Calathus

***Calathus*** er en slekt av billene i delgruppen (underfamilien) Platyninae i familiegruppen løpebiller. Seks arter er registret fra Norge.

## Utseende
Middelsstore (gjerne 7-12 millimeter), avlangt-ovale, svarte løpebiller, pronotum kan være lysere. Oversiden er ofte litt fløyelsaktig halvmatt. Hodet er forholdsvis lite, pronotum rundet med markerte bakhjørner. Dekkvingene er litt rundede med fine punktstriper. Beina er forholdsvis lange, klørne kamtannede.

## Levevis
Mange av artene er vanlige på skogbunnen. Artene i denne slekten ser ut til å være mer tolerante overfor skogmaur enn de fleste andre løpebiller, og kan være tallrike i nærheten av maurtuer.

## Utbredelse

## Systematisk inndeling
Systematikken følger Carabidae of the World. Artene er satt opp alfabetisk.

* ordenen biller,** Coleoptera** Linnaeus, 1758
** underordenen Adephaga Schellenberg, 1806
*** overfamilien løpebiller og vannkalver, **[[Caraboidea]]** Latreille, 1802
**** familien  ** [[løpebiller]]**, Carabidae Latreille, 1802
***** underfamilie Platyninae Bonelli, 1810
****** slekten *Calathus*  
******* underslekt *[[Calathus (Acalathus)]]* - ofte regnet som en egen slekt
******** *[[Calathus advena]]* (LeConte, 1846)
******** *[[Calathus businskyi]]* (Lassalle, 1999)
******** *[[Calathus drolmae]]* Lassalle, 1999
******** *[[Calathus fallax]]* (Semenov, 1889)
******** *[[Calathus langmusiensis]]* (Lassalle, 1999)
******** *[[Calathus luhuoensis]]* Lassalle, 1999
******** *[[Calathus nanpingensis]]* (Lassalle, 1999)
******** *[[Calathus semirufescens]]* Semenov, 1889
******** *[[Calathus validulus]]* Tschitscherine, 1895
******** *[[Calathus wrzecionkoi]]* Lassalle, 1999
******** *[[Calathus xiahensis]]* Lassalle, 1999
******* underslekt *[[Calathus (Amphyginus)]]*
******** *[[Calathus rotundicollis]]* Dejean, 1828
******* underslekt *[[Calathus (Bedelinus)]]*
******** *[[Calathus circumseptus]]* Germar, 1824
******* underslekt *[[Calathus (Calathus)]]*
******** *[[Calathus baeticus]]* Rambur, 1837
******** *[[Calathus bosnicus]]* Ganglbauer, 1891
******** *[[Calathus brevis]]* Gautier des Cottes, 1866
******** *[[Calathus distinguendus]]* Chaudoir, 1846
******** *[[Calathus ellipticus]]* Reitter, 1889
******** *[[Calathus focarilei]]* Schatzmayr, 1947
******** *[[Calathus fracassii]]* Heyden, 1908
******** *[[Calathus fuscipes]]* (Goeze, 1777)
******** *[[Calathus giganteus]]* Dejean, 1828
******** *[[Calathus glabricollis]]* Dejean, 1828
******** *[[Calathus hispanicus]]* Gautier des Cottes, 1866
******** *[[Calathus jakupicensis]]* Gueorgiev, 2008
******** *[[Calathus korax]]* Reitter, 1889
******** *[[Calathus longicollis]]* Motschulsky, 1865
******** *[[Calathus luctuosus]]* (Latreille, 1804)
******** *[[Calathus macedonicus]]* Maran, 1935
******** *[[Calathus malacensis]]* Nègre, 1966
******** *[[Calathus minutus]]* Gautier des Cottes, 1866
******** *[[Calathus mirei]]* Nègre, 1966
******** *[[Calathus montivagus]]* Dejean, 1831
******** *[[Calathus