Ataenius

* Se systematisk inndeling

***Ataenius*** er en artsrik slekt av biller som hører til familien skarabider (Scarabaeidae).

## Utseende
Små (1,5 -7 millimeter), avlange, blanke, mørke gjødselbiller. Hodet er stort og bredt, noe hvelvet, gjerne innbuktet foran.

## Levevis
Disse billene har larver som utvikler seg i jorda, der de etter på gressrøtter. De kan gjøre en del skade på golfbaner og lignende 

## Utbredelse
Slekten er utbredt i alle verdensdeler bortsett fra Antarktis.

## Systematisk inndeling

* Ordenen biller, Coleoptera
** Underordenen Polyphaga
*** Overfamilien skarabider, Scarabaeoidea
**** Familien skarabider, (Scarabaeidae) Latreille, 1806
***** Underfamilien Gjødselbiller, Aphodiinae Leach, 1815
****** Stammen Eupariini Schmidt, 1910 
******* Slekten *Ataenius* Harold, 1867
### Liste over artene
* *[[Ataenius abancay]]* Stebnicka, 2005
* *[[Ataenius abditus]]* (Haldeman, 1848)
* *[[Ataenius abdominalis]]* Petrovitz, 1973
* *[[Ataenius aciculus]]* Hinton, 1937
* *[[Ataenius acutulus]]* Schmidt, 1913
* *[[Ataenius aequalis]]* Harold, 1880
* *[[Ataenius aequatorialis]]* Petrovitz, 1961
* *[[Ataenius africanus]]* Endrödi, 1967
* *[[Ataenius alternatus]]* (Melsheimer, 1845)
* *[[Ataenius ambaritae]]* Stebnicka, 1988
* *[[Ataenius angulatus]]* Petrovitz, 1964
* *[[Ataenius annanus]]* Stebnicka & Howden, 1997
* *[[Ataenius aphodiiformis]]* Bordat & Théry, 2012
* *[[Ataenius apicalis]]* Hinton, 1937
* *[[Ataenius arenosus]]* Harold, 1868
* *[[Ataenius argillaceus]]* Schmidt, 1916
* *[[Ataenius arrowi]]* Hinton, 1936
* *[[Ataenius asper]]* Petrovitz, 1975
* *[[Ataenius atramentarius]]* (Erichson, 1847)
* *[[Ataenius attenuator]]* Harold, 1874
* *[[Ataenius australasiae]]* Boheman, 1858
* *[[Ataenius australis]]* Harold, 1875
* *[[Ataenius balthasari]]* Petrovitz, 1973
* *[[Ataenius barberi]]* Cartwright, 1974
* *[[Ataenius basiceps]]* Lea, 1923
* *[[Ataenius beattyi]]* Chapin, 1940
* *[[Ataenius benarabyensis]]* Stebnicka & Howden, 1997
* *[[Ataenius bicolor]]* Petrovitz, 1963
* *[[Ataenius biroi]]* Endrödi, 1951
* *[[Ataenius bispinulosus]]* Schmidt, 1911
* *[[Ataenius blapoides]]* Balthasar, 1947
* *[[Ataenius bolivarensis]]* Stebnicka, 2007
* *[[Ataenius bolivari]]* Stebnicka, 2001
* *[[Ataenius borgmeieri]]* Hinton, 1936
* *[[Ataenius borjae]]* Stebnicka, 2005
* *[[Ataenius brevicollis]]* (Wollaston, 1854)
* *[[Ataenius brevis]]* Fall, 1930
* *[[Ataenius brevitarsis]]* Petrovitz, 1964
* *[[Ataenius brouni]]* Sharp, 1876
* *[[Ataenius brunneus]]* Schmidt, 1922
* *[[Ataenius buenavistae]]* Stebnicka, 2001
* *[[Ataenius caesoides]]* Fairmaire, 1899
* *[[Ataenius caicarae]]* Stebnicka, 2005
* *[[Ataenius californicus]]* Horn, 1887
* *[[Ataenius canoasus]]* Stebnicka, 2007
* *[[Ataenius carinatipennis]]* Petrovitz, 1973
* *[[Ataenius carinator]]* Harold, 1874
* *[[Ataenius cartago]]* Stebnicka, 2007
* *[[Ataenius cartwrighti]]* Chalumeau & Gruner, 1974
