Scopaeus

***Scopaeus*** er en gruppe (slekt) av biller som hører til underfamilien smalkortvinger (Paederinae) i den store familien kortvinger (Staphylinidae).

## Utseende
Små (gjerne 2,5-3 millimeter), slanke kortvinger. Hodet er stort, tvert avskåret bak med markerte bakhjørner – dette er det beste kjennetegnet på slekten. "Halsen" som forbinder hodet med kroppen er ganske smal. Dekkvingene er korte.

## Levevis
Disse lange og smale billene lever i stor grad underjordisk, der de graver i jorda.

## Systematisk oversikt / norske arter
Disse artene ble tidligere gjerne regnet til den artsrike slekten *[[Lathrobium]]*.

* Ordenen biller, **Coleoptera**
** Underordenen Polyphaga
*** Overfamilien åtsel- og rovbiller, **Staphylinoidea**
**** Familien kortvinger (Staphylinidae) Latreille, 1802
***** Underfamilien smalkortvinger Paederinae Fleming, 1821
****** Slekten *Scopaeus* Erichson, 1839
******* *[[Scopaeus laevigatus]]* (Gyllenhal, 1827)
******* *[[Scopaeus cognatus]]* Mulsant & Rey, 1855
******* *[[Scopaeus pusillus]]* Kiesenwetter, 1843
### Liste over artene

