan være til nyttige for en sikrere artsbetemmelse.

## Levevis
Både larvene og de voksne (imago) billene lever av å spise blader på næringsplanten og er særlig aktive i varme og solskinn. Artene i denne slekten kan finnes på en rekke ulike plantearter, både løv- og bartrær og ulike slags urter.
De voksne kan også finnes på blomster.

## Systematisk inndeling med fokus på norske arter
Listen er ganske ufullstendig.
* orden **[[biller]]** (Coleoptera)
** underorden **[[Polyphaga]]**
*** overfamilie **[[Chrysomeloidea]]** Latreille, 1802 (=Phytophaga) 
**** familien  bladbiller (Chrysomelidae Latreille, 1802) 
***** underfamilien Hettebladbiller Cryptocephalinae  Gyllenhal, 1813
****** stamme Cryptocephalini
******* Slekten ***Cryptocephalus*** O.F. Müller, 1764  – 21 arter i Norge (+ 2 usikre)
******** *[[Cryptocephalus aureolus]]*
******** *[[Cryptocephalus biguttatus]]*
******** stripehettebladbille *Cryptocephalus bilineatus* (Linnaeus 1767) – finnes i Norge
******** toflekket hettebladbille *Cryptocephalus bipunctatus* (Linnaeus 1758) – finnes i Norge
******** hasselhettebladbille *Cryptocephalus coryli* (Linnaeus 1758) – finnes i Norge
******** myrhettebladbille *Cryptocephalus decemmaculatus* (Linnaeus 1758) – finnes i Norge
******** firflekket hettebladbille *Cryptocephalus distinguendus* Schneider 1792 – finnes i Norge
******** kurvhettebladbille *Cryptocephalus exiguus* Schneider 1792 – finnes i Norge
******** berghettebladbille *Cryptocephalus frontalis* Marsham 1802 – finnes i Norge
******** gulhettebladbille *Cryptocephalus fulvus* (Goeze 1777) – finnes i Norge
******** enghettebladbille *Cryptocephalus hypochoeridis* (Linnaeus 1758) – finnes i Norge
******** dverghettebladbille *Cryptocephalus labiatus* (Linnaeus 1761) – finnes i Norge
******** perikumhettebladbille *Cryptocephalus moraei* (Linnaeus 1758) – finnes i Norge
******** glanshettebladbille *Cryptocephalus nitidus* (Linnaeus 1758) – finnes i Norge
******** *[[Cryptocephalus nitidulus]]* Fabricius 1787 – finnes i Norge
******** blåhettebladbille *Cryptocephalus parvulus* O.F. Muller 1776 – finnes i Norge
******** Furuhettebladbille, *Cryptopcephalus pini* (Linnaeus, 1758) – finnes i Norge
******** *[[Cryptocephalus primarius]]*
******** bjørkehettebladbille *Cryptocephalus punctiger* Paykull 1799 – finnes i Norge
******** høsthettebladbille *Cryptocephalus pusillus* Fabricius 1777 – finnes i Norge
******** fireflekket furunålbille *Cryptocephalus quadripustulatus* Gyllenhal 1813 – finnes i Norge
******** eikehettebladbille *Cryptocephalus querceti* Suffrian 1848 – finnes i Norge
******** prakthettebladbille *Cryptocephalus sericeus* (Linnaeus 1758) – finnes i Norge
******** seksflekket hettebladbille *Cryptocephalus sexpunctatus* (Linnaeus 1758) – finnes i Norge

## Liste over artene
**[[Cryptocephalus abdominalis]]*
**[[Cryptocephalus acupunctatus]]*
**[[Cryptocephalus albicans]]* Haldeman, 1849
**[[Cryptocephalus albolineatus]]*
**[[Cryptocephalus alnicola]]*
**[[Cryptocephalus alternans]]* Suffrian, 1852
**[[Cryptocephalus amasiensis]]*
**[[Cryptocephalus amatus]]* Haldeman, 1849
**[[Cryptocephalus androgyne]]*
**[[Cryptocephalus angorensis]]*
