Engviklere

**Engviklere** (***Eucosma*** spp.) er en artsrik slekt i sommerfuglfamilien viklere (Tortricidae) som hører til undergruppen Olethreutinae. Det er beskrevet ca. 280 arter på verdensbasis, men det oppdages stadig nye, 13 av dem er funnet i Norge.

## Utseende
Små til nokså store (de norske artene har vingespenn 10-22 millimeter) viklere med forholdsvis smale vinger. Mange arter er nokså ensfarget brunlige, men det er også noen som er kontrastrikt tegnede.

## Levevis
Disse viklerne lever på planter i kurvplantefamilien.
De fleste av artene lever på blomsterhodene, men det er også noen som borer inne i stengel eller røtter. Disse artene er vanligvis spesialiserte på én enkelt planteart (monofage), på grunn av dette er ganske mange av artene sjeldne og truede.

## Utbredelse
Slekten er utbredt over hele Palearktis og i Nord-Amerika.

## Systematisk inndeling

* sommerfugler (Lepidoptera)
** gruppe **[[Heteroneura]]** 
*** gruppe **[[Apoditrysia]]**
**** gruppe / overfamilie **[[Tortricoidea|viklere]]** (Tortricoidea) 
***** familie **[[viklere]]** (Tortricidae) 
****** Underfamilien Olethreutinae
******* gruppe Eucosmini
******** slekten engviklere, *[[Eucosma]]* Hübner, 1823 – 13 arter i Norge
********* dylleengvikler, *[[Eucosma obumbratana]]* (Lienig & Zeller, 1846)
********* grisøreengvikler, *[[Eucosma scorzonerana]]* (Benander, 1942)
********* tistelengvikler, *[[Eucosma cana]]* (Haworth, 1811)
********* mørk knoppurtengvikler, *[[Eucosma hohenwartiana]]* (Denis & Schiffermüller, 1775)
********* landøydaengvikler, *[[Eucosma campoliliana]]* (Denis & Schiffermüller, 1775)
********* lys gullrisengvikler, *Eucosma aemulana* (Schläger, 1849)
********* strandstjerneengvikler, *[[Eucosma tripoliana]]* (Barrett, 1880)
********* hvit engvikler, *[[Eucosma lacteana]]* (Treitschke, 1835)
********* lys knoppurtengvikler, *[[Eucosma fulvana]]* (Stephens, 1834)
********* strandengvikler, *[[Eucosma guentheri]]* (Tengström, 1869)
********* skogsalatengvikler, *[[Eucosma conterminana]]* (Guenée, 1845)
********* mørk gullrisengvikler, *Eucosma aspidiscana* (Hübner, 1817)
********* malurtengvikler, *[[Eucosma pupillana]]* (Clerck, 1759)
********* fjellengvikler, *[[Eucosma saussureana]]* (Benander, 1928) 
********* oker gullrisengvikler, *[[Eucosma suomiana]]* (A. Hoffmann, 1893)
******** Arter som er funnet i Europa, men ikke i Norge:
********* *[[Eucosma abacana]]* (Erschoff, 1877)
********* *[[Eucosma agnatana]]* (Christoph, 1872)
********* *[[Eucosma albarracina]]* Hartig, 1941
********* *[[Eucosma albicosta]]* Falkovitsh, 1964
********* *[[Eucosma albuneana]]* (Zeller, 1847)
********* *[[Eucosma apocrypha]]* Falkovitsh, 1964
********* *[[Eucosma apocryphoides]]* Budashkin, 2009
********* *[[Eucosma balatonana]]* (Osthelder, 1937)
********* *[[Eucosma caliacrana]]* (Caradja, 1931)
********* *[[Eucosma clarescens]]* Kuznetzov, 1964
********* *[[Eucosma conformana]]* (Mann, 1872)
********* *[[Eucosma cretaceana]]* (Kennel, 1899)
********* *[[Eucosma culmana]]* (Muller-Rutz, 1932)
********* *[[Eucosma cumulana]]* (Guenee, 1845)
********* *[[Eucosma diakonoffi]]* Gibeaux, 1984
********* *[[Eucosma disjectana]]* (Kennel, 1921)
********* *[[Eucosma explicatana]]* (Kennel, 1900)
********* *[[Eucosma fervidana]]* (Zeller, 1847)