* *[[Ataenius castaniellus]]* Bates, 1887
* *[[Ataenius catarinaensis]]* Stebnicka, 2007
* *[[Ataenius catenulatus]]* (Erichson, 1847)
* *[[Ataenius cavagnaroi]]* Cartwright, 1970
* *[[Ataenius ceylonensis]]* Schmidt, 1912
* *[[Ataenius chapini]]* Hinton, 1937
* *[[Ataenius chilensis]]* (Solier, 1851)
* *[[Ataenius chinacotae]]* Stebnicka, 2007
* *[[Ataenius circulusbrumalis]]* Petrovitz, 1964
* *[[Ataenius clavatus]]* Schmidt, 1916
* *[[Ataenius clitellarius]]* Petrovitz, 1973
* *[[Ataenius cochabambae]]* Stebnicka, 2005
* *[[Ataenius cognatus]]* (LeConte, 1858)
* *[[Ataenius columbicus]]* Harold, 1880
* *[[Ataenius communis]]* Hinton, 1936
* *[[Ataenius complicatus]]* Harold, 1869
* *[[Ataenius confertus]]* Fall, 1909
* *[[Ataenius convexus]]* Robinson, 1940
* *[[Ataenius coriarius]]* Schmidt, 1912
* *[[Ataenius corporaali]]* Boucomont, 1924
* *[[Ataenius corrosus]]* Chapin, 1940
* *[[Ataenius costulifer]]* Balthasar, 1951
* *[[Ataenius crenaticollis]]* Petrovitz, 1973
* *[[Ataenius crenatipennis]]* MacLeay, 1871
* *[[Ataenius crenator]]* Harold, 1876
* *[[Ataenius crenatostriatus]]* (Blanchard, 1851)
* *[[Ataenius crenulatus]]* Schmidt, 1910
* *[[Ataenius cribrithorax]]* Bates, 1887
* *[[Ataenius cristobalensis]]* Cook & Peck, 2000
* *[[Ataenius cucutae]]* Stebnicka, 2007
* *[[Ataenius cylindrus]]* Horn, 1871
* *[[Ataenius dentatus]]* Koshantshikov, 1916
* *[[Ataenius depilis]]* Petrovitz, 1976
* *[[Ataenius deserti]]* Blackburn, 1894
* *[[Ataenius desertus]]* Horn, 1871
* *[[Ataenius ecruensis]]* Stebnicka & Lago, 2005
* *[[Ataenius edungalbae]]* Stebnicka & Howden, 1997
* *[[Ataenius elegans]]* Harold, 1868
* *[[Ataenius elisaensis]]* Stebnicka, 2002
* *[[Ataenius elongatulus]]* MacLeay, 1888
* *[[Ataenius elongatus]]* (Palisot de Beauvois, 1811)
* *[[Ataenius endroedyyoungai]]* Endrödi, 1973
* *[[Ataenius erinaceus]]* Petrovitz, 1961
* *[[Ataenius eringundae]]* Stebnicka & Howden, 1997
* *[[Ataenius erratus]]* Fall, 1930
* *[[Ataenius eurynotus]]* Lea, 1923
* *[[Ataenius excisicollis]]* Petrovitz, 1961
* *[[Ataenius exiguus]]* Brown, 1932
* *[[Ataenius fattigi]]* Cartwright, 1948
* *[[Ataenius floreanae]]* Cook & Peck, 2000
* *[[Ataenius forgiei]]* Bordat & Théry, 2012
* *[[Ataenius forsteri]]* Balthasar, 1960
* *[[Ataenius freyi]]* Petrovitz, 1961
* *[[Ataenius gammonensis]]* Stebnicka & Howden, 1997
* *[[Ataenius garamas]]* Peyerimhoff, 1929
* *[[Ataenius gascoyneensis]]* Stebnicka & Howden, 1997
* *[[Ataenius gibbus]]* Blackburn, 1904
* *[[Ataenius gilesi]]* Stebnicka & Howden, 1997
* *[[Ataenius glabriventris]]* Schmidt, 1911
* *[[Ataenius glaseri]]* Cartwright, 1974
* *[[Ataenius gracilis]]* (Melsheimer, 1845)
* *[[Ataenius granchacoensis]]* Balthasar, 1939
* *[[Ataenius granocostatus]]* Schmidt, 1912
* *[[Ataenius griffini]]* Cartwright, 1974
* *[[Ataenius gruneri]]* Chalumeau, 1979
* *[[Ataenius guanacastae]]* Stebnicka, 2005
