Coleus

***Coleus*** ialah genus bagi herba atau pokok renek tahunan atau saka, yang kadang-kadang bersukulen, kadang-kadang dicirikan dengan pokok penanti yang berisi atau berubi, dan ditemui di kawasan tropika dan subtropika Afro-Eurasia.

## Taksonomi
Genus *Coleus* julung kali didefinisikan oleh seorang ahli Botani Portugal, João de Loureiro pada tahun 1790. Dengan menggunakan aksara morfologi sebagai bantuan, *Coleus* dibezakan dengan *Plectranthus* (pertama kali diterangkan oleh Charles L'Héritier pada tahun 1788) dengan meletakkan empat stamennya bersama-sama dan bukannya dibiarkan bebas ke pangkal. Pada tahun 1962, J.K. Morton menyatakan bahawastamen yang bercamtum lebih meluas daripada apa yang difikirkan sebelum ini, dan lantaran itu beliau menggabungkan *Coleus* ke dalam *Plectranthus*, dengan mengekalkan *Solenostemon* dan beberapa genera lain sebagai genus yang berbeza. Kebanyakan sumber mengikuti Morton dalam gabungan *Coleus*; sesetengahnya pula mengekalkan *Solenostemon* sebagai satu takrif yang berbeza, walaupun yang lain membenamkan ke dalam *Plectranthus* bersama-sama dengan *Coleus*.

Kajian awal suku Ocimeae pada tahun 2004 telah menunjukkan bahawa subsuku Plectranthinae adalah monofiletik, dengan dua klad utama: klad pertama mengandungi spesies jenis *Coleus* dan termasuk *Solenostemon*, manakala klad kedua pula mengandungi spesies jenis *Plectranthus* bersama beberapa genera lain, agar *Plectranthus* dapat ditakrifkan secara meluas sebagai bukan monofiletik. Kajian yang lebih terperinci pada tahun 2018 juga memperoleh kesimpulan yang serupa, dan mencadangkan bahawa *Coleus* (termasuk *Solenostemon*) harus diiktiraf semula, skop takrifan bagi *Plectranthus* perlulah lebih kecil, dan genus baharu, *[[Equilabium]]* diperkenalkan untuk klad spesies *Plectranthus* dahulu yang dari Afrika tropika. Banyak kombinasi binomial baharu yang diperlukan untuk melaksanakan pendekatan ini telah disediakan pada 2019.

## Senarai spesies
Setakat April 2024, *[[Plants of the World Online]]* telah menyenaraikan sebanyak 302 spesies dalam genus *Colues*.

