Megarthrus

***Megarthrus*** er en gruppe (slekt) av små biller som er vanlige i kompost og annet råtnende plantemateriale. De hører til gruppen bredkortvinger (Proteininae) i den svært artsrike familien kortvinger (Staphylinidae). Det er kjent sju arter fra Norge.

## Utseende
Små (vanligvis 1 – 2&nbsp;mm), brede, litt flate biller, gulaktige, brunaktige eller svarte. **Hodet** er bredere enn langt med små fasettøyne. Antennene ender i en svakt antydet kølle. **Brystskjoldet** (pronotum) er bredere enn langt, sidene er innskåret ved bakhjørnene. **Dekkvingene** er ganske lange, mye lengre enn brystskjoldet, og dekker de fremste ryggleddene. **Bakkroppen** er bred og noe flattrykt. **Beina** er korte og ganske kraftige, leggene (*tibiae*) er noe utvidet særlig hos hannene.

## Levevis
Artene finnes ofte tallrikt i kompost og på sopp. Disse billene flyr godt og man kan finne dem svermende i solskinn.

## Systematisk inndeling / norske arter

* Ordenen biller, **Coleoptera**
** Underordenen Polyphaga
*** Overfamilien åtsel- og rovbiller, **Staphylinoidea**
**** Familien kortvinger (Staphylinidae)
***** Underfamilien **bredkortvinger, Proteininae** Erichson, 1839
****** Stammen Proteinini Erichson, 1839
******* Slekten ***Megarthrus*** Curtis, 1829
******** ***[[Megarthrus denticollis]]*** (Beck, 1817)
******** ***[[Megarthrus depressus]]*** (Paykull, 1789)
******** ***[[Megarthrus fennicus]]*** Lahtinen, 1938
******** ***[[Megarthrus nigrinus]]*** J. Sahlberg, 1876
******** ***[[Megarthrus nitidulus]]*** Kraatz, 1858
******** ***[[Megarthrus sinuatocollis]]*** (Lacordaire, 1835)
******** ***[[Megarthrus strandi]]*** Scheerpeltz, 1931

