Harpalus

* *Acardystus*
* *Artabas*
* *Harpalophonus*
* *Harpalus*
* *Licinoderus*

***Harpalus*** er en slekt av billene i delgruppen (underfamilien) markløpere (Harpalinae) i familiegruppen løpebiller.  Systematikken følger Carabidae of the World.

## Systematisk inndeling
Oversikten har med de europeiske artene, men den kan ha feil, som at både en god art og dens synonym er oppført (dobbeloppføring). 15 (?) arter er registret fra Norge, men ved søk i Fauna Europaea listes bare 12 arter, inkludert noen underarter, for Norge.

* ordenen biller,** Coleoptera** Linnaeus, 1758
** underordenen Adephaga Schellenberg, 1806
*** overfamilien løpebiller og vannkalver, **[[Caraboidea]]** Latreille, 1802
**** familien  ** [[løpebiller]]**, Carabidae Latreille, 1802
***** underfamilie **[[Markløpere (insekt)|markløpere]]** Harpalinae Bonelli, 1810
****** stammen Harpalini Bonelli, 1810 
******* slekten *Harpalus* Latreille, 1802
******** underslekten *Acardystus*
********* *[[Harpalus flavescens]]* (Piller & Mitterpacher 1783)
******** underslekten *Artabas*
********* *[[Harpalus dispar]]* Dejean 1829
********* *[[Harpalus petri]]* Tschitschérine 1902
********* *[[Harpalus punctatostriatus]]* Dejean 1829
********* *[[Harpalus rumelicus]]* Apfelbeck 1904
******** underslekten *Harpalophonus*
********* *[[Harpalus circumpunctatus]]* Chaudoir 1846
********* *[[Harpalus hospes]]* Sturm 1818
********* *[[Harpalus italus]]* Schaum 1860
********* *[[Harpalus stevenii]]* Dejean 1829
********* *[[Harpalus terrestris]]* (Motschulsky 1844)
******** underslekten *Harpalus*
********* *[[Harpalus affinis]]* (Schrank, 1781) 
********* *[[Harpalus akinini]]* Tschitschérine 1895
********* *[[Harpalus albanicus]]* Reitter 1900
********* *[[Harpalus amplicollis]]* Ménétriés 1848
********* *[[Harpalus angulatus]]* Putzeys 1878
********* *[[Harpalus anxius]]* (Duftschmid, 1812) 
********* *[[Harpalus atratus]]* Latreille 1804
********* *[[Harpalus attenuatus]]* Stephens, 1828 
********* *[[Harpalus autumnalis]]* (Duftschmid 1812)
********* *[[Harpalus bellieri]]* Reiche 1861
********* *[[Harpalus brevis]]* Motschulsky 1844
********* *[[Harpalus calathoides]]* Motschulsky 1844
********* *[[Harpalus calceatus]]* (Duftschmid, 1812) 
********* *[[Harpalus cardoni]]* Antoine 1922
********* *[[Harpalus caspius]]* (Steven 1806)
********* *[[Harpalus cisteloides]]* Motschulsky 1844
********* *[[Harpalus compressus]]* Motschulsky 1844
********* *[[Harpalus contemptus]]* Dejean 1829
********* *[[Harpalus cupreus]]* Dejean, 1829 
********* *[[Harpalus cyclogonus]]* Chaudoir 1844
********* *[[Harpalus danieli]]* Reitter 1900
********* *[[Harpalus decipiens]]* Dejean 1829
********* *[[Harpalus dimidiatus]]* (P. Rossi, 1790) 
********* *[[Harpalus distinguendus]]* (Duftschmid 1812)
********* *[[Harpalus ebeninus]]* Heyden 1870
********* *[[Harpalus euchlorus]]* Ménétriés 1836
********* *[[Harpalus flavicornis]]* Dejean 1829
********* *[[Harpalus foveiger]]* Tschitschérine 1895
********* *[[Harpalus froelichii]]* Sturm, 1818 
********* *[[Harpalus fuscicornis]]* Ménétriés 1832
********* *[[Harpalus fuscipalpis]]* Sturm 1818
********* *[[Harpalus griseus]]* Panzer, 1797 
********* *[[Harpalus hirtipes]]* (Panzer 1796)
********* *[[Harpalus honestus]]* (Duftschmid, 1812) 
********* *[[Harpalus inexpectatus]]* Kataev 1989
********* *[[Harpalus karamani]]* Apfelbeck 1902
********* *[[Harpalus kirgisicus]]* (Motschulsky 1844)
********* *[[Harpalus laevipes]]* Zetterstedt 1828   [syn.  *[[Harpalus quadripunctatus]]* Dejean, 1829 ]
********* *[[Harpalus lateralis]]* Dejean 1829
********* *[[Harpalus latus]]* (Linnaeus, 1758) 
********* *[[Harpalus lethierryi]]* Reiche 1860
********* *[[Harpalus luteicornis]]* (Duftschmid 1812)
********* *[[Harpalus lutshniki]]* Schauberger 1932
********* *[[Harpalus marginellus]]* Dejean 1829
********* *[[Harpalus melancholicus]]* Dejean, 1829 
********* *[[Harpalus metallinus]]* Ménétriés 1836
********* *[[Harpalus microthorax]]* (Motschulsky 1849)
********* *[[Harpalus mitridati]]* Pliginski 1915
********* *[[Harpalus modestus]]* Dejean 1829
********* *[[Harpalus neglectus]]* Serville, 1821
********* *[[Harpalus nevadensis]]* K. Daniel & J. Daniel 1898
********* *[[Harpalus nigritarsis]]* C.R. Sahlberg 1827
********* *[[Harpalus numidicus]]* Bedel 1893
********* *[[Harpalus oblitus]]* Dejean 1829
********* *[[Harpalus oodioides]]* Dejean 1829
********* *[[Harpalus optabilis]]* Dejean 1829
********* *[[Harpalus picipennis]]* (Duftschmid 1812)
********* *[[Harpalus politus]]* Dejean 1829
********* *[[Harpalus polyglyptus]]* Schaum 1862
********* *[[Harpalus progrediens]]* Schauberger 1922
********* *[[Harpalus pulvinatus]]* Ménétriés 1848
********* *[[Harpalus pumilus]]* Sturm, 1818 
********* *[[Harpalus punctipennis]]* Mulsant 1852
********* *[[Harpalus pygmaeus]]* Dejean 1829
********* *[[Harpalus rubripes]]* (Duftschmid, 1812) 
********* *[[Harpalus rufipalpis]]* Sturm, 1818 
********* *[[Harpalus rufiscapus]]* Gebler 1833
********* *[[Harpalus rufitarsoides]]* Schauberger 1934
********* *[[Harpalus rufipes]]* (De Geer, 1774) 
********* *[[Harpalus rufitarsis]]* (Duftschmid, 1812) non Illiger, 1801
********* *[[Harpalus salinus]]* Dejean 1829
********* *[[Harpalus saxicola]]* Dejean 1829
********* *[[Harpalus serripes]]* (Quensel in Schoenherr, 1806) 
********* *[[Harpalus servus]]* (Duftschmid, 1812) 
********* *[[Harpalus siculus]]* Dejean 1829
********* *[[Harpalus smaragdinus]]* (Duftschmid, 1812) 
********* *[[Harpalus solitaris]]* Dejean 1829
********* *[[Harpalus subcylindricus]]* Dejean 1829
********* *[[Harpalus sulphuripes]]* Germar 1824
********* *[[Harpalus taciturnus]]* Dejean 1829
********* *[[Harpalus tardus]]* (Panzer, 1796)
********* *[[Harpalus tenebrosus]]* Dejean, 1829 
********* *[[Harpalus tarsalis]]* Mannerheim 1825
********* *[[Harpalus tichonis]]* Jakobson 1907
********* *[[Harpalus torridoides]]* Reitter 1900
********* *[[Harpalus triseriatus]]* Fleischer 1897
********* *[[Harpalus vernalis]]* (Duftschmid, 1812) non Panzer, 1796 
********* *[[Harpalus xanthopus]]* Gemminger & Harold 1868
********* *[[Harpalus zabroides]]* Dejean 1829
******** underslekten *Licinoderus*
********* *[[Harpalus chobautianus]]* Lutshnik 1922
********* *[[Harpalus franzi]]* Mateu 1954
******** ikke plassert til underslekt
********* †*[[Harpalus mckayi]]* McKay & Mnguni, 2026
********* †*[[Harpalus mnguni]]* McKay & Mnguni, 2026
********* †*[[Harpalus nkqwili]]* McKay & Mnguni, 2026
********* †*[[Harpalus peteni]]* McKay & Mnguni, 2026

## Referanser


## Litteratur
* Bertram G. Rye. 1908. *Biller I. Løpebiller.* Danmarks Fauna Bind 3. G.E.C. GAD, København.

## Eksterne lenker
* 
* ''Harpalus'' – Database Carabidae of the World – engelskspråklig
* ''Harpalus'' - Fauna Europaea - engelskspråklig
* Norsk Entomologisk forening