********* *[[Eucosma flavispecula]]* Kuznetzov, 1964
********* *[[Eucosma fuscana]]* Kearfott, 1907
********* *[[Eucosma gonzalezalvarezi]]* Agenjo, 1970
********* *[[Eucosma gradensis]]* (Galvagni, 1909)
********* *[[Eucosma halophilana]]* Budashkin, 2009
********* *[[Eucosma incinerana]]* (Constant, 1888)
********* *[[Eucosma individiosana]]* (Kennel, 1901)
********* *[[Eucosma krygeri]]* (Rebel, 1937)
********* *[[Eucosma luciana]]* (Chrtien, 1908)
********* *[[Eucosma lugubrana]]* (Treitschke, 1830)
********* *[[Eucosma messingiana]]* (Fischer von Röslerstamm, 1837)
********* *[[Eucosma metzneriana]]* (Treitschke, 1830)
********* *[[Eucosma mirificana]]* (Peyerimhoff, 1877)
********* *[[Eucosma monstratana]]* (Rebel, 1906)
********* *[[Eucosma paetulana]]* (Kennel, 1900)
********* *[[Eucosma parvulana]]* (Wilkinson, 1859)
********* *[[Eucosma rubescana]]* (Constant, 1895
********* *[[Eucosma sardoensis]]* (Rebel, 1935)
********* *[[Eucosma sublucidana]]* (Kennel, 1901)
********* *[[Eucosma tetraplana]]* (Moschler, 1866)
********* *[[Eucosma tundrana]]* (Kennel, 1900)
********* *[[Eucosma ukrainica]]* Budashkin, 2009
********* *[[Eucosma urbana]]* (Kennel, 1901)
********* *[[Eucosma wimmerana]]* (Treitschke, 1835)
******** Arter som ikke er funnet i Europa:
********* *[[Eucosma abstemia]]* Meyrick, 1932
********* *[[Eucosma adamantana]]* (Guenee, 1845)
********* *[[Eucosma aeana]]* McDunnough, 1942
********* *[[Eucosma agassizi]]* (Robinson, 1869)
********* *[[Eucosma agricolana]]* (Walsingham, 1879)
********* *[[Eucosma albicostella]]* (Turati & Krger, 1936)
********* *[[Eucosma albidulana]]* (Herrich-Schffer, 1851)
********* *[[Eucosma albiguttana]]* (Zeller, 1875)
********* *[[Eucosma alphabetica]]* Walsingham, 1914
********* *[[Eucosma amellana]]* (Preissecker, 1930)
********* *[[Eucosma aprilana]]* (Grote, 1877)
********* *[[Eucosma argentifera]]* Razowski, 1972
********* *[[Eucosma aspidana]]* (Walsingham, 1884)
********* *[[Eucosma atascosana]]* Blanchard, 1980
********* *[[Eucosma atomosana]]* (Walsingham, 1879)
********* *[[Eucosma aurantiradix]]* Kuznetzov, 1962
********* *[[Eucosma aurilineana]]* Ferris, 2005
********* *[[Eucosma australis]]* Kuznetzov, 1988
********* *[[Eucosma austrina]]* Miller, 1985
********* *[[Eucosma avalona]]* McDunnough, 1938
********* *[[Eucosma bactromorpha]]* Diakonoff, 1992
********* *[[Eucosma barbara]]* Miller, 1974
********* *[[Eucosma betana]]* McDunnough, 1942
********* *[[Eucosma bilineana]]* Kearfott, 1907
********* *[[Eucosma biplagata]]* (Walsingham, 1895)
********* *[[Eucosma bipunctella]]* (Walker, 1863)
********* *[[Eucosma biquadrana]]* (Walsingham, 1879)
********* *[[Eucosma bobana]]* Kearfott, 1907
********* *[[Eucosma bolanderana]]* (Walsingham, 1879)
********* *[[Eucosma brachysticta]]* Meyrick, in Caradja & Meyrick, 1935
********* *[[Eucosma brigittana]]* (Kennel, 1919)
********* *[[Eucosma canana]]* (Walsingham, 1879)
********* *[[Eucosma canariana]]* Kearfott, 1907
********* *[[Eucosma caniceps]]* (Walsingham, 1884)
********* *[[Eucosma capnoleuca]]* Meyrick, 1932
********* *[[Eucosma cataclystiana]]* (Walker, 1863)
********* *[[Eucosma catharaspis]]* (Meyrick, 1922)
********* *[[Eucosma catoptrana]]* (Rebel, 1903)
********* *[[Eucosma certana]]* Kuznetzov, 1967
********* *[[Eucosma