Dichotomius

***Dichotomius*** er en artsrik slekt av biller som hører til familien skarabider (Scarabaeidae) i gruppen skarabider (Scarabaeoidea). Artene, som lever av kumøkk, kan minne om små neshornbiller, men er mindre, mer tettbygd og hvelvet, og dekkvingene har tydelige lengdestriper.

## Utseende
Middelsstore, oftest svarte, blanke skarabider. Kroppen er kraftig og hvelvet. Slekten minner om slekten *[[Heliocopris]]* fra den gamle verden, men disse artene blir ikke like store som *Heliocopris*-artene. Hannen har lange horn på hodet og knøler ved fremkanten av pronotum.

## Levevis
Disse billene finnes helst i tilknytning til gjødsel. Den graver dype sjakter under en kuruke og graver ned store mengder møkk som de legger egg på. Hannen graver gangene, hunnen arbeider møkka til glatte kuler som hun legger egg på. Hun passer disse i rundt fire måneder mens larvene utvikler seg. De voksne billene ses helst vår og høst.

## Utbredelse
Slekten er utbredt i Sør- og Mellom-Amerika og det sørlige Nord-Amerika.

## Systematisk plassering

* Ordenen biller, Coleoptera
** Underordenen Polyphaga
*** Overfamilien skarabider, Scarabaeoidea
**** Familien skarabider, (Scarabaeidae) Latreille, 1806
***** Underfamilien gjødselgravere (Scarabaeinae) Latreille, 1806
****** Slektsgruppen Coprini
******* Slekten *Dichotomius* Hope, 1838
******** *[[Dichotomius acuticornis]]*
******** *[[Dichotomius adrastus]]*
******** *[[Dichotomius affinis]]*
******** *[[Dichotomius agenor]]*
******** *[[Dichotomius agesilaus]]*
******** *[[Dichotomius alutaceus]]*
******** *[[Dichotomius alyattes]]*
******** *[[Dichotomius amicitiae]]*
******** *[[Dichotomius amplicollis]]*
******** *[[Dichotomius annae]]*
******** *[[Dichotomius anthrax]]*
******** *[[Dichotomius apicalis]]*
******** *[[Dichotomius ascanius]]*
******** *[[Dichotomius assifer]]*
******** *[[Dichotomius batesi]]*
******** *[[Dichotomius bechynei]]*
******** *[[Dichotomius belus]]*
******** *[[Dichotomius bicornis]]*
******** *[[Dichotomius bicuspis]]*
******** *[[Dichotomius bitiensis]]*
******** *[[Dichotomius boreus]]* (Olivier, 1789)
******** *[[Dichotomius borgmeieri]]*
******** *[[Dichotomius bos]]*
******** *[[Dichotomius bosqui]]*
******** *[[Dichotomius bucki]]*
******** *[[Dichotomius buqueti]]*
******** *[[Dichotomius calcaratus]]*
******** *[[Dichotomius camargoi]]*
******** *[[Dichotomius camposeabrai]]*
******** *[[Dichotomius carbonarius]]*
******** *[[Dichotomius carinatus]]*
******** *[[Dichotomius carolinus]]*
******** *[[Dichotomius centralis]]*
******** *[[Dichotomius chamuco]]* Solís & Kohlmann, 2025
******** *[[Dichotomius coenosus]]*
******** *[[Dichotomius colonicus]]*
******** *[[Dichotomius comarapensis]]*
******** *[[Dichotomius conicollis]]* Blanchard, 1846
******** *[[Dichotomius costaricensis]]*
******** *[[Dichotomius cotopaxi]]*
******** *[[Dichotomius crinicollis]]*
******** *[[Dichotomius cuprinus]]*
******** *[[Dichotomius dahli]]*
******** *[[Dichotomius danieli]]*
******** *[[Dichotomius depressicollis]]*
******** *[[Dichotomius deyrollei]]*
******** *[[Dichotomius diabolicus]]*
******** *[[Dichotomius divisor]]* Moura *et al*., 2025
******** *[[Dichotomius dynastus]]*
******** *[[Dichotomius enrietti]]*
******** *[[Dichotomius eucranioides]]*
******** *[[Dichotomius fallax]]*
******** *[[Dichotomius favi]]*
******** *[[Dichotomius femoratus]]*
******** *[[Dichotomius fimbriatus]]*
******** *[[Dichotomius fissiceps]]*
******** *[[Dichotomius fissus]]*
******** *[[Dichotomius fonsecae]]*
******** *[[Dichotomius fortestriatus]]*
******** *[[Dichotomius gamboaensis]]*
******** *[[Dichotomius geminatus]]*
******** *[[Dichotomius glaucus]]*
******** *[[Dichotomius globulus]]*
******** *[[Dichotomius guaranii]]*
******** *[[Dichotomius haroldi]]*
******** *[[Dichotomius hempeli]]*
******** *[[Dichotomius horridus]]*
******** *[[Dichotomius horvathi]]*
******** *[[Dichotomius imitator]]*
******** *[[Dichotomius inachoides]]*
******** *[[Dichotomius inachus]]*
