Heteronyx

***Heteronyx*** er en artsrik slekt av biller som har larver som eter planterøtter. De hører til underfamilien oldenborrer (Melolonthinae) i den store gruppen skarabider (Scarabaeoidea). Disse billene lever i Australia.

## Utseende
Små til middelsstore (5-12 millimeter), avlange, noe blanke, rødbrune til svarte oldenborrer. Oversiden er mer eller mindre tett kledt med korte hår.

## Levevis
Larvene lever i jorda der de eter røtter og råtnende plantemateriale. Larveutviklingen tar vanligvis ett år, men kan også vare to. De voksne billene kan gjøre noe skade ved å gnage på blader på busker og trær. Noen arter er nattaktive, andre dagaktive.

## Utbredelse
Artene lever i Australia og omkringliggende øyer i alle fall vest til Molukkene. Fem arter fra i Bolivia og Brasil ble tidligere ført til denne slekten, men er nå overført til slekten *[[Blepharotoma]]*.

## Systematisk inndeling

* Ordenen biller, Coleoptera
** Underordenen Polyphaga
*** Overfamilien skarabider, Scarabaeoidea
**** Familien skarabider, Scarabaeidae Latreille, 1806
***** Underfamilien oldenborrer, Melolonthinae Samouelle, 1819
****** Stammen Liparetrini Burmeister, 1855
******* Slekten *Heteronyx* Guérin-Méneville, 1831
******** *[[Heteronyx acutifrons]]* Blackburn, 1889
******** *[[Heteronyx additus]]* Blackburn, 1909
******** *[[Heteronyx advena]]* Blackburn, 1890
******** *[[Heteronyx aequaliceps]]* Blackburn, 1890
******** *[[Heteronyx aequalis]]* Blackburn, 1889
******** *[[Heteronyx affinis]]* Blackburn, 1909
******** *[[Heteronyx agrestis]]* Burmeister, 1855
******** *[[Heteronyx agricola]]* Blackburn, 1910
******** *[[Heteronyx alienus]]* Blackburn, 1892
******** *[[Heteronyx alpicola]]* Blackburn, 1892
******** *[[Heteronyx ambiguus]]* Blackburn, 1910
******** *[[Heteronyx amboinensis]]* Moser, 1920
******** *[[Heteronyx anomalus]]* Blackburn, 1908
******** *[[Heteronyx apertus]]* Blackburn, 1910
******** *[[Heteronyx aphodioides]]* Blanchard, 1851
******** *[[Heteronyx apicispinosa]]* Heller, 1914
******** *[[Heteronyx arcanus]]* Blackburn, 1892
******** *[[Heteronyx aridus]]* Blackburn, 1889
******** *[[Heteronyx aruanus]]* Moser, 1920
******** *[[Heteronyx asperatus]]* Moser, 1920
******** *[[Heteronyx aspericollis]]* Blackburn, 1889
******** *[[Heteronyx asperifrons]]* Blackburn, 1909
******** *[[Heteronyx augustae]]* Blackburn, 1889
******** *[[Heteronyx australis]]* Guérin-Méneville, 1831
******** *[[Heteronyx badius]]* MacLeay, 1888
******** *[[Heteronyx beltanae]]* Blackburn, 1889
******** *[[Heteronyx bidentatus]]* Blackburn, 1889
******** *[[Heteronyx blandus]]* Blackburn, 1909
******** *[[Heteronyx borealis]]* Blackburn, 1889
******** *[[Heteronyx bovilli]]* Blackburn, 1890
******** *[[Heteronyx brevior]]* Fairmaire, 1883
******** *[[Heteronyx caledoniae]]* Fauvel, 1903
******** *[[Heteronyx calidus]]* Blackburn, 1910
******** *[[Heteronyx callabonnae]]* Blackburn, 1909
******** *[[Heteronyx capillatus]]* MacLeay, 1888
******** *[[Heteronyx castanescens]]* Moser, 1924
******** *[[Heteronyx castaneus]]* Blanchard, 1846
******** *[[Heteronyx cervina]]* (Boisduval, 1835)
******** *[[Heteronyx chlorotica]]* (Gyllenhal, 1817)
******** *[[Heteronyx cliens]]* Blackburn, 1910
******** *[[Heteronyx collaris]]* Blackburn, 1890
******** *[[Heteronyx colossus]]* Arrow, 1915
