Onthophagus (i snever forstand)

***Onthophagus*** (i snever forstand) er en underslekt av biller som hører til den artsrike slekten *[[Onthophagus]]* i familien skarabider (Scarabaeidae) i gruppen skarabider (Scarabaeoidea). De lever av møkk som de graver dypt ned i jorda.

## Utseende

Små til middelsstore, kraftige biller. Kroppsformen er kort, kraftig og hvelvet. Hodet er spadeformet og bredt, med små fasettøyne og korte antenner. Hos denne underslekten har hannene gjerne et par lange, tynne, krumme, bakoverrettede horn i pannen. Pronotum er stort og bredt (vanligvis tydelig bredere enn dekkvingene), gjerne med tydelig mikroskulptur, av og til metallglinsende. Dekkvingene er korte og hvelvede, ofte med tydelige lengdestriper.
Beina er kraftige gravebein.

## Levevis
Gjødselgraverne er knyttet til møkk. De graver tunneler under møkka, som ofte kan være overraskende lange sammenlignet med billens størrelse, og leder ned til et ynglekammer der hunnen arbeider møkka til en ball som hun legger egg på.

## Utbredelse

## Systematisk inndeling
Listen er temmelig ufullstendig.

* Ordenen biller, Coleoptera
** Underordenen Polyphaga
*** Overfamilien skarabider, Scarabaeoidea
**** Familien skarabider, (Scarabaeidae) Latreille, 1806
***** Underfamilien gjødselgravere (Scarabaeinae) Latreille, 1806
****** slektsgruppen Onthophagini
******* slekten gjødselgravere, *[[Onthophagus]]* Latreille, 1802
******** Underslekten *Onthophagus (i snever forstand)* Latreille, 1802
******** I Europa:
********* *[[Onthophagus binodis|Onthophagus (Onthophagus) binodis]]* (Thunberg, 1818)
********* *[[Onthophagus (Onthophagus) illyricus]]* (Scopoli, 1763)
********* *[[Onthophagus (Onthophagus) taurus]]* (Schreber, 1759)
******** Utenfor Europa:
********* *[[Onthophagus (Onthophagus) abacus]]* Boucomont, 1921
********* *[[Onthophagus (Onthophagus) abreui]]* Arrow, 1931
********* *[[Onthophagus (Onthophagus) aciculatus]]* Blatchley, 1928
********* *[[Onthophagus (Onthophagus) acuticollis]]* Gillet, 1927
********* *[[Onthophagus (Onthophagus) acuticornis]]* Endrödi, 1973
********* *[[Onthophagus (Onthophagus) aegrotus]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) afghanus]]* Petrovitz, 1961
********* *[[Onthophagus (Onthophagus) agnes]]* Gillet
********* *[[Onthophagus (Onthophagus) akhaus]]* Masumoto, Ochi & Hanboonsong, 2007
********* *[[Onthophagus (Onthophagus) alluvius]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) alternans]]* Raffray, 1877
********* *[[Onthophagus (Onthophagus) amicus]]* Gillet, 1925
********* *[[Onthophagus (Onthophagus) amirus]]* Kabakov, 1982
********* *[[Onthophagus (Onthophagus) anamalaiensis]]* Balthasar, 1974
********* *[[Onthophagus (Onthophagus) anguicornis]]* Boucomont
********* *[[Onthophagus (Onthophagus) angustatus]]* Boucomont, 1914
********* *[[Onthophagus (Onthophagus) antivagans]]* Howden & Génier, 2004
********* *[[Onthophagus (Onthophagus) apilularius]]* Masumoto, 1995
********* *[[Onthophagus (Onthophagus) apunneeae]]* Masumoto, Ochi & Hanboonsong, 2007
********* *[[Onthophagus (Onthophagus) armatus]]* Blanchard, 1853
********* *[[Onthophagus (Onthophagus) arnetti]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) arnoldii]]* Kabakov, 1982
********* *[[Onthophagus (Onthophagus) azusae]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) baloghi]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) bartosi]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) batesi]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) bengali]]* Gordon & Oppenheimer, 1976
********* *[[Onthophagus (Onthophagus) bicolensis]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) bicuneus]]* Kabakov, 2008
********* *[[Onthophagus (Onthophagus) biplagiatus]]* Thomson, 1858
********* *[[Onthophagus (Onthophagus) bistiniocelloides]]* Krikken, 1986
********* *[[Onthophagus (Onthophagus) bivertex]]* Heyden, 1887
********* *[[Onthophagus (Onthophagus) bokiaunus]]* Masumoto, 1996
********* *[[Onthophagus (Onthophagus) bomberaianus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) borneomontanus]]* Ochi & Kon, 2005
********* *[[Onthophagus (Onthophagus) brazzavillanus]]* Balthasar, 1974
********* *[[Onthophagus (Onthophagus) brevicollis]]* Arrow, 1931
********* *[[Onthophagus (Onthophagus) brevifrons]]* Horn, 1881
********* *[[Onthophagus (Onthophagus) browni]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) busiris]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) caelator]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) calchas]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) canelasensis]]* Howden & Génier, 2004
********* *[[Onthophagus (Onthophagus) caprai]]* Frey, 1956
********* *[[Onthophagus (Onthophagus) cartwrighti]]* Howden, 1973
********* *[[Onthophagus (Onthophagus) cavernicollis]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) centricornis]]* (Fabricius, 1798)
********* *[[Onthophagus (Onthophagus) cervus]]* (Fabricius, 1798)
********* *[[Onthophagus (Onthophagus) chaiyaphumensis]]* Masumoto, Ochi & Hanboonsong, 2002
********* *[[Onthophagus (Onthophagus) chevrolati]]* von Harold, 1869
********* *[[Onthophagus (Onthophagus) chremes]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) cicer]]* Balthasar, 1964
********* *[[Onthophagus (Onthophagus) clypeatus]]* Blanchard, 1845
********* *[[Onthophagus (Onthophagus) coahuilae]]* Zunino & Halftter, 1988
********* *[[Onthophagus (Onthophagus) cochisus]]* Brown, 1927
********* *[[Onthophagus (Onthophagus) concinnus]]* Laporte de Castelnau, 1840
********* *[[Onthophagus (Onthophagus) coproides]]* Horn, 1881
********* *[[Onthophagus (Onthophagus) crantor]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) cratippus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) cribricollis]]* Horn, 1881
********* *[[Onthophagus (Onthophagus) cuevensis]]* Howden, 1973
********* *[[Onthophagus (Onthophagus) cupreopastillatus]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) cyanellus]]* Bates, 1887
********* *[[Onthophagus (Onthophagus) cynomysi]]* Brown, 1927
********* *[[Onthophagus (Onthophagus) deemaak]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) dejongi]]* Krikken, 1986
********* *[[Onthophagus (Onthophagus) depilis]]* d'Orbigny, 1902
********* *[[Onthophagus (Onthophagus) depressus]]* von Harold, 1871
********* *[[Onthophagus (Onthophagus) dicax]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) dichropygus]]* Balthasar, 1963
********* *[[Onthophagus (Onthophagus) dissentaneus]]* Balthasar, 1964
********* *[[Onthophagus (Onthophagus) dlabolai]]* Balthasar, 1964
********* *[[Onthophagus (Onthophagus) doiinthanonensis]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) doipuiensis]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) doitungensis]]* Masumoto, Ochi & Hanboonsong , 2002
********* *[[Onthophagus (Onthophagus) duporti]]* Boucomont, 1914
********* *[[Onthophagus (Onthophagus) eliptamimus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) embersoni]]* Masumoto, Hanboonsong & Ochi, 2002
********* *[[Onthophagus (Onthophagus) falcarius]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) falcifer]]* von Harold, 1880
********* *[[Onthophagus (Onthophagus) falsivigilans]]* Masumoto, 1995
********* *[[Onthophagus (Onthophagus) fasciatus]]* Boucomont, 1914
********* *[[Onthophagus (Onthophagus) finschi]]* von Harold, 1877
********* *[[Onthophagus (Onthophagus) fissicornis]]* von Steven, 1809
********* *[[Onthophagus (Onthophagus) formosanus]]* Gillet, 1924
********* *[[Onthophagus (Onthophagus) fuscidorsis]]* d'Orbigny, 1902
********* *[[Onthophagus (Onthophagus) fuscus]]* Boucomont, 1932
********* *[[Onthophagus (Onthophagus) geryon]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) ghanensis]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) gibbulus]]* (Pallas, 1781)
********* *[[Onthophagus (Onthophagus) gorochovi]]* Kabakov, 1998
********* *[[Onthophagus (Onthophagus) gosoli]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) grandivigilans]]* Masumoto, 1995
********* *[[Onthophagus (Onthophagus) gratus]]* Arrow, 1931
********* *[[Onthophagus (Onthophagus) griseosetosus]]* Arrow, 1931
********* *[[Onthophagus (Onthophagus) hajimai]]* Masumoto, 1984
********* *[[Onthophagus (Onthophagus) hayashii]]* Masumoto, 1991
********* *[[Onthophagus (Onthophagus) hippopotamus]]* Bates
********* *[[Onthophagus (Onthophagus) hecate]]* (Panzer, 1794)
********* *[[Onthophagus (Onthophagus) hidalgus]]* Zunino & Halftter, 1988
********* *[[Onthophagus (Onthophagus) hoberlandti]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) hoepfneri]]* von Harold, 1869
********* *[[Onthophagus (Onthophagus) horii]]* Ochi, Kon & Tsubaki, 2009
********* *[[Onthophagus (Onthophagus) horribilis]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) hyrcanicus]]* Kabakov, 1982
********* *[[Onthophagus (Onthophagus) ifugaoensis]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) incantatus]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) incensus]]* Say, 1835
********* *[[Onthophagus (Onthophagus) inermifrons]]* d'Orbigny, 1902
********* *[[Onthophagus (Onthophagus) irianus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) isanus]]* Masumoto, Ochi & Hanboonsong , 2008
********* *[[Onthophagus (Onthophagus) ishii]]* Ochi & Kon, 1995
********* *[[Onthophagus (Onthophagus) iumenius]]* Masumoto, Ochi & Hanboonsong , 2007
********* *[[Onthophagus (Onthophagus) ivae]]* Balthasar, 1963
********* *[[Onthophagus (Onthophagus) javaecola]]* Balthasar, 1959
********* *[[Onthophagus (Onthophagus) kachinicus]]* Kabakov, 2006
********* *[[Onthophagus (Onthophagus) kanyaayonus]]* Masumoto, 1992
********* *[[Onthophagus (Onthophagus) kaosoidowensis]]* Masumoto, Ochi & Hanboonsong , 2008
********* *[[Onthophagus (Onthophagus) kapuri]]* Endrödi, 1974
********* *[[Onthophagus (Onthophagus) kashizakii]]* Ochi & Kon, 2005
********* *[[Onthophagus (Onthophagus) kashmirensis]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) kaszabi]]* Balthasar, 1963
********* *[[Onthophagus (Onthophagus) kawaharai]]* Ochi & Kon, 2007
********* *[[Onthophagus (Onthophagus) khonmiinitnoi]]* Masumoto, 1990
********* *[[Onthophagus (Onthophagus) kiuchii]]* Masumoto, 1995
********* *[[Onthophagus (Onthophagus) knausi]]* Brown, 1927
********* *[[Onthophagus (Onthophagus) knulli]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) konsarnensis]]* Masumoto, Ochi & Hanboonsong , 2008
********* *[[Onthophagus (Onthophagus) koryoensis]]* Kim, 1985
********* *[[Onthophagus (Onthophagus) krakadaakhomus]]* Masumoto, 1992
********* *[[Onthophagus (Onthophagus) kryzhanovskii]]* Kabakov, 1982
********* *[[Onthophagus (Onthophagus) labdacus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) laboranus]]* Arrow, 1931
********* *[[Onthophagus (Onthophagus) lakyim]]* Masumoto, 1990
********* *[[Onthophagus (Onthophagus) lamizanai]]* Cambefort, 1983
********* *[[Onthophagus (Onthophagus) landolti]]* von Harold, 1880
********* *[[Onthophagus (Onthophagus) lassulus]]* Balthasar, 1964
********* *[[Onthophagus (Onthophagus) lecontei]]* von Harold, 1871
********* *[[Onthophagus (Onthophagus) lefiniensis]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) lindaae]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) ludicrus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) maculatus]]* (Fabricius)
********* *[[Onthophagus (Onthophagus) magnioculus]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) mairuu]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) maleengnaafon]]* Masumoto, 1990
********* *[[Onthophagus (Onthophagus) maleengnoi]]* Masumoto, 1990
********* *[[Onthophagus (Onthophagus) malevolus]]* Balthasar, 1964
********* *[[Onthophagus (Onthophagus) maniti]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) mankonoensis]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) margaritifer]]* d'Orbigny, 1898
********* *[[Onthophagus (Onthophagus) masumotoi]]* Ochi, 1985
********* *[[Onthophagus (Onthophagus) matsudai]]* Ochi & Kon, 2004
********* *[[Onthophagus (Onthophagus) matsuii]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) mcclevei]]* Howden & Génier, 2004
********* *[[Onthophagus (Onthophagus) medorensis]]* Brown, 1929
********* *[[Onthophagus (Onthophagus) medvedevi]]* Kabakov, 1982
********* *[[Onthophagus (Onthophagus) megapacificus]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) metalliceps]]* Arrow
********* *[[Onthophagus (Onthophagus) mextexus]]* Howden & Cartwright, 1970
********* *[[Onthophagus (Onthophagus) mimikanus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) mopsus]]* (Fabricius, 1792)
********* *[[Onthophagus (Onthophagus) moroni]]* Zunino & Halftter, 1988
********* *[[Onthophagus (Onthophagus) mulleri]]* Lansberge
********* *[[Onthophagus (Onthophagus) naaroon]]* Masumoto, 1990
********* *[[Onthophagus (Onthophagus) namnaoensis]]* Masumoto, Ochi & Hanboonsong , 2002
********* *[[Onthophagus (Onthophagus) nasonis]]* Kabakov, 1998
********* *[[Onthophagus (Onthophagus) niasensis]]* Boucomont, 1914
********* *[[Onthophagus (Onthophagus) novaeirlandiae]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) nurestanicus]]* Kabakov, 1982
********* *[[Onthophagus (Onthophagus) obenbergeri]]* Balthasar, 1965
********* *[[Onthophagus (Onthophagus) occipitalis]]* Lansberge
********* *[[Onthophagus (Onthophagus) ochii]]* Masumoto, 1988
********* *[[Onthophagus (Onthophagus) ohkuboi]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) okahandjanus]]* Balthasar, 1974
********* *[[Onthophagus (Onthophagus) oklahomensis]]* Brown, 1924
********* *[[Onthophagus (Onthophagus) olsoufieffi]]*
********* *[[Onthophagus (Onthophagus) orientalis]]* von Harold, 1868
********* *[[Onthophagus (Onthophagus) orissanus]]* Arrow, 1931
********* *[[Onthophagus (Onthophagus) orpheus]]* (Panzer, 1794)
********* *[[Onthophagus (Onthophagus) orthocerus]]* Thomson, 1858
********* *[[Onthophagus (Onthophagus) otai]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) otjivarongus]]* Balthasar, 1967
********* *[[Onthophagus (Onthophagus) parafasciatus]]* Balthasar, 1974
********* *[[Onthophagus (Onthophagus) parapedisequus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) paraprotzenkoi]]* Tesar, 1967
********* *[[Onthophagus (Onthophagus) parcepilosus]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) pedisequus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) penicillatus]]* von Harold, 1879
********* *[[Onthophagus (Onthophagus) pennsylvanicus]]* von Harold, 1871
********* *[[Onthophagus (Onthophagus) perroti]]* Paulian, 1978
********* *[[Onthophagus (Onthophagus) phatoensis]]* Masumoto, Ochi & Hanboonsong , 2008
********* *[[Onthophagus (Onthophagus) phraoensis]]* Masumoto, 1990
********* *[[Onthophagus (Onthophagus) phrixus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) phrutsaphaakhomus]]* Masumoto, 1992
********* *[[Onthophagus (Onthophagus) phukhieoensis]]* Masumoto, Ochi & Hanboonsong , 2007
********* *[[Onthophagus (Onthophagus) pleurogonus]]* d'Orbigny, 1913
********* *[[Onthophagus (Onthophagus) poephagus]]* Kabakov, 1979
********* *[[Onthophagus (Onthophagus) polyphemi]]* Hubbard, 1894
********* *[[Onthophagus (Onthophagus) popovi]]* Kabakov
********* *[[Onthophagus (Onthophagus) protzenkoi]]* Tesar, 1967
********* *[[Onthophagus (Onthophagus) pseudofuscus]]* Zunino & Halftter, 1988
********* *[[Onthophagus (Onthophagus) punctator]]* Reitter, 1892
********* *[[Onthophagus (Onthophagus) punneeae]]* Masumoto, 1989
********* *[[Onthophagus (Onthophagus) ratchasimaensis]]* Masumoto, Hanboonsong & Ochi, 2002
********* *[[Onthophagus (Onthophagus) rechingeri]]* Petrovitz, 1980
********* *[[Onthophagus (Onthophagus) retecornutus]]* van Lansberge, 1883
********* *[[Onthophagus (Onthophagus) robertopoggii]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) roubali]]* Balthasar, 1935
********* *[[Onthophagus (Onthophagus) rubripennis]]* Arrow, 1907
********* *[[Onthophagus (Onthophagus) rudis]]* Sharp, 1875
********* *[[Onthophagus (Onthophagus) rugulosus]]* Harold, 1886
********* *[[Onthophagus (Onthophagus) rutriceps]]* Krikken & Huijbregts, 2012
********* *[[Onthophagus (Onthophagus) sabahensis]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) sakaeratensis]]* Masumoto, Hanboonsong & Ochi, 2002
********* *[[Onthophagus (Onthophagus) sakainoi]]* Masumoto, 1991
********* *[[Onthophagus (Onthophagus) salavazai]]* Paulian, 1978
********* *[[Onthophagus (Onthophagus) sangwalus]]* Masumoto, Ochi & Hanboonsong, 2008
********* *[[Onthophagus (Onthophagus) sauteri]]* Gillet, 1924
********* *[[Onthophagus (Onthophagus) schaefferi]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) sciron]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) seabrai]]* Gomes Alves, 1944
********* *[[Onthophagus (Onthophagus) semipacificus]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) sepilokensis]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) setchan]]* Masumoto, 1984
********* *[[Onthophagus (Onthophagus) silus]]* Balthasar, 1960
********* *[[Onthophagus (Onthophagus) simboroni]]* Ochi & Kon, 2006
********* *[[Onthophagus (Onthophagus) singhaakhomus]]* Masumoto, 1992
********* *[[Onthophagus (Onthophagus) sobrius]]* Balthasar, 1959