* *[[Ataenius guayasi]]* Stebnicka, 2001
* *[[Ataenius gungareei]]* Stebnicka & Howden, 1997
* *[[Ataenius guriensis]]* Stebnicka, 2005
* *[[Ataenius heinekeni]]* (Wollaston, 1854)
* *[[Ataenius hesperius]]* Cartwright, 1974
* *[[Ataenius hirsutus]]* Horn, 1871
* *[[Ataenius hispaniolae]]* Chalumeau, 1982
* *[[Ataenius hispidus]]* Harold, 1867
* *[[Ataenius holopubescens]]* Hinton, 1938
* *[[Ataenius horticola]]* Harold, 1869
* *[[Ataenius howdeni]]* Chalumeau, 1978
* *[[Ataenius hrubanti]]* Balthasar, 1967
* *[[Ataenius huanus]]* Stebnicka, 2007
* *[[Ataenius humptydooensis]]* Stebnicka & Howden, 1997
* *[[Ataenius hygrophilus]]* Paulian, 1947
* *[[Ataenius icanus]]* Balthasar, 1941
* *[[Ataenius illaetabilis]]* Lea, 1923
* *[[Ataenius imbricatoides]]* Schmidt, 1909
* *[[Ataenius imbricatus]]* (Melsheimer, 1845)
* *[[Ataenius imparilis]]* Blackburn, 1904
* *[[Ataenius impiger]]* Schmidt, 1916
* *[[Ataenius impressus]]* (Petrovitz, 1963)
* *[[Ataenius incanus]]* Balthasar, 1951
* *[[Ataenius indutus]]* Petrovitz, 1961
* *[[Ataenius inquisitus]]* Horn, 1887
* *[[Ataenius insculptus]]* Horn, 1887
* *[[Ataenius insolitus]]* Schmidt, 1909
* *[[Ataenius insulae]]* Chalumeau & Gruner, 1974
* *[[Ataenius insularis]]* Lea, 1923
* *[[Ataenius integer]]* Harold, 1868
* *[[Ataenius integricollis]]* Lea, 1923
* *[[Ataenius intermedius]]* Bates, 1887
* *[[Ataenius iquitosae]]* Stebnicka, 2007
* *[[Ataenius isabelae]]* Franz, 1985
* *[[Ataenius jamaicensis]]* Chapin, 1940
* *[[Ataenius jardinensis]]* Stebnicka, 2002
* *[[Ataenius jelineki]]* Chalumeau, 1982
* *[[Ataenius kapalgaensis]]* Stebnicka & Howden, 1997
* *[[Ataenius klapperichi]]* Howden, 1978
* *[[Ataenius kochi]]* Balthasar, 1941
* *[[Ataenius koebelei]]* Blackburn, 1904
* *[[Ataenius koelleri]]* Balthasar, 1963
* *[[Ataenius koghianus]]* Paulian, 1991
* *[[Ataenius lamarensis]]* Stebnicka, 2007
* *[[Ataenius lamotteiroyi]]* Bordat, 1992
* *[[Ataenius lanei]]* Petrovitz, 1973
* *[[Ataenius languidus]]* Schmidt, 1911
* *[[Ataenius latus]]* Petrovitz, 1963
* *[[Ataenius lenkoi]]* Petrovitz, 1973
* *[[Ataenius liogaster]]* Bates, 1887
* *[[Ataenius lobatus]]* Horn, 1871
* *[[Ataenius londrinae]]* Stebnicka, 2007
* *[[Ataenius longiclavus]]* Petrovitz, 1970
* *[[Ataenius lucanus]]* Horn, 1871
* *[[Ataenius luctuosus]]* (Burmeister, 1877)
* *[[Ataenius luteomargo]]* Chapin, 1940
* *[[Ataenius madagassicus]]* Bordat, 1990
* *[[Ataenius maghribinicus]]* Baraud, 1985
* *[[Ataenius martinezi]]* Petrovitz, 1973
* *[[Ataenius miamii]]* Cartwright, 1934
* *[[Ataenius michelii]]* Chalumeau, 1978
* *[[Ataenius microtrichopterus]]* Lea, 1923
* *[[Ataenius millstreamae]]* Stebnicka & Howden, 1997
* *[[Ataenius montanus]]* Schmidt, 1911
* *[[Ataenius monteithi]]* Paulian, 1991
* *[[Ataenius monticola]]* Paulian, 1940
* *[[Ataenius montreuili]]* Bordat & Théry, 2012
* *[[Ataenius morator]]* Harold, 1869