******** *[[Heteronyx comans]]* Blackburn, 1909
******** *[[Heteronyx concolor]]* MacLeay, 1871
******** *[[Heteronyx confertus]]* Blackburn, 1910
******** *[[Heteronyx constans]]* Blackburn, 1889
******** *[[Heteronyx corpulentus]]* MacLeay, 1888
******** *[[Heteronyx costulatus]]* Blackburn, 1910
******** *[[Heteronyx crassus]]* Blackburn, 1889
******** *[[Heteronyx cribriceps]]* Blackburn, 1892
******** *[[Heteronyx crinitus]]* Blackburn, 1910
******** *[[Heteronyx cunnamullae]]* Blackburn, 1910
******** *[[Heteronyx cygneus]]* Blackburn, 1889
******** *[[Heteronyx darlingensis]]* Blackburn, 1889
******** *[[Heteronyx declaratus]]* Blackburn, 1909
******** *[[Heteronyx dentipes]]* Blackburn, 1889
******** *[[Heteronyx dimidiata]]* (Erichson, 1842)
******** *[[Heteronyx disjectus]]* Blackburn, 1910
******** *[[Heteronyx diversiceps]]* Blackburn, 1890
******** *[[Heteronyx doctus]]* Blackburn, 1890
******** *[[Heteronyx doddi]]* Blackburn, 1908
******** *[[Heteronyx electus]]* Blackburn, 1889
******** *[[Heteronyx elongatus]]* Blanchard, 1851
******** *[[Heteronyx elytrurus]]* Blackburn, 1909
******** *[[Heteronyx excisus]]* Blackburn, 1890
******** *[[Heteronyx exsectus]]* Blackburn, 1909
******** *[[Heteronyx fervidus]]* Blackburn, 1892
******** *[[Heteronyx fimbriatus]]* Moser, 1924
******** *[[Heteronyx flavus]]* Blackburn, 1890
******** *[[Heteronyx fortis]]* Blackburn, 1898
******** *[[Heteronyx fraserensis]]* Blackburn, 1892
******** *[[Heteronyx frenchi]]* Blackburn, 1909
******** *[[Heteronyx froggatti]]* MacLeay, 1888
******** *[[Heteronyx frontalis]]* Blackburn, 1889
******** *[[Heteronyx fulvohirtus]]* Blackburn, 1889
******** *[[Heteronyx fumata]]* (Erichson, 1842)
******** *[[Heteronyx furvus]]* Blackburn, 1892
******** *[[Heteronyx glabratus]]* (Erichson, 1842)
******** *[[Heteronyx grandis]]* Blackburn, 1900
******** *[[Heteronyx granulatus]]* Blackburn, 1910
******** *[[Heteronyx granulifer]]* Blackburn, 1889
******** *[[Heteronyx granum]]* Burmeister, 1855
******** *[[Heteronyx helmsi]]* Blackburn, 1892
******** *[[Heteronyx hirsutosetosus]]* Moser, 1920
******** *[[Heteronyx hirsutus]]* Blackburn, 1910
******** *[[Heteronyx hirtuosus]]* Blackburn, 1890
******** *[[Heteronyx hispidulus]]* Blackburn, 1908
******** *[[Heteronyx horridus]]* Blackburn, 1889
******** *[[Heteronyx humilis]]* Blackburn, 1910
******** *[[Heteronyx imitator]]* Blackburn, 1909
******** *[[Heteronyx impar]]* Blackburn, 1910
******** *[[Heteronyx incognitus]]* Blackburn, 1892
******** *[[Heteronyx incola]]* Blackburn, 1889
******** *[[Heteronyx infirmus]]* Blackburn, 1910
******** *[[Heteronyx infuscatus]]* MacLeay, 1871
******** *[[Heteronyx inornatus]]* (Blackburn, 1900)
******** *[[Heteronyx insignis]]* Blackburn, 1889
******** *[[Heteronyx insularis]]* Fairmaire, 1883
******** *[[Heteronyx interioris]]* Blackburn, 1909
******** *[[Heteronyx intrusus]]* Blackburn, 1910
******** *[[Heteronyx iridiventris]]* Blackburn, 1890
******** *[[Heteronyx irrasus]]* Lea, 1926
******** *[[Heteronyx jejunus]]* Blackburn, 1889
******** *[[Heteronyx johannis]]* Blackburn, 1912
******** *[[Heteronyx labratus]]* Moser, 1926
******** *[[Heteronyx laeviceps]]* Blackburn, 1889
******** *[[Heteronyx laminatus]]* Blackburn, 1890
******** *[[Heteronyx lateritius]]* Blackburn, 1889
******** *[[Heteronyx