### Liste over arter som ikke forekommer i Norge
* *[[Megarthrus abessinus]]* Bernhauer, 1931
* *[[Megarthrus adelphus]]* Bierig, 1940
* *[[Megarthrus africanus]]* Eichelbaum, 1913
* *[[Megarthrus aino]]* Cuccodoro, 1996
* *[[Megarthrus alatorreorum]]* Rodríguez & Navarrete-Heredia, 2015
* *[[Megarthrus alesi]]* Cuccodoro, 2003
* *[[Megarthrus alienus]]* Cuccodoro, 1998
* *[[Megarthrus alticola]]* Cameron, 1924
* *[[Megarthrus altivagans]]* Bernhauer, 1929
* *[[Megarthrus americanus]]* Sachse, 1852
* *[[Megarthrus andinus]]* López-García, Méndez-Rojas & Navarrete-Heredia, 2011
* *[[Megarthrus anggiensis]]* Cuccodoro, 1998
* *[[Megarthrus angulicollis]]* Mäklin, 1852
* *[[Megarthrus antennalis]]* Cameron, 1941
* *[[Megarthrus apicicornis]]* Cameron, 1950
* *[[Megarthrus arcuatus]]* Hatch, 1957
* *[[Megarthrus ashei]]* Cuccodoro & Löbl, 1996
* *[[Megarthrus atratus]]* Mäklin, 1852
* *[[Megarthrus auricola]]* Cuccodoro, 1995
* *[[Megarthrus balensis]]* Cuccodoro, 1999
* *[[Megarthrus baliemensis]]* Cuccodoro, 1998
* *[[Megarthrus bantu]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus basicornis]]* Fauvel, 1904
* *[[Megarthrus basilewskyi]]* Fagel, 1957
* *[[Megarthrus bellevoyei]]* Saulcy, 1862
* *[[Megarthrus bierigi]]* Cuccodoro, 2011
* *[[Megarthrus bimaculatus]]* Fauvel, 1904
* *[[Megarthrus birmanus]]* Fauvel, 1895
* *[[Megarthrus borealis]]* Cuccodoro & Löbl, 1996
* *[[Megarthrus budai]]* Liu & Cuccodoro, 2020
* *[[Megarthrus calcaratus]]* Coiffait, 1977
* *[[Megarthrus cavianae]]* Rodriguez, Navarrete-Heredia & Arriaga-Varela, 2020
* *[[Megarthrus chiapas]]* Rodríguez, Navarrete-Heredia, Arriaga-Varela & Cuccodoro, 2020
* *[[Megarthrus chinese]]* Li & Jingke, 1993
* *[[Megarthrus chobauti]]* Fauvel, 1902
* *[[Megarthrus chujiao]]* Liu & Cuccodoro, 2020
* *[[Megarthrus clarkei]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus con]]* Cuccodoro, 2011
* *[[Megarthrus conformis]]* K.Sawada, 1962
* *[[Megarthrus congoensis]]* Cameron, 1950
* *[[Megarthrus conspirator]]* Cuccodoro, 1996
* *[[Megarthrus constrictus]]* Cuccodoro, 1996
* *[[Megarthrus convexus]]* Sharp, 1874
* *[[Megarthrus coreanus]]* Kim, Myoung Hee & Cuccodoro, 2011
* *[[Megarthrus corticalis]]* Sharp, 1889
* *[[Megarthrus danieli]]* Cuccodoro, 1999
* *[[Megarthrus dentatus]]* Coiffait, 1977
* *[[Megarthrus dentipes]]* Bernhauer, 1938
* *[[Megarthrus dissymetricus]]* Coiffait, 1977
* *[[Megarthrus dominicae]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus elevatus]]* Coiffait, 1977
* *[[Megarthrus excisus]]* J.L.LeConte, 1863
* *[[Megarthrus fakir]]* Cuccodoro, 2003
* *[[Megarthrus falasha]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus festivus]]* Cuccodoro, 2011
* *[[Megarthrus fijianus]]* Cuccodoro, 1998
* *[[Megarthrus flavolimbatus]]* Cameron, 1924
* *[[Megarthrus flavosignatus]]* Bierig, 1940
* *[[Megarthrus geginati]]* Cuccodoro, 2010
* *[[Megarthrus gigas]]* Fagel, 1957
* *[[Megarthrus globulus]]* Cuccodoro, 2011
* *[[Megarthrus gressitti]]* Cuccodoro, 1998
* *[[Megarthrus harennaensis]]* Cuccodoro, 1999
* *[[Megarthrus heise]]* Zhang, Cuccodoro, Chen & Liu, 2021
* *[[Megarthrus hemipterus]]* (Illiger, 1794)
* *[[Megarthrus horticola]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus hutu]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus impressicollis]]* Eppelsheim, 1893
* *[[Megarthrus inaequalis]]* Bierig, 1940
* *[[Megarthrus incubifer]]* Cuccodoro, 1996
* *[[Megarthrus integricollis]]* Coiffait, 1977
* *[[Megarthrus ivani]]* Cuccodoro, 2003
* *[[Megarthrus japonicus]]* Sharp, 1874
* *[[Megarthrus kamerunensis]]* Bernhauer, 1942
* *[[Megarthrus kurbatovi]]* Cuccodoro, 2018
* *[[Megarthrus kuscheli]]* Cuccodoro, 1998
* *[[Megarthrus lanka]]* Cuccodoro & Zhiping Liu, 2016
* *[[Megarthrus lisae]]* Cuccodoro, 2011
* *[[Megarthrus loebli]]* Cuccodoro, 2018
* *[[Megarthrus longicornis]]* Wollaston, 1854
* *[[Megarthrus machu]]* Cuccodoro, 2011
* *[[Megarthrus magnicaudatus]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus magnificus]]* Cuccodoro, 2011
* *[[Megarthrus mahnerti]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus major]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus malaisei]]* Scheerpeltz, 1965
* *[[Megarthrus mammiger]]* Bierig, 1940