chlorobathra]]* Meyrick, 1911
********* *[[Eucosma chrysyphis]]* Razowski, 1972
********* *[[Eucosma circulana]]* (Hubner, 1823)
********* *[[Eucosma coagulana]]* (Kennel, 1901)
********* *[[Eucosma cocana]]* Kearfott, 1907
********* *[[Eucosma comatulana]]* (Zeller, 1876)
********* *[[Eucosma confunda]]* Kuznetzov, 1966
********* *[[Eucosma coniogramma]]* Clarke, 1976
********* *[[Eucosma consobrinana]]* Heinrich, 1923
********* *[[Eucosma consociana]]* Heinrich, 1923
********* *[[Eucosma conspiciendana]]* Heinrich, 1923
********* *[[Eucosma corosana]]* (Walsingham, 1884)
********* *[[Eucosma costastrigulana]]* Kearfott, 1907
********* *[[Eucosma crambitana]]* (Walsingham, 1879)
********* *[[Eucosma crymalana]]* Powell, 1968
********* *[[Eucosma dapsilis]]* Heinrich, 1929
********* *[[Eucosma denigratana]]* (Kennel, 1901)
********* *[[Eucosma denverana]]* Kearfott, 1907
********* *[[Eucosma derelicta]]* Heinrich, 1929
********* *[[Eucosma diabolana]]* Blanchard, 1980
********* *[[Eucosma dilatana]]* (Walsingham, 1895)
********* *[[Eucosma directa]]* Meyrick, 1912
********* *[[Eucosma discernata]]* Kuznetzov, 1966
********* *[[Eucosma dodana]]* Kearfott, 1907
********* *[[Eucosma doiinthanonensis]]* Kawabe, 1989
********* *[[Eucosma dorsisignatana]]* (Clemens, 1860)
********* *[[Eucosma eburata]]* Heinrich, 1929
********* *[[Eucosma ephedrana]]* (Christoph, 1877)
********* *[[Eucosma esmodes]]* Meyrick, 1937
********* *[[Eucosma excerptionana]]* Heinrich, 1923
********* *[[Eucosma exclusoriana]]* Heinrich, 1923
********* *[[Eucosma excusabilis]]* Heinrich, 1923
********* *[[Eucosma fandana]]* Kearfott, 1907
********* *[[Eucosma fernaldana]]* (Grote, 1880)
********* *[[Eucosma fiskeana]]* Kearfott, 1905
********* *[[Eucosma floridana]]* Kearfott, 1907
********* *[[Eucosma fofana]]* Kearfott, 1907
********* *[[Eucosma franclemonti]]* Powell, 1968
********* *[[Eucosma fraudabilis]]* Heinrich, 1923
********* *[[Eucosma fraudulentana]]* (Kennel, 1901)
********* *[[Eucosma fritillana]]* Blanchard & Knudson, 1982
********* *[[Eucosma fulminana]]* (Walsingham, 1879)
********* *[[Eucosma fuscida]]* Kuznetzov, 1966
********* *[[Eucosma galenapunctana]]* Kearfott, 1905
********* *[[Eucosma gandana]]* Kearfott, 1907
********* *[[Eucosma getonia]]* Razowski, 1972
********* *[[Eucosma giarabubensis]]* Turati, 1930
********* *[[Eucosma giganteana]]* (Riley, 1881)
********* *[[Eucosma gilletteana]]* Dyar, 1903
********* *[[Eucosma glebana]]* (Snellen, 1883)
********* *[[Eucosma glomerana]]* (Walsingham, 1879)
********* *[[Eucosma gloriola]]* Heinrich, 1931
********* *[[Eucosma gomonana]]* Kearfott, 1907
********* *[[Eucosma gorodkovi]]* Kuznetzov, 1979
********* *[[Eucosma graciliana]]* Kearfott, 1905
********* *[[Eucosma graduatana]]* (Walsingham, 1879)
********* *[[Eucosma grandiflavana]]* (Walsingham, 1879)
********* *[[Eucosma graziella]]* Blanchard, 1968
********* *[[Eucosma griselda]]* Blanchard & Knudson, 1982
********* *[[Eucosma guttulana]]* Blanchard, 1980
********* *[[Eucosma gypsatana]]* (Kennel, 1921)
********* *[[Eucosma haberhaueri]]* (Kennel, 1901)
********* *[[Eucosma handana]]* Kearfott, 1907
********* *[[Eucosma hasseanthi]]* Clarke, 1952
********* *[[Eucosma haydenae]]* Wright, 2006
********* *[[Eucosma hazelana]]* Klots, 1936