********* *[[Onthophagus (Onthophagus) spinifex]]* (Fabricius, 1781)
********* *[[Onthophagus (Onthophagus) stehliki]]* Balthasar, 1966
********* *[[Onthophagus (Onthophagus) sternax]]* Balthasar, 1959
********* *[[Onthophagus (Onthophagus) stomachosus]]* Krell, 2009
********* *[[Onthophagus (Onthophagus) striatulus]]* (Palisot de Beauvois, 1809)
********* *[[Onthophagus (Onthophagus) strictestriatus]]* d'Orbigny, 1913
********* *[[Onthophagus (Onthophagus) subaeneus]]* (Palisot de Beauvois, 1811)
********* *[[Onthophagus (Onthophagus) subalternans]]* d'Orbigny, 1902
********* *[[Onthophagus (Onthophagus) subopacus]]* Robinson, 1940
********* *[[Onthophagus (Onthophagus) subtropicus]]* Howden & Cartwright, 1963
********* *[[Onthophagus (Onthophagus) sunanicus]]* Stebnicka, 1973
********* *[[Onthophagus (Onthophagus) taayai]]* Masumoto, 1995
********* *[[Onthophagus (Onthophagus) taxillus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) telegonus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) telephus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) tenax]]* Balthasar, 1964
********* *[[Onthophagus (Onthophagus) tholaayi]]* Masumoto, 1990
********* *[[Onthophagus (Onthophagus) tiniocelloides]]* Boucomont, 1925
********* *[[Onthophagus (Onthophagus) totutlaensis]]* Sánchez-Huerta, Moctezuma & Halffter, 2025
********* *[[Onthophagus (Onthophagus) triptolemus]]* Balthasar, 1969
********* *[[Onthophagus (Onthophagus) tritinctus]]' <small>Boucomont, 1914</small>
********* *Onthophagus (Onthophagus) truncaticornis* <small>(Schaller, 1783)</small>
********* *Onthophagus (Onthophagus) tschadensis* <small>Balthasar, 1963</small>
********* *Onthophagus (Onthophagus) tuberculifrons* <small>von Harold, 1871</small>
********* *Onthophagus (Onthophagus) tumami* <small>Masumoto, Ochi & Hanboonsong, 2002</small>
********* *Onthophagus (Onthophagus) tungkamangensis* <small>Masumoto, Ochi & Hanboonsong, 2007</small>
********* *Onthophagus (Onthophagus) turfanicus* <small>Balthasar, 1971</small>
********* *Onthophagus (Onthophagus) umbratus* <small>d'Orbigny, 1902</small>
********* *Onthophagus (Onthophagus) ulula* <small>Balthasar, 1966</small>
********* *Onthophagus (Onthophagus) unifasciatus* <small>(Schaller, 1783)</small>
********* *Onthophagus (Onthophagus) usurpator* <small>Balthasar, 1959</small>
********* *Onthophagus (Onthophagus) velutinus* <small>Horn, 1875</small>
********* *Onthophagus (Onthophagus) vesanus* <small>Balthasar, 1967</small>
********* *Onthophagus (Onthophagus) vespertilio* <small>Howden, Cartwright & Halftter</small>
********* *Onthophagus (Onthophagus) vitulus* <small>([[Fabricius]], 1776)</small>
********* *Onthophagus (Onthophagus) wangnamkhieoensis* <small>Masumoto, Hanboonsong & Ochi, 2002</small>
********* *Onthophagus (Onthophagus) xiquenus* <small>Sánchez-Huerta, Moctezuma & Halffter, 2025</small>
********* *Onthophagus (Onthophagus) yamaokai* <small>Masumoto, Ochi & Hanboonsong, 2002</small>
********* *Onthophagus (Onthophagus) yangi* <small>Masumoto, Tsai & Ochi, 2006</small>
********* *Onthophagus (Onthophagus) zebra* <small>Arrow, 1931</small>
********* *Onthophagus (Onthophagus) zicsii* <small>Balthasar, 1967</small>
********* *Onthophagus (Onthophagus) zimmermanni'' Balthasar, 1959

## Referanser


## Kilder
* Liste over artene 
* Fauna Europaea Web Service Version 2.4 (2011) Utbredelsesdatabase for europeiske dyr 

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