**[[Cryptocephalus anticus]]*
**[[Cryptocephalus apicalis]]*
**[[Cryptocephalus aquitanus]]*
**[[Cryptocephalus araxidis]]*
**[[Cryptocephalus areolatus]]* Suffrian, 1852
**[[Cryptocephalus arizonensis]]* Schaeffer, 1904
**[[Cryptocephalus astracanicus]]*
**[[Cryptocephalus astralosus]]* White, 1968
**[[Cryptocephalus asturiensis]]*
**[[Cryptocephalus atrifrons]]*
**[[Cryptocephalus atriplicis]]*
**[[Cryptocephalus atrofasciatus]]* Jacoby, 1880
**[[Cryptocephalus augustalisi]]*
**[[Cryptocephalus aulicus]]* Haldeman, 1849
**[[Cryptocephalus aureolus]]*
**[[Cryptocephalus ayvazi]]*
**[[Cryptocephalus azurescens]]*
**[[Cryptocephalus baborensis]]*
**[[Cryptocephalus badius]]* Suffrian, 1852
**[[Cryptocephalus baeticus]]*
**[[Cryptocephalus bahilloi]]* José Ignacio López Colón,, 2003
**[[Cryptocephalus bailundensis]]* Weise
**[[Cryptocephalus bameuli]]*
**[[Cryptocephalus barii]]*
**[[Cryptocephalus basalis]]* Suffrian, 1852
**[[Cryptocephalus bicolor]]*
**[[Cryptocephalus bidorsalis]]*
**[[Cryptocephalus biguttatus]]*
**[[Cryptocephalus biledjekensis]]*
*stripehettebladbille *Cryptocephalus bilineatus*
**[[Cryptocephalus bimaculatithoracalis]]* Duan, Wang & Zhu, 2025
**[[Cryptocephalus bimaculatus]]*
**[[Cryptocephalus binominis]]* Newman, 1841
**[[Cryptocephalus binotatus]]* White, 1968
**[[Cryptocephalus biondii]]*
*toflekket hettebladbille *Cryptocephalus bipunctatus*
**[[Cryptocephalus bispinus]]* Suffrian, 1858
**[[Cryptocephalus bivius]]* Newman, 1840
**[[Cryptocephalus blanduloides]]*
**[[Cryptocephalus bohemius]]*
**[[Cryptocephalus borowieci]]*
**[[Cryptocephalus brevisignaticollis]]*
**[[Cryptocephalus brunneovittatus]]* Schaeffer, 1904
**[[Cryptocephalus calidus]]* Suffrian, 1852
**[[Cryptocephalus cantabricus]]*
**[[Cryptocephalus carinthiacus]]*
**[[Cryptocephalus carpathicus]]*
**[[Cryptocephalus castaneus]]* Leconte, 1880
**[[Cryptocephalus castilianus]]*
**[[Cryptocephalus celtibericus]]*
**[[Cryptocephalus cerinus]]* White, 1937
**[[Cryptocephalus chrysopus]]*
**[[Cryptocephalus cicatricosus]]*
**[[Cryptocephalus cognatus]]*
**[[Cryptocephalus concolor]]*
**[[Cryptocephalus confluentus]]* Say, 1824
**[[Cryptocephalus connexus]]*
**[[Cryptocephalus contextus]]* White, 1968
**[[Cryptocephalus convergens]]*
**[[Cryptocephalus cordiger]]*
**[[Cryptocephalus coronatus]]*
*hasselhettebladbille *Cryptocephalus coryli*
**[[Cryptocephalus corynetes]]*
**[[Cryptocephalus cowaniae]]* Schaeffer, 1933
**[[Cryptocephalus crassus]]*
**[[Cryptocephalus crenatus]]*
**[[Cryptocephalus creticus]]*
**[[Cryptocephalus cribratus]]*
**[[Cryptocephalus cribripennis]]* Leconte, 1880
**[[Cryptocephalus cristula]]*
**[[Cryptocephalus cuneatus]]* Fall, 1932
**[[Cryptocephalus cupressi]]* Schaeffer, 1933
**[[Cryptocephalus curda]]*
**[[Cryptocephalus curvilinea]]*
**[[Cryptocephalus cyanipes]]*
**[[Cryptocephalus cynarae]]*
**[[Cryptocephalus czwalinae]]*
**[[Cryptocephalus daccordii]]*
**[[Cryptocephalus dahdah]]*
**[[Cryptocephalus danieli]]*
*myrhettebladbille *Cryptocephalus decemmaculatus*
**[[Cryptocephalus defectus]]* Leconte, 1880
**[[Cryptocephalus dinae]]*
**[[Cryptocephalus discicollis]]*
**[[Cryptocephalus disruptus]]* White, 1968
**[[Cryptocephalus distinctenotatus]]*
*firflekket hettebladbille *Cryptocephalus distinguendus*