moralesi]]* Nègre, 1966
******** *[[Calathus oertzeni]]* Jeanne & F.Battoni, 1987
******** *[[Calathus oreades]]* Nègre, 1966
******** *[[Calathus pirazzolii]]* Putzeys, 1873
******** *[[Calathus ravasinii]]* G.Müller, 1935
******** *[[Calathus reflexus]]* Schaum, 1858
******** *[[Calathus rubripes]]* Dejean, 1831
******** *[[Calathus sirentensis]]* D'Amore Fracassi, 1908
******** *[[Calathus syriacus]]* Chaudoir, 1863
******** *[[Calathus tombesii]]* F.Battoni, 1976
******** *[[Calathus uniseriatus]]* Vuillefroy, 1866
******** *[[Calathus vivesi]]* Nègre, 1966
******** *[[Calathus vuillefroyi]]* Gautier des Cottes, 1867
******* underslekt *[[Calathus (Iberocalathus)]]*
******** *[[Calathus rotundatus]]* Jacquelin du Val, 1857
******* underslekt *[[Calathus (Lauricalathus)]]*
******** *[[Calathus abaxoides]]* Brullé, 1839
******** *[[Calathus amplius]]* Escalera, 1921
******** *[[Calathus angularis]]* Brullé, 1839
******** *[[Calathus angustulus]]* Wollaston, 1862
******** *[[Calathus appendiculatus]]* Wollaston, 1862
******** *[[Calathus ascendens]]* Wollaston, 1862
******** *[[Calathus auctus]]* Wollaston, 1862
******** *[[Calathus canariensis]]* Harold, 1868
******** *[[Calathus carinatus]]* Brullé, 1839
******** *[[Calathus carvalhoi]]* Serrano & Borges, 1986
******** *[[Calathus ciliatus]]* Wollaston, 1862
******** *[[Calathus cognatus]]* Wollaston, 1864
******** *[[Calathus colasianus]]* Mateu, 1970
******** *[[Calathus complanatus]]* Dejean, 1828
******** *[[Calathus depressus]]* Brullé, 1836
******** *[[Calathus fimbriatus]]* Wollaston, 1858
******** *[[Calathus freyi]]* Colas, 1941
******** *[[Calathus gomerensis]]* Colas, 1943
******** *[[Calathus laureticola]]* Wollaston, 1865
******** *[[Calathus marcellae]]* Colas, 1943
******** *[[Calathus pecoudi]]* Colas, 1938
******** *[[Calathus rectus]]* Wollaston, 1862
******** *[[Calathus rufocastaneus]]* Wollaston, 1862
******** *[[Calathus spretus]]* Wollaston, 1862
******** *[[Calathus vividus]]* (Fabricius, 1801)
******* underslekt *[[Lindrothius|Calathus (Lindrothius)]]* Kurnakov, 1961 - ofte plassert som en egen slekt
******** *[[Calathus aeneocupreus]]* (Heinz, 1971)
******** *[[Calathus aequistriatus]]* Kurnakov, 1961
******** *[[Calathus caucasicus]]* Chaudoir, 1846
******** *[[Calathus grandiceps]]* (Kurnakov, 1961)
******** *[[Calathus horsti]]* (Reitter, 1888)
******** *[[Calathus laticaudis]]* (Kurnakov, 1961)
******** *[[Calathus mandibularis]]* (Kurnakov, 1961)
******** *[[Calathus praestans]]* (Heyden, 1885)
******** *[[Calathus pseudopraestans]]* (Kurnakov, 1961)
******** *[[Calathus recticaudis]]* (Kurnakov, 1961)
******** *[[Calathus robustus]]* (Kurnakov, 1961)
******** *[[Calathus sotshiensis]]* (Zamotajlov, 1997)
******** *[[Calathus stricticaudis]]* (Kurnakov, 1961)
******** *[[Calathus subpraestans]]* (Kurnakov, 1961)
******* underslekt *[[Calathus (Neocalathus)]]*
******** *[[Calathus albanicus]]* Apfelbeck, 1906
******** *[[Calathus ambiguus]]* (Paykull, 