laticeps]]* Burmeister, 1855
******** *[[Heteronyx leai]]* Blackburn, 1909
******** *[[Heteronyx liliputanus]]* Blackburn, 1890
******** *[[Heteronyx lindi]]* Blackburn, 1889
******** *[[Heteronyx lividus]]* Blackburn, 1889
******** *[[Heteronyx lobatus]]* Blackburn, 1889
******** *[[Heteronyx luteolus]]* Blackburn, 1909
******** *[[Heteronyx maculatus]]* Blackburn, 1889
******** *[[Heteronyx magnus]]* Moser, 1924
******** *[[Heteronyx major]]* Blackburn, 1910
******** *[[Heteronyx maluensis]]* Brenske, 1889
******** *[[Heteronyx marcidus]]* Blackburn, 1892
******** *[[Heteronyx marginatus]]* Blackburn, 1890
******** *[[Heteronyx merus]]* Blackburn, 1892
******** *[[Heteronyx metropolitanus]]* Blackburn, 1909
******** *[[Heteronyx mimus]]* Blackburn, 1890
******** *[[Heteronyx minimus]]* Lea, 1926
******** *[[Heteronyx minutus]]* Blackburn, 1910
******** *[[Heteronyx moluccanus]]* Moser, 1920
******** *[[Heteronyx montanus]]* Blackburn, 1890
******** *[[Heteronyx mulwalensis]]* Blackburn, 1889
******** *[[Heteronyx mundus]]* Blackburn, 1909
******** *[[Heteronyx nasutus]]* Blackburn, 1889
******** *[[Heteronyx neglectus]]* Blackburn, 1910
******** *[[Heteronyx nigella]]* (Erichson, 1842)
******** *[[Heteronyx nigrescens]]* Blackburn, 1909
******** *[[Heteronyx nigricans]]* Burmeister, 1855
******** *[[Heteronyx nigrita]]* Blanchard, 1851
******** *[[Heteronyx nigritulus]]* Moser, 1926
******** *[[Heteronyx normalis]]* Blackburn, 1889
******** *[[Heteronyx novitius]]* Blackburn, 1910
******** *[[Heteronyx obesa]]* Burmeister, 1855
******** *[[Heteronyx oblongus]]* Blanchard, 1851
******** *[[Heteronyx orbus]]* Blackburn, 1909
******** *[[Heteronyx ovatus]]* Blanchard, 1851
******** *[[Heteronyx pallidulus]]* MacLeay, 1871
******** *[[Heteronyx paniei]]* Paulian, 1991
******** *[[Heteronyx papuanus]]* Moser, 1920
******** *[[Heteronyx parvulus]]* MacLeay, 1888
******** *[[Heteronyx pauxillus]]* Blackburn, 1910
******** *[[Heteronyx pellucida]]* Burmeister, 1855
******** *[[Heteronyx peregrinus]]* Blackburn, 1890
******** *[[Heteronyx perkinsi]]* Blackburn, 1909
******** *[[Heteronyx piceoniger]]* MacLeay, 1888
******** *[[Heteronyx piceus]]* Blanchard, 1851
******** *[[Heteronyx pilosellus]]* Blanchard, 1851
******** *[[Heteronyx pilosus]]* Blanchard, 1846
******** *[[Heteronyx placidus]]* Blackburn, 1910
******** *[[Heteronyx planatus]]* Burmeister, 1855
******** *[[Heteronyx planiceps]]* Blackburn, 1910
******** *[[Heteronyx posticalis]]* Blackburn, 1890
******** *[[Heteronyx praecox]]* (Erichson, 1842)
******** *[[Heteronyx proditor]]* Blackburn, 1892
******** *[[Heteronyx proprius]]* Blackburn, 1910
******** *[[Heteronyx prosper]]* Blackburn, 1909
******** *[[Heteronyx protervus]]* Blackburn, 1892
******** *[[Heteronyx proxima]]* Burmeister, 1855
******** *[[Heteronyx pubescens]]* (Erichson, 1842)
******** *[[Heteronyx pumilus]]* Sharp, 1877
******** *[[Heteronyx puncticollis]]* Blackburn, 1890
******** *[[Heteronyx punctipennis]]* Blackburn, 1889
******** *[[Heteronyx pustulosus]]* Blackburn, 1890
******** *[[Heteronyx pygidialis]]* Blackburn, 1889
******** *[[Heteronyx pygmaeus]]* Blackburn, 1910
******** *[[Heteronyx quadraticollis]]* Blackburn, 1890
******** *[[Heteronyx queenlandicus]]* Blackburn, 1909
******** *[[Heteronyx randalli]]* Blackburn, 1890
******** *[[Heteronyx