**[[Cryptocephalus dogueti]]*
**[[Cryptocephalus dorsatus]]* White, 1968
**[[Cryptocephalus dumonti]]*
**[[Cryptocephalus duplicatus]]*
**[[Cryptocephalus duryi]]* Schaeffer, 1906
**[[Cryptocephalus egregius]]* Schaeffer, 1933
**[[Cryptocephalus elatus]]*
**[[Cryptocephalus elegantulus]]*
**[[Cryptocephalus elongatus]]*
**[[Cryptocephalus emiliae]]*
**[[Cryptocephalus equiseti]]*
**[[Cryptocephalus ergenensis]]*
**[[Cryptocephalus eridani]]*
**[[Cryptocephalus espanoli]]*
**[[Cryptocephalus etruscus]]*
**[[Cryptocephalus euchirus]]*
**[[Cryptocephalus excisus]]*
*kurvhettebladbille *Cryptocephalus exiguus*
**[[Cryptocephalus falzonii]]*
**[[Cryptocephalus fasciatus]]* Say, 1824
**[[Cryptocephalus fausti]]*
**[[Cryptocephalus flavicinctus]]* Jacoby, 1892
**[[Cryptocephalus flavicollis]]*
**[[Cryptocephalus flavipes]]*
**[[Cryptocephalus flexuosus]]*
**[[Cryptocephalus floralis]]*
**[[Cryptocephalus floribundus]]*
**[[Cryptocephalus freidbergi]]*
**[[Cryptocephalus frenatus]]*
*berghettebladbille *Cryptocephalus frontalis*
**[[Cryptocephalus fulgurans]]*
**[[Cryptocephalus fulguratus]]* Leconte, 1880
*gulhettebladbille *Cryptocephalus fulvus*
**[[Cryptocephalus gamma]]*
**[[Cryptocephalus gibbicollis]]* Haldeman, 1849
**[[Cryptocephalus globicollis]]*
**[[Cryptocephalus gloriosus]]*
**[[Cryptocephalus gridellii]]*
**[[Cryptocephalus guttulatellus]]* Schaeffer, 1919
**[[Cryptocephalus guttulatus]]* Olivier, 1808
**[[Cryptocephalus heinigi]]*
**[[Cryptocephalus hypochaeridis]]* L.
**[[Cryptocephalus ilicis]]*
**[[Cryptocephalus illyricus]]*
**[[Cryptocephalus imperialis]]*
**[[Cryptocephalus implacidus]]* White, 1968
**[[Cryptocephalus incertus]]* Olivier, 1808
**[[Cryptocephalus infirmior]]*
**[[Cryptocephalus informis]]*
**[[Cryptocephalus ingamma]]*
**[[Cryptocephalus insertus]]* Haldeman, 1849
**[[Cryptocephalus janthinus]]*
**[[Cryptocephalus jocularius]]*
**[[Cryptocephalus katranus]]*
*dverghettebladbille *Cryptocephalus labiatus*
**[[Cryptocephalus laetus]]*
**[[Cryptocephalus laevicollis]]*
**[[Cryptocephalus langxianus]]* Duan, Wang & Zhu, 2025
**[[Cryptocephalus laosensis]]* Pic, 1928
**[[Cryptocephalus lateralis]]*
**[[Cryptocephalus lateritius]]* Newman, 1841
**[[Cryptocephalus lederi]]*
**[[Cryptocephalus leonhardi]]*
**[[Cryptocephalus leucomelas]]* Suffrian, 1852
**[[Cryptocephalus limbifer]]*
**[[Cryptocephalus limoniastri]]*
**[[Cryptocephalus lineellus]]*
**[[Cryptocephalus literatifer]]*
**[[Cryptocephalus lividimanus]]*
**[[Cryptocephalus loebli]]*
**[[Cryptocephalus longwangshani]]* Duan, Wang & Zhou, 2025
**[[Cryptocephalus loreyi]]*
**[[Cryptocephalus lostiai]]*
**[[Cryptocephalus lunatus]]* White, 1968
**[[Cryptocephalus luridicollis]]*
**[[Cryptocephalus lusitanicus]]*
**[[Cryptocephalus luteolus]]* Newman, 1840
**[[Cryptocephalus maccus]]* White, 1968
**[[Cryptocephalus macellus]]*
**[[Cryptocephalus majoricensis]]*
**[[Cryptocephalus marginatus]]*
*berghettebladbille *Cryptocephalus marginellus*