********* *[[Eucosma heathiana]]* Kearfott, 1907
********* *[[Eucosma hebescana]]* Kuznetzov, 1986
********* *[[Eucosma heinrichi]]* McDunnough, 1925
********* *[[Eucosma hennei]]* Clarke, 1947
********* *[[Eucosma hohana]]* Kearfott, 1907
********* *[[Eucosma hyponomeutana]]* (Walsingham, 1895)
********* *[[Eucosma ignotana]]* (Caradja, 1916)
********* *[[Eucosma immaculana]]* Kearfott, 1907
********* *[[Eucosma inquadrana]]* (Walsingham, 1884)
********* *[[Eucosma intermediana]]* (Kennel, 1900)
********* *[[Eucosma invicta]]* (Walsingham, 1895)
********* *[[Eucosma iographa]]* Diakonoff, 1968
********* *[[Eucosma irroratana]]* (Walsingham, 1879)
********* *[[Eucosma jansei]]* Diakonoff, 1956
********* *[[Eucosma jejunana]]* McDunnough, 1942
********* *[[Eucosma jerusalemana]]* (Amsel, 1935)
********* *[[Eucosma juncticiliana]]* (Walsingham, 1879)
********* *[[Eucosma kurdistana]]* (Amsel, 1959)
********* *[[Eucosma landana]]* Kearfott, 1907
********* *[[Eucosma langstoni]]* Powell, 1963
********* *[[Eucosma larana]]* (Walsingham, 1879)
********* *[[Eucosma lathami]]* Forbes, 1937
********* *[[Eucosma laticurva]]* Heinrich, 1929
********* *[[Eucosma lignana]]* (Snellen, 1883)
********* *[[Eucosma lioplintha]]* Meyrick, 1920
********* *[[Eucosma liturana]]* (Walsingham, 1879)
********* *[[Eucosma lolana]]* Kearfott, 1907
********* *[[Eucosma louisana]]* McDunnough, 1944
********* *[[Eucosma luridana]]* (Walsingham, 1879)
********* *[[Eucosma lyrana]]* (Snellen, 1883)
********* *[[Eucosma maculatana]]* (Walsingham, 1879)
********* *[[Eucosma magnana]]* Kuznetzov, 1978
********* *[[Eucosma magnidicana]]* Heinrich, 1923
********* *[[Eucosma matutina]]* (Grote, 1873)
********* *[[Eucosma mediostriata]]* (Walsingham, 1895)
********* *[[Eucosma metagrapta]]* Meyrick, 1932
********* *[[Eucosma metana]]* (Kennel, 1919)
********* *[[Eucosma micropterana]]* (Turati, 1930)
********* *[[Eucosma microsignata]]* Heinrich, 1929
********* *[[Eucosma mobilensis]]* Heinrich, 1923
********* *[[Eucosma momana]]* Kearfott, 1907
********* *[[Eucosma monitorana]]* Heinrich, 1920
********* *[[Eucosma monoensis]]* Powell, 1968
********* *[[Eucosma monogrammana]]* (Zeller, 1875)
********* *[[Eucosma morrisoni]]* (Walsingham, 1884)
********* *[[Eucosma muguraxana]]* Kostyuk, 1975
********* *[[Eucosma muliebris]]* Meyrick, 1922
********* *[[Eucosma nandana]]* Kearfott, 1907
********* *[[Eucosma nitorana]]* Kuznetzov, 1962
********* *[[Eucosma niveicaput]]* (Walsingham, 1900)
********* *[[Eucosma nordini]]* Wright, 2005
********* *[[Eucosma notanthes]]* Meyrick, 1936
********* *[[Eucosma notialis]]* Miller, 1985
********* *[[Eucosma nuntia]]* Heinrich, 1929
********* *[[Eucosma ochricostana]]* Razowski, 1972
********* *[[Eucosma oculatana]]* (Kennel, 1900)
********* *[[Eucosma ommatoptera]]* Falkovitsh, 1965
********* *[[Eucosma ophionana]]* McDunnough, 1925
********* *[[Eucosma optimana]]* Dyar, 1903
********* *[[Eucosma ottoniana]]* (Kennel, 1919)
********* *[[Eucosma palabundana]]* Heinrich, 1923
********* *[[Eucosma pamirana]]* Kuznetzov, 1972
********* *[[Eucosma pandana]]* Kearfott, 1907
********* *[[Eucosma pediasios]]* Miller, 1985
********* *[[Eucosma pedisignata]]* Diakonoff, 1948
********* *[[Eucosma perdricana]]* (Walsingham, 1879)
********* *[[Eucosma