1790)
******** *[[Calathus asturiensis]]* Vuillefroy, 1866
******** *[[Calathus calceus]]* Ball and Nègre, 1972
******** *[[Calathus cinctus]]* Motschulsky, 1850
******** *[[Calathus erratus]]* (C.R.Sahlberg, 1827)
******** *[[Calathus extensicollis]]* Putzeys, 1873
******** *[[Calathus gonzalezi]]* Mateu, 1956
******** *[[Calathus granatensis]]* Vuillefroy, 1866
******** *[[Calathus gregarius]]* (Say, 1823)
******** *[[Calathus ingratus]]* Dejean, 1828
******** *[[Calathus lundbladi]]* Colas, 1938
******** *[[Calathus melanocephalus]]* (Linnaeus, 1758)
******** *[[Calathus metallicus]]* Dejean, 1828
******** *[[Calathus micropterus]]* (Duftschmid, 1812)
******** *[[Calathus mollis]]* (Marsham, 1802)
******** *[[Calathus opaculus]]* Leconte, 1854
******** *[[Calathus peltatus]]* Kolenati, 1845
******** *[[Calathus peropacus]]* Casey, 1920
******** *[[Calathus ruficollis]]* Dejean, 1828
******** *[[Calathus simplicicollis]]* Wollaston, 1862
******** *[[Calathus solieri]]* Bassi, 1834
******** *[[Calathus subfuscus]]* Wollaston, 1865
******** *[[Calathus vicenteorum]]* Schatzmayr, 1937
******* underslekt *[[Calathus (Trichocalathus)]]*
******** *[[Calathus obliteratus]]* Wollaston, 1865
******** *[[Calathus pilosipennis]]* Machado, 1992
******** *[[Calathus refleximargo]]* Machado, 1992
### Arter ikke plassert til underslekt
* *[[Calathus acuticollis]]* Putzeys, 1873 c g
* *[[Calathus aethiopicus]]* Alluaud, 1918 c g
* *[[Calathus aethiops]]* Alluaud, 1937 c g
* *[[Calathus afroalpinus]]* Lassalle, 2016 c g
* *[[Calathus algens]]* Andrewes, 1934 c g
* *[[Calathus alternans]]* Faldermann, 1836 c g
* *[[Calathus ambigens]]* Bates, 1891 i c g
* *[[Calathus anatolicus]]* Jedlicka, 1969 c g
* *[[Calathus anistschenkoi]]* Berlov & Ippolitova, 2005 c g
* *[[Calathus ankoberensis]]* Lassalle, 2016 c g
* *[[Calathus annapurnae]]* J.Schmidt, 1999 c g
* *[[Calathus arcuatus]]* Gautier des Cottes, 1870 c g
* *[[Calathus atitari]]* Novoa, 1999 c g
* *[[Calathus aztec]]* Ball and Nègre, 1972 i c g
* *[[Calathus baleensis]]* Lassalle, 2016 c g
* *[[Calathus balli]]* Novoa; Gañan & Baselga, 2015 c g
* *[[Calathus bolivari]]* Nègre, 1970 i c g
* *[[Calathus busii]]* F.Battoni, 1984 c g
* *[[Calathus carballalae]]* Novoa & Gañán, 2014
* *[[Calathus casalei]]* F.Battoni, 1986 c g
* *[[Calathus chioriae]]* Lassalle, 2016 c g
* *[[Calathus chokeensis]]* Lassalle, 2016 c g
* *[[Calathus clauseni]]* Ball and Nègre, 1972 i c g
* *[[Calathus deliae]]* Morvan, 1999 c g
* *[[Calathus deplanatus]]* Chaudoir, 1843 c g
* *[[Calathus deyrollei]]* Gautier des Cottes, 1870 c g
* *[[Calathus dubaulti]]* Lassalle, 2016 c g
* *[[Calathus durango]]* Ball and Nègre, 1972 i c g
* *[[Calathus elpis]]* Ortuño & Arillo, 2009 c g
* *[[Calathus erwini]]* Ball and Nègre, 1972 i c g
* *[[Calathus erzeliki]]* Schweiger, 1977 c g
* *[[Calathus femoralis]]* Chaudoir, 1846 c g
* *[[Calathus gansuensis]]* (Jedlicka, 1937) c g
* *[[Calathus gelascens]]* Andrewes, 1934 c g