* *[[Scopaeus abyssinicus]]*
* *[[Scopaeus admixtus]]*
* *[[Scopaeus alaniensis]]*
* *[[Scopaeus alaschiacus]]*
* *[[Scopaeus albertvillensis]]*
* *[[Scopaeus alborzensis]]*
* *[[Scopaeus aliiceps]]*
* *[[Scopaeus allardianus]]*
* *[[Scopaeus amos]]*
* *[[Scopaeus amphionis]]*
* *[[Scopaeus ampliatus]]*
* *[[Scopaeus andrewesi]]*
* *[[Scopaeus angolanus]]*
* *[[Scopaeus angustatus]]*
* *[[Scopaeus angusticeps]]*
* *[[Scopaeus angusticollis]]*
* *[[Scopaeus angustissimus]]*
* *[[Scopaeus anlasi]]*
* *[[Scopaeus annuliphallus]]*
* *[[Scopaeus antennalis]]*
* *[[Scopaeus antoniensis]]*
* *[[Scopaeus anxius]]*
* *[[Scopaeus apicicornis]]*
* *[[Scopaeus apiculatus]]*
* *[[Scopaeus apterus]]*
* *[[Scopaeus arcuatus]]*
* *[[Scopaeus arena]]*
* *[[Scopaeus argonauta]]*
* *[[Scopaeus arizonae]]*
* *[[Scopaeus asiaticus]]*
* *[[Scopaeus asirensis]]*
* *[[Scopaeus assingi]]*
* *[[Scopaeus auripilis]]*
* *[[Scopaeus australiensis]]*
* *[[Scopaeus azerbaidzhanus]]*
* *[[Scopaeus baliensis]]*
* *[[Scopaeus bamaniaensis]]*
* *[[Scopaeus beesoni]]*
* *[[Scopaeus beieri]]*
* *[[Scopaeus bertiae]]*
* *[[Scopaeus bicolor]]*
* *[[Scopaeus bicuspis]]*
* *[[Scopaeus bilaminulatus]]*
* *[[Scopaeus binaghii]]*
* *[[Scopaeus bingervillensis]]*
* *[[Scopaeus biskrensis]]*
* *[[Scopaeus bituberculatus]]*
* *[[Scopaeus blackburni]]*
* *[[Scopaeus borbonicus]]*
* *[[Scopaeus borneensis]]*
* *[[Scopaeus borumandi]]*
* *[[Scopaeus boxi]]*
* *[[Scopaeus brachypterus]]*
* *[[Scopaeus brasiliensis]]*
* *[[Scopaeus bredoanus]]*
* *[[Scopaeus brevicuspis]]*
* *[[Scopaeus brevipennis]]*
* *[[Scopaeus brunnescens]]*
* *[[Scopaeus brunnipes]]*
* *[[Scopaeus burgeonianus]]*
* *[[Scopaeus calidus]]*
* *[[Scopaeus californicus]]*
* *[[Scopaeus camelaensis]]*
* *[[Scopaeus cameroni]]*
* *[[Scopaeus cariensis]]*
* *[[Scopaeus carissimus]]*
* *[[Scopaeus carolinae]]*
* *[[Scopaeus caseyi]]*
* *[[Scopaeus caspius]]*
* *[[Scopaeus celisianus]]*
* *[[Scopaeus cervicula]]*
* *[[Scopaeus ceylonensis]]*
* *[[Scopaeus chalcodactylus]]*
* *[[Scopaeus championi]]*
* *[[Scopaeus chapini]]*
* *[[Scopaeus chatkalensis]]*
* *[[Scopaeus chinensis]]*
* *[[Scopaeus cinctipennis]]*
* *[[Scopaeus complex]]* Sharp, 1874
* *[[Scopaeus concavus]]*
* *[[Scopaeus concolor]]*
* *[[Scopaeus confusoides]]*
* *[[Scopaeus confusus]]*
* *[[Scopaeus consimilis]]*
* *[[Scopaeus convexiceps]]*
* *[[Scopaeus corpulentus]]*
* *[[Scopaeus courtoisi]]*
* *[[Scopaeus crassipes]]*
* *[[Scopaeus crassulus]]*
* *[[Scopaeus creticus]]*
* *[[Scopaeus ctenocryptus]]*
* *[[Scopaeus cupiens]]*
* *[[Scopaeus currax]]*
* *[[Scopaeus curraxoides]]*
* *[[Scopaeus curtipennis]]*
* *[[Scopaeus cylindricus]]*
* *[[Scopaeus cyprius]]*
* *[[Scopaeus darlingtoni]]*
* *[[Scopaeus debilis]]*
* *[[Scopaeus decelleanus]]*
* *[[Scopaeus decipiens]]* Kraatz, 1859
* *[[Scopaeus degener]]*
* *[[Scopaeus delicatissimus]]*
* *[[Scopaeus delicatulus]]*
* *[[Scopaeus delicatus]]*
* *[[Scopaeus densicollis]]*
* *[[Scopaeus desaegerianus]]*
* *[[Scopaeus didymus]]*
* *[[Scopaeus digitalis]]*
* *[[Scopaeus dilutus]]*
* *[[Scopaeus discoidalis]]*
* *[[Scopaeus dissentaneus]]*
* *[[Scopaeus dissimilis]]*
* *[[Scopaeus distans]]*
* *[[Scopaeus diversiceps]]*
* *[[Scopaeus dolosus]]*