**[[Cryptocephalus mariae]]*
**[[Cryptocephalus mayeti]]*
**[[Cryptocephalus merus]]* Fall, 1932
**[[Cryptocephalus modestus]]*
**[[Cryptocephalus moehringi]]*
**[[Cryptocephalus monticola]]*
*perikumhettebladbille *Cryptocephalus moraei*
**[[Cryptocephalus moroderi]]*
**[[Cryptocephalus mucoreus]]* Leconte, 1859
**[[Cryptocephalus muellerianus]]*
**[[Cryptocephalus multisignatus]]* Schaeffer, 1933
**[[Cryptocephalus mutabilis]]* Melsheimer, 1847
**[[Cryptocephalus mystacatus]]*
**[[Cryptocephalus nanus]]* Fabricius, 1801
**[[Cryptocephalus nigellus]]*
**[[Cryptocephalus nitidicollis]]*
**[[Cryptocephalus nitidulus]]*
*glanshettebladbille *Cryptocephalus nitidus*
**[[Cryptocephalus notatus]]* Fabricius, 1787
**[[Cryptocephalus nubigena]]*
**[[Cryptocephalus numidicus]]*
**[[Cryptocephalus obsoletus]]* Germar, 1824
**[[Cryptocephalus ocellatus]]*
*gulhettebladbille *Cryptocephalus ochraceus* Fall, 1932 (Synonym for *Cryptocephalus fulvus*)
**[[Cryptocephalus ochroleucus]]*
**[[Cryptocephalus octacosmus]]*
**[[Cryptocephalus octoguttatus]]*
**[[Cryptocephalus octomaculatus]]*
**[[Cryptocephalus octopunctatus]]*
**[[Cryptocephalus oranensis]]*
**[[Cryptocephalus paganensis]]*
**[[Cryptocephalus pallidicinctus]]* Fall, 1932
**[[Cryptocephalus pallidocinctus]]*
**[[Cryptocephalus pallifrons]]*
**[[Cryptocephalus paphlagonius]]*
*blåhettebladbille *Cryptocephalus parvulus*
**[[Cryptocephalus pelleti]]*
**[[Cryptocephalus perrisi]]*
**[[Cryptocephalus pexicollis]]*
**[[Cryptocephalus peyroni]]*
**[[Cryptocephalus phaleratus]]*
**[[Cryptocephalus piceoverticalis]]*
*furuhettebladbille *Cryptocephalus pini*
**[[Cryptocephalus pinicola]]* Schaeffer, 1919
**[[Cryptocephalus planifrons]]*
**[[Cryptocephalus plantaris]]*
**[[Cryptocephalus podager]]*
**[[Cryptocephalus politus]]*
**[[Cryptocephalus pominorum]]*
**[[Cryptocephalus populi]]*
**[[Cryptocephalus praticola]]*
**[[Cryptocephalus primarius]]*
**[[Cryptocephalus prusias]]*
**[[Cryptocephalus pseudolusitanicus]]*
**[[Cryptocephalus pseudomaccus]]* White, 1968
**[[Cryptocephalus pseudoreitteri]]*
**[[Cryptocephalus pseudosindonicus]]*
**[[Cryptocephalus pubicollis]]* Linell, 1898
**[[Cryptocephalus pubiventris]]* Schaeffer, 1919
**[[Cryptocephalus pulchellus]]*
**[[Cryptocephalus pullus]]*
**[[Cryptocephalus pumilus]]* Haldeman, 1849
**[[Cryptocephalus punctatissimus]]*
**[[Cryptocephalus puncticollis]]*
*bjørkehettebladbille *Cryptocephalus punctiger*
**[[Cryptocephalus punctipes]]* Say, 1824
*høsthettebladbille *Cryptocephalus pusillus*
**[[Cryptocephalus pygmaeus]]*
**[[Cryptocephalus quadriguttatus]]*
**[[Cryptocephalus quadripunctatus]]*
*fireflekket furunålbille *Cryptocephalus quadripustulatus*
**[[Cryptocephalus quadruplex]]* Newman, 1841
**[[Cryptocephalus quatuordecimmaculatus]]*
*eikehettebladbille *Cryptocephalus querceti*
**[[Cryptocephalus quercus]]* Schaeffer, 1906
**[[Cryptocephalus quinquepunctatus]]*
**[[Cryptocephalus rabatensis]]*
**[[Cryptocephalus ramburii]]*
**[[Cryptocephalus reichei]]*
**[[Cryptocephalus reitteri]]*
**[[Cryptocephalus renatae]]*
**[[Cryptocephalus richteri]]*
**[[Cryptocephalus rubi]]*
**[[Cryptocephalus rufilabris]]*
**[[Cryptocephalus rufipes]]*
**[[Cryptocephalus rufofasciatus]]*
**[[Cryptocephalus rugicollis]]*
*enghettebladbille (*Cryptocephalus rugulipennis*er synonym)