******** *[[Dichotomius inflaticollis]]*
******** *[[Dichotomius ingens]]*
******** *[[Dichotomius interstitialis]]*
******** *[[Dichotomius irinus]]*
******** *[[Dichotomius joelus]]*
******** *[[Dichotomius laevicollis]]*
******** *[[Dichotomius larseni]]* Arias-Buriticá & Vaz-de-Mello, 2024
******** *[[Dichotomius latistriatus]]*
******** *[[Dichotomius longiceps]]*
******** *[[Dichotomius lucasi]]*
******** *[[Dichotomius lucifer]]* Solís & Kohlmann, 2025
******** *[[Dichotomius luctuosioides]]*
******** *[[Dichotomius luctuosus]]*
******** *[[Dichotomius lycas]]*
******** *[[Dichotomius machadoi]]*
******** *[[Dichotomius mamillatus]]*
******** *[[Dichotomius maya]]*
******** *[[Dichotomius melzeri]]*
******** *[[Dichotomius micans]]*
******** *[[Dichotomius missionus]]*
******** *[[Dichotomius monstrosus]]*
******** *[[Dichotomius mormon]]* (Ljungh, 1799)
******** *[[Dichotomius motai]]*
******** *[[Dichotomius mundus]]*
******** *[[Dichotomius muticus]]*
******** *[[Dichotomius mysticus]]*
******** *[[Dichotomius nemoricola]]*
******** *[[Dichotomius nevermanni]]*
******** *[[Dichotomius nimuendaju]]*
******** *[[Dichotomius nisus]]*
******** *[[Dichotomius nitidissimus]]*
******** *[[Dichotomius nobilis]]*
******** *[[Dichotomius nutans]]*
******** *[[Dichotomius ocellatopunctatus]]*
******** *[[Dichotomius ohausi]]* Luederwaldt, 1923
******** *[[Dichotomius opacipennis]]*
******** *[[Dichotomius opalescens]]*
******** *[[Dichotomius paraguayanus]]*
******** *[[Dichotomius parcepunctatus]]*
******** *[[Dichotomius pauloensis]]*
******** *[[Dichotomius pelamon]]* Harold, 1869
******** *[[Dichotomius pisuicas]]* Solís & Kohlmann, 2025
******** *[[Dichotomius planicollis]]*
******** *[[Dichotomius podalirius]]* Felsche, 1901
******** *[[Dichotomius prietoi]]* Martínez & Martínez, 1981
******** *[[Dichotomius problematicus]]*
******** *[[Dichotomius protectus]]*
******** *[[Dichotomius provisorius]]*
******** *[[Dichotomius punctatostriatus]]*
******** *[[Dichotomius puncticollis]]*
******** *[[Dichotomius punctulatipennis]]*
******** *[[Dichotomius pygidialis]]*
******** *[[Dichotomius quadraticeps]]*
******** *[[Dichotomius quadrilobatus]]* Chamorro, Lopera-Toro & Rossini, 2021
******** *[[Dichotomius quadrinodosus]]*
******** *[[Dichotomius quinquedens]]*
******** *[[Dichotomius quinquelobatus]]*
******** *[[Dichotomius reclinatus]]*
******** *[[Dichotomius ribeiroi]]*
******** *[[Dichotomius robustus]]*
******** *[[Dichotomius rodrigoi]]*
******** *[[Dichotomius rotundatus]]*
******** *[[Dichotomius rotundigena]]*
******** *[[Dichotomius rugatus]]*
******** *[[Dichotomius rugosicollis]]*
******** *[[Dichotomius rugosipennis]]*
******** *[[Dichotomius sagittarius]]*
******** *[[Dichotomius satanas]]*
******** *[[Dichotomius schiffleri]]*
******** *[[Dichotomius semiaeneus]]*
******** *[[Dichotomius semisquamosus]]*
******** *[[Dichotomius sericeus]]*
******** *[[Dichotomius sexdentatus]]*
******** *[[Dichotomius simplex]]*
******** *[[Dichotomius simplicicornis]]*
******** *[[Dichotomius simulans]]*
******** *[[Dichotomius singularis]]*
******** *[[Dichotomius smaragdinus]]*
******** *[[Dichotomius spadiceus]]*
******** *[[Dichotomius speciosus]]*
******** *[[Dichotomius subaeneus]]*
******** *[[Dichotomius superbus]]*
******** *[[Dichotomius talaus]]* Erichson, 1847
******** *[[Dichotomius telamon]]*
******** *[[Dichotomius texanus]]*
******** *[[Dichotomius torulosus]]*
******** *[[Dichotomius triangulariceps]]*
******** *[[Dichotomius triquetrus]]*
******** *[[Dichotomius tristis]]*
******** *[[Dichotomius uacari]]* Moura *et al*., 2025
******** *[[Dichotomius validipilosus]]*
******** *[[Dichotomius vidaurrei]]* Nunes & Vaz-De-Mello,2013
******** *[[Dichotomius virescens]]*
******** *[[Dichotomius worontzowi]]*
******** *[[Dichotomius yucatanus]]*
******** *[[Dichotomius zikani]]*
******** *[[Dichotomius ziliolii]]*

## Referanser


## Kilder
* Scarabaeinae del Nuovo Mondo - ''Dichotomius''

## Eksterne lenker
*