pergratana]]* (Rebel, 1914)
********* *[[Eucosma persiae]]* Razowski, 1963
********* *[[Eucosma persolita]]* Heinrich, 1929
********* *[[Eucosma phaecochyta]]* Bradley, 1965
********* *[[Eucosma piperata]]* Wright, 2005
********* *[[Eucosma plumbaginea]]* Meyrick, 1932
********* *[[Eucosma ponderosa]]* Powell, 1968
********* *[[Eucosma primulana]]* (Walsingham, 1879)
********* *[[Eucosma pulveratana]]* (Walsingham, 1879)
********* *[[Eucosma quinquemaculana]]* (Robinson, 1869)
********* *[[Eucosma ragonoti]]* (Walsingham, 1895)
********* *[[Eucosma recentana]]* (Zerny, 1933)
********* *[[Eucosma rescissoriana]]* Heinrich, 1920
********* *[[Eucosma rhymogramma]]* Meyrick, 1916
********* *[[Eucosma ridingsana]]* (Robinson, 1869)
********* *[[Eucosma rigidana]]* (Snellen, 1883)
********* *[[Eucosma rindgei]]* Miller, 1985
********* *[[Eucosma robinsonana]]* (Grote, 1872)
********* *[[Eucosma rosaocellana]]* Knudson, 1986
********* *[[Eucosma russeola]]* Heinrich, 1929
********* *[[Eucosma rusticana]]* Kearfott, 1905
********* *[[Eucosma salaciana]]* Blanchard & Knudson, 1982
********* *[[Eucosma sandiego]]* Kearfott, 1908
********* *[[Eucosma scutiformis]]* Meyrick, in Caradja, 1931
********* *[[Eucosma serapicana]]* Heinrich, 1923
********* *[[Eucosma serpentana]]* (Walsingham, 1895)
********* *[[Eucosma shastana]]* (Walsingham, 1879)
********* *[[Eucosma sierrae]]* Blanchard & Knudson, 1983
********* *[[Eucosma similiana]]* (Clemens, 1860)
********* *[[Eucosma simplex]]* McDunnough, 1925
********* *[[Eucosma siskiyouana]]* (Kearfott, 1907)
********* *[[Eucosma smithiana]]* (Walsingham, 1895)
********* *[[Eucosma snyderana]]* Kearfott, 1907
********* *[[Eucosma sombreana]]* Kearfott, 1905
********* *[[Eucosma sonomana]]* Kearfott, 1907
********* *[[Eucosma spaldingana]]* Kearfott, 1907
********* *[[Eucosma sparsana]]* (Rebel, 1935)
********* *[[Eucosma sperryana]]* McDunnough, 1942
********* *[[Eucosma sphalerodes]]* Meyrick, 1934
********* *[[Eucosma striatiradix]]* Kuznetzov, 1964
********* *[[Eucosma striatulana]]* (Walsingham, 1900)
********* *[[Eucosma suadana]]* Heinrich, 1923
********* *[[Eucosma subflavana]]* (Walsingham, 1879)
********* *[[Eucosma subinvicta]]* Kearfott, 1907
********* *[[Eucosma sybillana]]* (Kennel, 1919)
********* *[[Eucosma taosana]]* Wright, 2005
********* *[[Eucosma tocullionana]]* Heinrich, 1920
********* *[[Eucosma tonitrualis]]* Meyrick, 1934
********* *[[Eucosma totana]]* Kearfott, 1907
********* *[[Eucosma urnigera]]* Meyrick, 1937
********* *[[Eucosma ursulana]]* (Kennel, 1919)
********* *[[Eucosma vagana]]* McDunnough, 1925
********* *[[Eucosma victoriana]]* (Kennel, 1919)
********* *[[Eucosma vulpecularis]]* Meyrick, 1932
********* *[[Eucosma wandana]]* Kearfott, 1907
********* *[[Eucosma watertonana]]* McDunnough, 1925
********* *[[Eucosma williamsi]]* Powell, 1963
********* *[[Eucosma yasudai]]* Nasu, 1982

## Kilder
* Fauna Europaea Web Service (2004) Fauna Europaea version 1.1, www.faunaeur.org 
* Aarvik, Leif, Berggren, Kai, Hansen, Lars Ove. 2000. *Norges Sommerfugler. Catalogus Lepidopterorum  Norvegiae.* Norsk entomologisk forening. 192 sider.

## Eksterne lenker
* 
*  www.nhm.uio.no/fakta/zoologi/insekter/norlep/  Norges sommerfugler – Et norskspråklig nettsted om norske sommerfugler.