* *[[Ataenius murchisoni]]* Stebnicka & Howden, 1997
* *[[Ataenius nakpandurii]]* Endrödi, 1973
* *[[Ataenius nanus]]* (DeGeer, 1774)
* *[[Ataenius napoensis]]* Stebnicka, 2007
* *[[Ataenius nigricans]]* Paulian, 1933
* *[[Ataenius noques]]* Stebnicka, 2007
* *[[Ataenius noronhai]]* Stebnicka, 2007
* *[[Ataenius nudus]]* Blackburn, 1904
* *[[Ataenius nugator]]* Harold, 1880
* *[[Ataenius oaxacaensis]]* Stebnicka & Lago, 2005
* *[[Ataenius ocumarensis]]* Stebnicka, 2001
* *[[Ataenius oklahomensis]]* Brown, 1930
* *[[Ataenius onkonensis]]* Stebnicka, 2005
* *[[Ataenius opacipennis]]* Schmidt, 1910
* *[[Ataenius opatrinus]]* Harold, 1867
* *[[Ataenius opatroides]]* (Balthasar, 1947)
* *[[Ataenius ovatulus]]* Horn, 1871
* *[[Ataenius pacificus]]* (Sharp, 1879)
* *[[Ataenius palmaritoensis]]* Stebnicka, 2007
* *[[Ataenius palmerstoni]]* Blackburn, 1891
* *[[Ataenius palustris]]* (Montrouzier, 1860)
* *[[Ataenius panamensis]]* Hinton, 1936
* *[[Ataenius parallelipennis]]* Petrovitz, 1973
* *[[Ataenius parvus]]* Lea, 1923
* *[[Ataenius pearlensis]]* Stebnicka, 2006
* *[[Ataenius peregianensis]]* Stebnicka & Howden, 1997
* *[[Ataenius peregrinator]]* Harold, 1877
* *[[Ataenius pereirai]]* Petrovitz, 1970
* *[[Ataenius perforatus]]* Harold, 1867
* *[[Ataenius pertuga]]* Balthasar, 1961
* *[[Ataenius petrovitzi]]* Balthasar, 1960
* *[[Ataenius picinus]]* Harold, 1867
* *[[Ataenius platensis]]* (Blanchard, 1847)
* *[[Ataenius plaumanni]]* Petrovitz, 1973
* *[[Ataenius polyglyptus]]* Bates, 1887
* *[[Ataenius pseudimparilis]]* Stebnicka & Howden, 1997
* *[[Ataenius pseudocarinator]]* Balthasar, 1947
* *[[Ataenius pseudoclavatus]]* Stebnicka, 2005
* *[[Ataenius pseudocommunis]]* Stebnicka, 2001
* *[[Ataenius pseudostercorator]]* Stebnicka, 2003
* *[[Ataenius pseudousingeri]]* Galante, Stebnicka & Verdú, 2003
* *[[Ataenius punctatohirsutus]]* Schmidt, 1909
* *[[Ataenius puncticollis]]* LeConte, 1868
* *[[Ataenius punctipennis]]* Harold, 1868
* *[[Ataenius purator]]* Harold, 1868
* *[[Ataenius quintanaroo]]* Stebnicka, 2006
* *[[Ataenius raccurti]]* Chalumeau, 1978
* *[[Ataenius rakovici]]* Bordat & Théry, 2012
* *[[Ataenius raslani]]* Bordat & Théry, 2012
* *[[Ataenius raucus]]* Schmidt, 1908
* *[[Ataenius robustus]]* Horn, 1871
* *[[Ataenius rosinae]]* Balthasar, 1967
* *[[Ataenius rubrotessellatus]]* (Blanchard, 1843)
* *[[Ataenius sabinoi]]* Cartwright, 1974
* *[[Ataenius saltae]]* Stebnicka, 2007
* *[[Ataenius santarosae]]* Stebnicka, 2007
* *[[Ataenius saramari]]* Cartwright, 1939
* *[[Ataenius saulensis]]* Stebnicka, 2006
* *[[Ataenius scabrelloides]]* Petrovitz, 1962
* *[[Ataenius scabrellus]]* Schmidt, 1909
* *[[Ataenius scalptifrons]]* Bates, 1887
* *[[Ataenius schmidti]]* Stebnicka, 2003
* *[[Ataenius sculptilis]]* Harold, 1868
* *[[Ataenius sculptor]]* Harold, 1868
* *[[Ataenius scutellaris]]* Harold, 1867
* *[[Ataenius seaforthensis]]* Stebnicka & Howden, 1997
* *[[Ataenius semicoecus]]* MacLeay, 1888