* *[[Megarthrus maniwaata]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus maronitus]]* Fagel, 1968
* *[[Megarthrus martensi]]* Coiffait, 1982
* *[[Megarthrus mastiger]]* Bierig, 1940
* *[[Megarthrus merabet]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus metanas]]* Cuccodoro, 2011
* *[[Megarthrus minor]]* Coiffait, 1977
* *[[Megarthrus mirabilis]]* Cuccodoro, 2011
* *[[Megarthrus montanus]]* K.Sawada, 1962
* *[[Megarthrus monticola]]* Cameron, 1942
* *[[Megarthrus mukankundiyeorum]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus mwami]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus nanus]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus narendrani]]* Cuccodoro & Zhiping Liu, 2016
* *[[Megarthrus negus]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus newtoni]]* Cuccodoro & Löbl, 1996
* *[[Megarthrus nigerrima]]* Cameron, 1941
* *[[Megarthrus nilgiriensis]]* Cuccodoro & Zhiping Liu, 2016
* *[[Megarthrus niloticus]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus notabilis]]* Cameron, 1941
* *[[Megarthrus occidentalis]]* Cuccodoro & Löbl, 1996
* *[[Megarthrus octopus]]* Cuccodoro, 2011
* *[[Megarthrus ogloblini]]* Bruch, 1940
* *[[Megarthrus oromo]]* Cuccodoro, 1999
* *[[Megarthrus ovalis]]* Cameron, 1950
* *[[Megarthrus panga]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus parallelus]]* Sharp, 1874
* *[[Megarthrus pecki]]* Cuccodoro & Löbl, 1996
* *[[Megarthrus peckorum]]* Cuccodoro, 1998
* *[[Megarthrus phoenix]]* Cuccodoro, 2011
* *[[Megarthrus pictus]]* Motschulsky, 1845
* *[[Megarthrus ping]]* Cuccodoro, 2011
* *[[Megarthrus primus]]* Cuccodoro, 1995
* *[[Megarthrus prosseni]]* Schatzmayr, 1904
* *[[Megarthrus ras]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus riedeli]]* Cuccodoro, 1998
* *[[Megarthrus rougemonti]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus rufomarginatus]]* Cameron, 1914
* *[[Megarthrus saddu]]* Cuccodoro, 2003
* *[[Megarthrus sawadai]]* Cuccodoro, 1996
* *[[Megarthrus scotti]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus scriptus]]* Sharp, 1889
* *[[Megarthrus sculpticollis]]* Coiffait, 1982
* *[[Megarthrus selenitus]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus septempunctatus]]* Champion, 1925
* *[[Megarthrus serrula]]* Wollaston, 1865
* *[[Megarthrus sexpunctatus]]* Cameron, 1941
* *[[Megarthrus shibatai]]* K.Sawada, 1962
* *[[Megarthrus simienensis]]* Fagel, 1957
* *[[Megarthrus smetanai]]* Cuccodoro & Löbl, 1996
* *[[Megarthrus solitarius]]* Sharp, 1887
* *[[Megarthrus spathuliformis]]* Assing & Wunderle, 1999
* *[[Megarthrus spinosus]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus splendidus]]* Cuccodoro, 2011
* *[[Megarthrus stercorarius]]* Mulsant & Rey, 1878
* *[[Megarthrus stylifer]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus sumatrensis]]* Cameron, 1928
* *[[Megarthrus tac]]* Cuccodoro, 2011
* *[[Megarthrus taiwanus]]* Cuccodoro, 2011
* *[[Megarthrus tibialis]]* Coiffait, 1977
* *[[Megarthrus tic]]* Cuccodoro, 2011
* *[[Megarthrus trisinuatus]]* Cameron, 1924
* *[[Megarthrus twa]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus uhligi]]* Cuccodoro & Löbl, 1997
* *[[Megarthrus umbonatus]]* Fauvel, 1895
* *[[Megarthrus vanschuytbroecki]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus vastus]]* Wendeler, 1926
* *[[Megarthrus watutsi]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus wayqecha]]* Pérez, Rodríguez & Asenjo, 2020
* *[[Megarthrus wittei]]* Cameron, 1950
* *[[Megarthrus wollastoni]]* Cuccodoro & Löbl, 1997
* *[[Megarthrus yeti]]* Cuccodoro, 2003
* *[[Megarthrus zekorum]]* Cuccodoro & Löbl, 1997
* *[[Megarthrus zerchei]]* Cuccodoro & Löbl, 1997
* *[[Megarthrus zulu]]* Cuccodoro & Löbl, 1995
* *[[Megarthrus zunilensis]]* Sharp, 1887

## Kilder
* Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (Liste over Nordens biller). Helsinki. ISBN 951-96510-0-4
* Fauna Europaea Web Service Version 2.4 (2011) Utbredelsesdatabase for europeiske dyr http://www.faunaeur.org/ 

## Eksterne lenker
* 
* Bilder av *Megarthrus depressus*: http://www.flickr.com/photos/coleoptera-us/4763964937/ og https://web.archive.org/web/20080925124815/http://www.colpolon.biol.uni.wroc.pl/megarthrus%20depressus.htm