* *[[Calathus gugheensis]]* Basilewsky, 1953 c g
* *[[Calathus gunaensis]]* Lassalle, 2016 c g
* *[[Calathus heinertzi]]* Deuve; Lassalle & Queinnec, 1985 c g
* *[[Calathus heinzianus]]* F.Battoni, 1986 c g
* *[[Calathus himalayae]]* Bates, 1891 c g
* *[[Calathus holzschuhi]]* Kirschenhofer, 1990 c g
* *[[Calathus hyrcanus]]* Heinz, 1970 c g
* *[[Calathus juan]]* Novoa & Gañán, 2014 c g
* *[[Calathus kebedei]]* Novoa; Gañan & Baselga, 2015 c g
* *[[Calathus kirschenhoferi]]* F.Battoni, 1982 c g
* *[[Calathus kirschenhoferianus]]* F.Battoni, 1986 c g
* *[[Calathus kollari]]* Putzeys, 1873 c g
* *[[Calathus kryzhanovskii]]* J.Schmidt, 1999 c g
* *[[Calathus leechi]]* Ball and Nègre, 1972 i c g
* *[[Calathus leptodactylus]]* Putzeys, 1873 c g
* *[[Calathus libanensis]]* Putzeys, 1873 c g
* *[[Calathus lissoderus]]* Putzeys, 1873 c g
* *[[Calathus manasluensis]]* J.Schmidt, 1999 c g
* *[[Calathus marmoreus]]* Ball and Nègre, 1972
* *[[Calathus mcclevei]]* Shpeley & Devender, 2022
* *[[Calathus mexicanus]]* Chaudoir, 1837 i c g
* *[[Calathus montanellus]]* Heinz, 2000 c g
* *[[Calathus montanus]]* Alluaud, 1937 c g
* *[[Calathus muchei]]* Jedlicka, 1961 c g
* *[[Calathus nitidus]]* Basilewsky, 1957 c g
* *[[Calathus ollivieri]]* Lassalle, 2016 c g
* *[[Calathus opacus]]* Lucas, 1846 c g
* *[[Calathus ordinatus]]* Gautier des Cottes, 1870 c g
* *[[Calathus oreobius]]* Alluaud, 1937 c g
* *[[Calathus orthomoides]]* Alluaud, 1932 c g
* *[[Calathus ovipennis]]* Putzeys, 1873 i c g
* *[[Calathus parvicollis]]* Fairmaire, 1882 c g
* *[[Calathus patrizii]]* Basilewsky, 1953 c g
* *[[Calathus pommeranzi]]* J.Schmidt, 1999 c g
* *[[Calathus potosi]]* Ball and Nègre, 1972 i c g
* *[[Calathus queinneci]]* Lassalle, 2016 c g
* *[[Calathus ras]]* Basilewsky, 1957 c g
* *[[Calathus reebae]]* Lassalle, 2016 c g
* *[[Calathus reflexicollis]]* Faldermann, 1839 c g
* *[[Calathus relictus]]* J.Schmidt & Tian, 2013 c g
* *[[Calathus rhaticus]]* Antoine, 1941 c g
* *[[Calathus roccai]]* F.Battoni, 1984 c g
* *[[Calathus rotgeri]]* Ball and Nègre, 1972 i c g
* *[[Calathus scotti]]* Alluaud, 1937 c g
* *[[Calathus scottianus]]* Basilewsky, 1953 c g
* *[[Calathus semisericeus]]* Fairmaire, 1879 c g
* *[[Calathus shoanus]]* Alluaud, 1932 c g
* *[[Calathus simienensis]]* Basilewsky, 1957 c g
* *[[Calathus suffuscus]]* Andrewes, 1934 c g
* *[[Calathus theodori]]* Ancey, 1882 c g
* *[[Calathus tigrinus]]* Shpeley & Devender, 2022
* *[[Calathus trapezicollis]]* Alluaud, 1937 c g
* *[[Calathus vagestriatus]]* Fairmaire, 1882 c g
* *[[Calathus vignatagliantii]]* F.Battoni, 1986 c g
* *[[Calathus whiteheadi]]* Ball and Nègre, 1972 i c g
* *[[Calathus wittmerianus]]* Deuve; Lassalle & Queinnec, 1985 c g
* *[[Calathus wolloensis]]* Lassalle, 2016 c g
* *[[Calathus zabroides]]* Putzeys, 1873 c g

## Referanser


## Eksterne lenker
* 
* ''Calathus'' - Fauna Europaea - engelskspråklig
* ''Calathus'' – Database Carabidae of the World – engelskspråklig
* Norsk Entomologisk forening