* *[[Ataenius semicornutus]]* MacLeay, 1871
* *[[Ataenius semipilosus]]* Van Dyke, 1928
* *[[Ataenius setiger]]* Bates, 1887
* *[[Ataenius setosus]]* Schmidt, 1909
* *[[Ataenius seydeli]]* Endrödi, 1964
* *[[Ataenius siminasus]]* Petrovitz, 1973
* *[[Ataenius simplicipes]]* Mulsant & Rey, 1870
* *[[Ataenius skelleyi]]* Stebnicka, 2007
* *[[Ataenius sparsicollis]]* Blackburn, 1904
* *[[Ataenius speculator]]* Blackburn, 1891
* *[[Ataenius spinipennis]]* Lea, 1923
* *[[Ataenius spretulus]]* (Haldeman, 1848)
* *[[Ataenius stebnickae]]* Bordat & Théry, 2012
* *[[Ataenius steinheili]]* Harold, 1874
* *[[Ataenius stephani]]* Cartwright, 1974
* *[[Ataenius stercorator]]* (Fabricius, 1775)
* *[[Ataenius striatocrenatus]]* Fairmaire, 1889
* *[[Ataenius strigatus]]* (Say, 1823)
* *[[Ataenius strigicaudus]]* Bates, 1887
* *[[Ataenius strigifrons]]* Schmidt, 1920
* *[[Ataenius suineata]]* Deloya, 2012
* *[[Ataenius synnotensis]]* Stebnicka & Howden, 1997
* *[[Ataenius talpoides]]* Stebnicka, 2001
* *[[Ataenius tambopatae]]* Stebnicka, 2001
* *[[Ataenius tarumensis]]* Stebnicka, 2007
* *[[Ataenius temperei]]* Chalumeau & Gruner, 1974
* *[[Ataenius terminalis]]* (Chevrolat, 1864)
* *[[Ataenius texanus]]* Harold, 1874
* *[[Ataenius tindalensis]]* Stebnicka & Howden, 1997
* *[[Ataenius tomentosus]]* Petrovitz, 1969
* *[[Ataenius torridus]]* Blackburn, 1892
* *[[Ataenius tovarensis]]* Stebnicka, 2001
* *[[Ataenius trox]]* Minkina & Skelley, 2025
* *[[Ataenius tuberculatus]]* Schmidt, 1911
* *[[Ataenius uriarrae]]* Stebnicka & Howden, 1997
* *[[Ataenius usingeri]]* Hinton, 1937
* *[[Ataenius utahensis]]* Cartwright, 1974
* *[[Ataenius vandykei]]* Cartwright, 1974
* *[[Ataenius variopunctatus]]* Schmidt, 1922
* *[[Ataenius versicolor]]* Schmidt, 1916
* *[[Ataenius vethianus]]* Schmidt, 1909
* *[[Ataenius vinacoensis]]* Stebnicka, 2006
* *[[Ataenius walkeri]]* Blackburn, 1904
* *[[Ataenius waltherhorni]]* Balthasar, 1938
* *[[Ataenius warisensis]]* Stebnicka, 1998
* *[[Ataenius wenzelii]]* Horn, 1887
* *[[Ataenius windjanae]]* Stebnicka & Howden, 1997
* *[[Ataenius yungasus]]* Stebnicka, 2001

## Referanser


## Kilder
* Generic Guide to New World Scarab Beetles - *Ataenius* http://museum.unl.edu/research/entomology/Guide/Scarabaeoidea/Scarabaeidae/Aphodiinae/AphodiinaeTribes/Eupariini/Ataenius/Ataenius.html
* Z. Stebnicka og P.K. Lago (1995) The New World species of *Ataenius* Harold, 1867. V. Revision of the *A. strigatus* group (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi 19: 55-83 http://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1091&context=insectamundi&sei-redir=1&referer=http%3A%2F%2Fwww.bing.com%2Fsearch%3Fq%3DAtaenius%26qs%3Dn%26sp%3D-1%26pq%3Dataenius%26sc%3D1-8%26sk%3D%26cvid%3D33CBB99F631F4CB39CE5F0476135B165%26first%3D11%26FORM%3DPERE#search=%22Ataenius%22
* biolib.cz - Oversikt over stammer og slekter

## Eksterne lenker

*