* *[[Scopaeus dominicanus]]*
* *[[Scopaeus dubius]]*
* *[[Scopaeus durus]]*
* *[[Scopaeus duryi]]*
* *[[Scopaeus ebneri]]*
* *[[Scopaeus efesi]]*
* *[[Scopaeus effundatus]]*
* *[[Scopaeus elaboratus]]*
* *[[Scopaeus elegans]]*
* *[[Scopaeus elegantulus]]*
* *[[Scopaeus elgonensis]]*
* *[[Scopaeus elisabethvillensis]]*
* *[[Scopaeus endrodyanus]]*
* *[[Scopaeus errans]]*
* *[[Scopaeus erraticus]]*
* *[[Scopaeus evagatus]]*
* *[[Scopaeus exiguus]]*
* *[[Scopaeus fageli]]*
* *[[Scopaeus fagelianus]]*
* *[[Scopaeus fallaciosus]]*
* *[[Scopaeus farsensis]]*
* *[[Scopaeus fasciatellus]]*
* *[[Scopaeus fauveli]]*
* *[[Scopaeus femineus]]*
* *[[Scopaeus femoralis]]*
* *[[Scopaeus femursetosus]]*
* *[[Scopaeus ferganensis]]*
* *[[Scopaeus filicornis]]*
* *[[Scopaeus filiformis]]*
* *[[Scopaeus filitarsis]]*
* *[[Scopaeus filum]]*
* *[[Scopaeus flavidulus]]*
* *[[Scopaeus flavocastaneus]]*
* *[[Scopaeus flavofasciatus]]*
* *[[Scopaeus fluviatilis]]*
* *[[Scopaeus fossiceps]]*
* *[[Scopaeus franzi]]*
* *[[Scopaeus frater]]*
* *[[Scopaeus frommeri]]*
* *[[Scopaeus fuliginosus]]*
* *[[Scopaeus fulvescens]]*
* *[[Scopaeus fulvus]]*
* *[[Scopaeus fusculus]]*
* *[[Scopaeus galapagosus]]*
* *[[Scopaeus galinae]]*
* *[[Scopaeus gangtokensis]]*
* *[[Scopaeus garambanus]]*
* *[[Scopaeus germanus]]*
* *[[Scopaeus gigantulus]]*
* *[[Scopaeus gilensis]]*
* *[[Scopaeus girardi]]*
* *[[Scopaeus girardianus]]*
* *[[Scopaeus gissarensis]]*
* *[[Scopaeus giulianii]]*
* *[[Scopaeus gladifer]]*
* *[[Scopaeus gomyi]]*
* *[[Scopaeus gracilis]]*
* *[[Scopaeus graecus]]*
* *[[Scopaeus grandicollis]]*
* *[[Scopaeus grandipennis]]*
* *[[Scopaeus graskopensis]]*
* *[[Scopaeus gusarovi]]*
* *[[Scopaeus gustavkraatzi]]*
* *[[Scopaeus haemusensis]]*
* *[[Scopaeus heinzi]]*
* *[[Scopaeus hercegovinensis]]*
* *[[Scopaeus hermani]]*
* *[[Scopaeus hiekei]]*
* *[[Scopaeus hispanicus]]*
* *[[Scopaeus hova]]*
* *[[Scopaeus hudsonicus]]*
* *[[Scopaeus hulstaertianus]]*
* *[[Scopaeus humidulus]]*
* *[[Scopaeus humilis]]*
* *[[Scopaeus hyrcanus]]*
* *[[Scopaeus illustris]]*
* *[[Scopaeus imbecillus]]*
* *[[Scopaeus indiensis]]*
* *[[Scopaeus insidiosus]]*
* *[[Scopaeus intermixtus]]*
* *[[Scopaeus interocularis]]*
* *[[Scopaeus iranensis]]*
* *[[Scopaeus jacobsoni]]*
* *[[Scopaeus janaki]]*
* *[[Scopaeus japonicus]]*
* *[[Scopaeus javanus]]*
* *[[Scopaeus jeanneli]]*
* *[[Scopaeus kabakovi]]*
* *[[Scopaeus kasongensis]]*
* *[[Scopaeus kastcheevi]]*
* *[[Scopaeus kaszabianus]]*
* *[[Scopaeus katanganus]]*
* *[[Scopaeus kermanensis]]*
* *[[Scopaeus khnzoriani]]*
* *[[Scopaeus kivuanus]]*
* *[[Scopaeus kivuensis]]*
* *[[Scopaeus klapperichi]]*
* *[[Scopaeus kokodanus]]*
* *[[Scopaeus korelli]]*
* *[[Scopaeus korgei]]*
* *[[Scopaeus kovaci]]*
* *[[Scopaeus kritschi]]* Frisch, 2025
* *[[Scopaeus kurdistanicoides]]*
* *[[Scopaeus kurdistanus]]*
* *[[Scopaeus laevigatus]]*
* *[[Scopaeus laevis]]*
* *[[Scopaeus lamtoensis]]*
* *[[Scopaeus lanceolatus]]*
* *[[Scopaeus latebricola]]*
* *[[Scopaeus laticollis]]*
* *[[Scopaeus latitarsis]]*
* *[[Scopaeus laxus]]*
* *[[Scopaeus leleupi]]*
* *[[Scopaeus leleupianus]]*
* *[[Scopaeus leopoldvillensis]]*
* *[[Scopaeus levasseuri]]*
* *[[Scopaeus ligulifer]]*
* *[[Scopaeus likovskyi]]*
* *[[Scopaeus limbatus]]*