rectangulus]]* Blackburn, 1910
******** *[[Heteronyx relictus]]* Blackburn, 1910
******** *[[Heteronyx rhinastus]]* Blackburn, 1890
******** *[[Heteronyx rhinocerus]]* Blackburn, 1892
******** *[[Heteronyx rothei]]* Blackburn, 1890
******** *[[Heteronyx rotundiceps]]* Blanchard, 1851
******** *[[Heteronyx rubescens]]* Blanchard, 1851
******** *[[Heteronyx ruficollis]]* MacLeay, 1871
******** *[[Heteronyx rufomarginatus]]* Blanchard, 1851
******** *[[Heteronyx rufopiceus]]* MacLeay, 1888
******** *[[Heteronyx rugosipennis]]* MacLeay, 1871
******** *[[Heteronyx rusticus]]* Blackburn, 1890
******** *[[Heteronyx salebrosus]]* Blackburn, 1909
******** *[[Heteronyx satellus]]* Blackburn, 1889
******** *[[Heteronyx scalptus]]* Blackburn, 1890
******** *[[Heteronyx sequens]]* Blackburn, 1894
******** *[[Heteronyx setifer]]* Blackburn, 1890
******** *[[Heteronyx sexualis]]* Blackburn, 1909
******** *[[Heteronyx siccus]]* Blackburn, 1892
******** *[[Heteronyx simius]]* Blackburn, 1890
******** *[[Heteronyx simplicicollis]]* Blackburn, 1909
******** *[[Heteronyx simulator]]* Blackburn, 1889
******** *[[Heteronyx sloanei]]* Blackburn, 1889
******** *[[Heteronyx solidus]]* Blackburn, 1889
******** *[[Heteronyx spadicea]]* Burmeister, 1855
******** *[[Heteronyx spretus]]* Blackburn, 1889
******** *[[Heteronyx squalidus]]* Blackburn, 1908
******** *[[Heteronyx striatus]]* Blackburn, 1909
******** *[[Heteronyx suavis]]* Blackburn, 1909
******** *[[Heteronyx subferruginea]]* Burmeister, 1855
******** *[[Heteronyx subfuscus]]* MacLeay, 1888
******** *[[Heteronyx subglaber]]* MacLeay, 1888
******** *[[Heteronyx substriatus]]* MacLeay, 1871
******** *[[Heteronyx sulcifrons]]* Blackburn, 1909
******** *[[Heteronyx sydneyanus]]* Blackburn, 1890
******** *[[Heteronyx tarsalis]]* Blackburn, 1909
******** *[[Heteronyx tasmanicus]]* Blackburn, 1909
******** *[[Heteronyx tenebrosus]]* Blackburn, 1909
******** *[[Heteronyx tepperi]]* Blackburn, 1889
******** *[[Heteronyx terrena]]* Blackburn, 1892
******** *[[Heteronyx tindalei]]* Britton, 1992
******** *[[Heteronyx torvus]]* Blackburn, 1889
******** *[[Heteronyx transversicollis]]* MacLeay, 1888
******** *[[Heteronyx tridentatus]]* Lea, 1924
******** *[[Heteronyx tristis]]* Blackburn, 1889
******** *[[Heteronyx umbilicatus]]* Fauvel, 1903
******** *[[Heteronyx unicolor]]* Blanchard, 1851
******** *[[Heteronyx unicus]]* Blackburn, 1900
******** *[[Heteronyx variegatus]]* Blackburn, 1889
******** *[[Heteronyx vicinus]]* Blackburn, 1910
******** *[[Heteronyx victoris]]* Blackburn, 1889
******** *[[Heteronyx viduus]]* Blackburn, 1910
******** *[[Heteronyx zalotus]]* Blackburn, 1910

## Kilder
* Britton, E.B. (2000) A review of *Heteronyx* Guerin-Meneville (Coleoptera : Scarabaeidae : Melolonthinae). *Invertebrate Taxonomy* 14(4) 465 - 589. http://www.publish.csiro.au/paper/IT98009.htm
* Generic Guide to New World Scarab Beetles - *Heteronyx* http://museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Melolonthinae/Melolonthinae-Generic-pages/Liparetrini/Heteronyx/Heteronyx.html
* Biolib.cz - liste over artene http://www.biolib.cz/en/taxontree/id663336/
* IPMG Pestnotes - *Heteronyx* beetles https://web.archive.org/web/20140126143906/http://plantationhealth.com.au/pdf/pestnotes/heteronyx.pdf

## Eksterne lenker
* 
* Bilde av ''Heteronyx'' sp.