**[[Cryptocephalus saintpierrei]]*
**[[Cryptocephalus saliceti]]*
**[[Cryptocephalus samniticus]]*
**[[Cryptocephalus sanguinicollis]]* Suffrian, 1852
**[[Cryptocephalus sareptanus]]*
**[[Cryptocephalus saucius]]*
**[[Cryptocephalus scapularis]]*
**[[Cryptocephalus schaefferi]]*
**[[Cryptocephalus schreibersii]]* Suffrian, 1852
**[[Cryptocephalus senegalensis]]*
*prakthettebladbille *Cryptocephalus sericeus*
*seksflekket hettebladbille *Cryptocephalus sexpunctatus*
**[[Cryptocephalus sexpustulatus]]*
**[[Cryptocephalus sexsignatus]]* (Fabricius, 1801)
**[[Cryptocephalus signatifrons]]*
**[[Cryptocephalus simulans]]* Schaeffer, 1906
**[[Cryptocephalus sinaita]]*
**[[Cryptocephalus snowi]]* Schaeffer, 1933
**[[Cryptocephalus solivagus]]*
**[[Cryptocephalus spilothorax]]*
**[[Cryptocephalus spurcus]]* Leconte, 1858
**[[Cryptocephalus stragula]]*
**[[Cryptocephalus striatulus]]* Leconte, 1880
**[[Cryptocephalus strigosus]]*
**[[Cryptocephalus suffriani]]*
**[[Cryptocephalus sulphureus]]*
**[[Cryptocephalus sultani]]*
**[[Cryptocephalus surdus]]*
**[[Cryptocephalus tappesi]]*
**[[Cryptocephalus tardus]]*
**[[Cryptocephalus telueticus]]*
**[[Cryptocephalus tetraspilus]]*
**[[Cryptocephalus texanus]]* Schaeffer, 1933
**[[Cryptocephalus therondi]]*
**[[Cryptocephalus tibialis]]*
**[[Cryptocephalus tinctus]]* Leconte, 1880
**[[Cryptocephalus tramuntanae]]*
**[[Cryptocephalus transiens]]*
**[[Cryptocephalus trapezicollis]]*
**[[Cryptocephalus tricolor]]*
**[[Cryptocephalus trimaculatus]]*
**[[Cryptocephalus tristigma]]*
**[[Cryptocephalus triundulatus]]* White, 1968
**[[Cryptocephalus trivittatus]]* Olivier, 1808
**[[Cryptocephalus trizonatus]]* Suffrian, 1858
**[[Cryptocephalus tshorumae]]*
**[[Cryptocephalus tsutsumiuchii]]* Shigetoh, 2025
**[[Cryptocephalus turcicus]]*
**[[Cryptocephalus umbonatus]]* Schaeffer, 1906
**[[Cryptocephalus undulatus]]*
**[[Cryptocephalus vapidus]]* White, 1968
**[[Cryptocephalus variceps]]*
**[[Cryptocephalus variegatus]]*
**[[Cryptocephalus venustus]]* Fabricius, 1787
**[[Cryptocephalus vidali]]*
**[[Cryptocephalus villosulus]]*
**[[Cryptocephalus violaceus]]*
**[[Cryptocephalus virens]]*
**[[Cryptocephalus virginiensis]]* White, 1968
**[[Cryptocephalus vittatus]]*
**[[Cryptocephalus volkovitshi]]*
**[[Cryptocephalus wehnckei]]*
**[[Cryptocephalus wenquanensis]]* Duan, Wang & Zhu, 2025
**[[Cryptocephalus xanthus]]*
**[[Cryptocephalus yangae]]* Duan, Wang & Zhou, 2025
**[[Cryptocephalus zambanellus]]*
**[[Cryptocephalus zhangi]]* Duan, Wang & Zhu, 2025
**[[Cryptocephalus zoiai]]*

## Referanser


## Litteratur
* Lindroth, C.H. 1993. *Våra skalbaggar och hur man kãnner igen dem* (opptrykk av opprinnelig tre hefter (1942-1945. Bonnier forlag.) Fãltbiologerna. ca. 240 sider. ISBN 91-85094-722

## Eksterne lenker
* 
* ''Cryptocephalus'' (Borowiec, Lech. ''European Chrysomelidae'') – www.biol.uni.wroc.pl
* ''Cryptocephalus'' - Fauna Europaea
* Bestemmelsesnøkkel til de Britiske slektene i Cryptocephalinae – Pdf-fil
* Oversikt over nordamerikanske arter
* bugguide.net - bildedatabase over nordamerikanske insekter
* Bestemmelsesnøkkel til de europeiske artene
* Norsk Entomologisk forening