* *[[Scopaeus linearis]]*
* *[[Scopaeus littoralis]]*
* *[[Scopaeus loebli]]*
* *[[Scopaeus longicollis]]*
* *[[Scopaeus longicornis]]*
* *[[Scopaeus longilobatus]]*
* *[[Scopaeus longipennis]]*
* *[[Scopaeus longiusculus]]*
* *[[Scopaeus loxias]]*
* *[[Scopaeus lucidus]]*
* *[[Scopaeus luctuosus]]*
* *[[Scopaeus ludificatorius]]*
* *[[Scopaeus ludificatus]]*
* *[[Scopaeus lugubris]]*
* *[[Scopaeus lungwensis]]*
* *[[Scopaeus luzonicus]]*
* *[[Scopaeus machadoanus]]*
* *[[Scopaeus macilentus]]*
* *[[Scopaeus madagascarensis]]*
* *[[Scopaeus major]]*
* *[[Scopaeus marginalis]]*
* *[[Scopaeus marginatus]]*
* *[[Scopaeus mariae]]*
* *[[Scopaeus masaicus]]*
* *[[Scopaeus mendosus]]*
* *[[Scopaeus menteshensis]]*
* *[[Scopaeus meridioafricanus]]*
* *[[Scopaeus meridioanatolicus]]*
* *[[Scopaeus methneri]]*
* *[[Scopaeus micropterus]]*
* *[[Scopaeus micros]]*
* *[[Scopaeus milkoi]]*
* *[[Scopaeus minimus]]*
* *[[Scopaeus minutoides]]*
* *[[Scopaeus minutulus]]*
* *[[Scopaeus minutus]]*
* *[[Scopaeus mitratus]]*
* *[[Scopaeus mixtus]]*
* *[[Scopaeus moerens]]*
* *[[Scopaeus mollis]]*
* *[[Scopaeus molo]]*
* *[[Scopaeus montalbanensis]]*
* *[[Scopaeus monticola]]*
* *[[Scopaeus moriturus]]*
* *[[Scopaeus muehlei]]*
* *[[Scopaeus muhiensis]]*
* *[[Scopaeus mulongoensis]]*
* *[[Scopaeus mutatus]]*
* *[[Scopaeus myrmecocephalus]]*
* *[[Scopaeus naomii]]*
* *[[Scopaeus nepalensis]]*
* *[[Scopaeus nepalicus]]*
* *[[Scopaeus nevermanni]]*
* *[[Scopaeus nicanor]]*
* *[[Scopaeus nicetius]]*
* *[[Scopaeus nigellus]]*
* *[[Scopaeus niger]]*
* *[[Scopaeus nigerrimus]]*
* *[[Scopaeus nimius]]*
* *[[Scopaeus nipponensis]]*
* *[[Scopaeus nitidiceps]]*
* *[[Scopaeus nitidicollis]]*
* *[[Scopaeus nitidipennis]]*
* *[[Scopaeus nitidulus]]*
* *[[Scopaeus nitidus]]*
* *[[Scopaeus notangulus]]*
* *[[Scopaeus notmani]]*
* *[[Scopaeus obscuripennis]]*
* *[[Scopaeus obscuripes]]*
* *[[Scopaeus obscurus]]*
* *[[Scopaeus ooderes]]*
* *[[Scopaeus opaciceps]]*
* *[[Scopaeus opacicollis]]*
* *[[Scopaeus opacus]]*
* *[[Scopaeus orbiceps]]*
* *[[Scopaeus ornatus]]*
* *[[Scopaeus orophilus]]*
* *[[Scopaeus overlaetianus]]*
* *[[Scopaeus oviceps]]*
* *[[Scopaeus pakistanensis]]*
* *[[Scopaeus palaestinus]]*
* *[[Scopaeus paliferus]]*
* *[[Scopaeus pallens]]*
* *[[Scopaeus paludicola]]*
* *[[Scopaeus parvicornis]]*
* *[[Scopaeus parvipterus]]*
* *[[Scopaeus pauliani]]*
* *[[Scopaeus pauper]]*
* *[[Scopaeus pellionis]]*
* *[[Scopaeus penangensis]]*
* *[[Scopaeus peregrinus]]*
* *[[Scopaeus perroti]]*
* *[[Scopaeus persicus]]*
* *[[Scopaeus persimilis]]*
* *[[Scopaeus philippinensis]]*
* *[[Scopaeus piceolus]]*
* *[[Scopaeus picipes]]*
* *[[Scopaeus poggii]]*
* *[[Scopaeus politus]]*
* *[[Scopaeus ponticus]]*
* *[[Scopaeus portai]]*
* *[[Scopaeus potamus]]*
* *[[Scopaeus proculus]]*
* *[[Scopaeus prolixipennis]]*
* *[[Scopaeus pruinosulus]]*
* *[[Scopaeus pseudomethneri]]*
* *[[Scopaeus puberulus]]*
* *[[Scopaeus pulchellus]]*
* *[[Scopaeus punctatellus]]*
* *[[Scopaeus puncticeps]]*
* *[[Scopaeus punctithorax]]*
* *[[Scopaeus puritanus]]*
* *[[Scopaeus pusilloides]]*
* *[[Scopaeus pusillus]]*
* *[[Scopaeus puthzi]]*
* *[[Scopaeus pygmaeus]]*
* *[[Scopaeus qohrudensis]]*
* *[[Scopaeus quadriceps]]*
* *[[Scopaeus quadripennis]]*
* *[[Scopaeus rambouseki]]*
* *[[Scopaeus reduncus]]*
* *[[Scopaeus ripicola]]*
* *[[Scopaeus rivularis]]*
* *[[Scopaeus rotundiceps]]*
* *[[Scopaeus ruandensis]]*
* *[[Scopaeus rubidus]]*
* *[[Scopaeus rubricollis]]*
* *[[Scopaeus rudis]]*
* *[[Scopaeus ruficollis]]*
* *[[Scopaeus rufulus]]*
* *[[Scopaeus rugegensis]]*
* *[[Scopaeus rugosulus]]*
* *[[Scopaeus ruguliceps]]*
* *[[Scopaeus rulomus]]*
* *[[Scopaeus ruwenzoriensis]]*
* *[[Scopaeus ruziziensis]]*
* *[[Scopaeus ryei]]*
* *[[Scopaeus saaberae]]* Frisch, 2025
* *[[Scopaeus saginellus]]*
* *[[Scopaeus salvini]]*
* *[[Scopaeus sarawakensis]]*
* *[[Scopaeus sareptanus]]*
* *[[Scopaeus satricus]]*
* *[[Scopaeus schawalleri]]*
* *[[Scopaeus schillhammeri]]*
* *[[Scopaeus schuelkei]]*
* *[[Scopaeus schusteri]]*
* *[[Scopaeus scotti]]*
* *[[Scopaeus semicornis]]*
* *[[Scopaeus senegalensis]]*
* *[[Scopaeus sericans]]*
* *[[Scopaeus serriae]]*
* *[[Scopaeus seydeli]]*
* *[[Scopaeus siamensis]]*
* *[[Scopaeus siculus]]*
* *[[Scopaeus signifer]]*
* *[[Scopaeus similaris]]*
* *[[Scopaeus similis]]*
* *[[Scopaeus simillimus]]*
* *[[Scopaeus simulator]]*
* *[[Scopaeus sinaicus]]*
* *[[Scopaeus socotrensis]]*
* *[[Scopaeus solomonensis]]*
* *[[Scopaeus sonoricus]]*
* *[[Scopaeus spathiferus]]*
* *[[Scopaeus spectralis]]*
* *[[Scopaeus spinosophallatus]]*
* *[[Scopaeus stramineus]]*
* *[[Scopaeus subconfusus]]*
* *[[Scopaeus subfasciatus]]*
* *[[Scopaeus subopacus]]*
* *[[Scopaeus subprocerus]]*
* *[[Scopaeus sulcicollis]]*
* *[[Scopaeus sumbaensis]]*
* *[[Scopaeus summicola]]*
* *[[Scopaeus sundaensis]]*
* *[[Scopaeus surdus]]*
* *[[Scopaeus suspectus]]*
* *[[Scopaeus tahitiensis]]*
* *[[Scopaeus tarsalis]]*
* *[[Scopaeus tauricus]]*
* *[[Scopaeus tchapembanus]]*
* *[[Scopaeus testaceipes]]*
* *[[Scopaeus testaceus]]*
* *[[Scopaeus texanus]]*
* *[[Scopaeus thoracicus]]*
* *[[Scopaeus tibialis]]*
* *[[Scopaeus tigreanus]]*
* *[[Scopaeus tonkinensis]]*
* *[[Scopaeus transvaalensis]]*
* *[[Scopaeus trapeziceps]]*
* *[[Scopaeus triangularis]]*
* *[[Scopaeus trifurcatus]]*
* *[[Scopaeus tristis]]*
* *[[Scopaeus truncaticeps]]*
* *[[Scopaeus tsaratananus]]*
* *[[Scopaeus tshuapaensis]]*
* *[[Scopaeus tucumanensis]]*
* *[[Scopaeus tumbaensis]]*
* *[[Scopaeus turkestanicus]]*
* *[[Scopaeus udus]]*
* *[[Scopaeus ulughdaghensis]]*
* *[[Scopaeus uluguruensis]]*
* *[[Scopaeus umbra]]*
* *[[Scopaeus unifasciatus]]*
* *[[Scopaeus vagans]]*
* *[[Scopaeus velutinus]]* Motschulsky, 1858
* *[[Scopaeus ventralis]]*
* *[[Scopaeus versicolor]]*
* *[[Scopaeus vietnamensis]]*
* *[[Scopaeus vilhenai]]*
* *[[Scopaeus vilis]]*
* *[[Scopaeus viriliformis]]*
* *[[Scopaeus virilis]]*
* *[[Scopaeus viriloides]]*
* *[[Scopaeus voltae]]*
* *[[Scopaeus weigeli]]*
* *[[Scopaeus witteanus]]*
* *[[Scopaeus wunderlei]]*
* *[[Scopaeus yoshidai]]*
* *[[Scopaeus zagrosensis]]*

## Referanser


## Kilder
* Silfverberg, H. 1992. *Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae* (Liste over Nordens biller). Helsinki.
* Fauna Europaea (2011) Fauna Europaea Web Service Version 2.4. Utbredelsesdatabase for europeiske dyr. http://www.faunaeur.org/ 

## Eksterne lenker
* 
* Bildegalleri over nordamerikanske arter: http://bugguide.net/node/view/101182/bgpage