**[[Coleus abyssinicus]]* (Fresen.) A.J.Paton
**[[Coleus acariformis]]* (P.I.Forst.) P.I.Forst.
**[[Coleus achankoviliensis]]* Smitha & A.J.Paton
**[[Coleus actites]]* (P.I.Forst.) P.I.Forst.
**[[Coleus adenanthus]]* (Dalzell & A.Gibson) A.J.Paton
**[[Coleus adenophorus]]* (Gürke) A.J.Paton
**[[Coleus aegyptiacus]]* (Forssk.) A.J.Paton
**[[Coleus affinis]]* (Gürke) A.J.Paton
**[[Coleus albicalyx]]* (Suddee) Suddee
**[[Coleus aliciae]]* (Codd) A.J.Paton
**[[Coleus alloplectus]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus alpinus]]* Vatke
**[[Coleus altanmouiensis]]* (T.C.Wilson, P.I.Forst. & M.A.M.Renner) T.C.Wilson & P.I.Forst.
**[[Coleus amboinicus]]* Lour.
**[[Coleus amicorum]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus amiculatus]]* (T.C.Wilson, P.I.Forst. & M.A.M.Renner) T.C.Wilson & P.I.Forst.
**[[Coleus amoenus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus anamudianus]]* (Smitha & Sunojk.) Smitha
**[[Coleus angolensis]]* (G.Taylor) A.J.Paton
**[[Coleus angulatus]]* (Hedge) A.J.Paton & Phillipson
**[[Coleus anthonyi]]* Jebin Joseph & J.Mathew
**[[Coleus apoensis]]* Elmer
**[[Coleus apreptus]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus apricus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus arabicus]]* Benth.
**[[Coleus arenicola]]* (P.I.Forst.) P.I.Forst.
**[[Coleus argentatus]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus argenteus]]* (Gamble) A.J.Paton
**[[Coleus argentifolius]]* (Ryding) A.J.Paton
**[[Coleus articulatus]]* (I.M.Johnst.) A.J.Paton
**[[Coleus ater]]* A.J.Paton
**[[Coleus auriglandulosus]]* (A.J.Paton) A.J.Paton
**[[Coleus australis]]* (R.Br.) A.J.Paton
**[[Coleus autranii]]* Briq.
**[[Coleus barbatus]]* (Andrews) Benth. ex G.Don
**[[Coleus bariensis]]* (Ryding) A.J.Paton
**[[Coleus batesii]]* (Baker) A.J.Paton
**[[Coleus batianoffii]]* (P.I.Forst.) P.I.Forst.
**[[Coleus bellus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus betonicifolius]]* (Baker) A.J.Paton
**[[Coleus bifidus]]* (A.J.Paton) A.J.Paton
**[[Coleus bipartitus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus bishopianus]]* (Gamble) Smitha & A.J.Paton
**[[Coleus blakei]]* (P.I.Forst.) P.I.Forst.
**[[Coleus bojeri]]* Benth.
**[[Coleus bolavenensis]]* Suddee, Tagane & Rueangr.
**[[Coleus botryosus]]* (A.J.Paton) A.J.Paton
**[[Coleus bourneae]]* (Gamble) Smitha & A.J.Paton
**[[Coleus bracteatus]]* Dunn
**[[Coleus brazzavillensis]]* A.Chev.
**[[Coleus buchananii]]* (Baker) Brenan
**[[Coleus burorum]]* Chiov.
**[[Coleus caillei]]* A.Chev. ex Hutch. & Dalziel
**[[Coleus calaminthoides]]* (Baker) A.J.Paton
**[[Coleus calcicola]]* Murata
**[[Coleus caldericola]]* (P.I.Forst.) P.I.Forst.
**[[Coleus calycinus]]* (Benth.) A.J.Paton
**[[Coleus cambodianus]]* (Murata) A.J.Paton
**[[Coleus caninus]]* (B.Heyne ex Roth) Vatke
**[[Coleus carnosifolius]]* (Hemsl.) Dunn
**[[Coleus cataractarum]]* (B.J.Pollard) A.J.Paton
**[[Coleus caudatus]]* (S.Moore) E.Downes & I.Darbysh.
**[[Coleus celsus]]* A.J.Paton
**[[Coleus centraliafricanus]]* A.J.Paton
**[[Coleus chevalieri]]* Briq.
**[[Coleus ciliatus]]* (Bramley) A.J.Paton
**[[Coleus circinnatus]]* (Hedge) A.J.Paton & Phillipson
**[[Coleus coeruleus]]* Gürke
**[[Coleus comosus]]* Hochst. ex Gürke
**[[Coleus confertiflorus]]* (A.J.Paton) A.J.Paton
**[[Coleus congensis]]* (Gürke) A.J.Paton
**[[Coleus congestus]]* (R.Br.) A.J.Paton
**[[Coleus conglomeratus]]* (T.C.E.Fr.) Robyns & Lebrun
**[[Coleus crassus]]* (N.E.Br.) Culham
**[[Coleus cremnus]]* (B.J.Conn) A.J.Paton
**[[Coleus cucullatus]]* (A.J.Paton) A.J.Paton
**[[Coleus cuneatus]]* Baker f.
**[[Coleus cyanophyllus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus cylindraceus]]* (Hochst. ex Benth.) A.J.Paton
**[[Coleus daviesii]]* E.A.Bruce
**[[Coleus decimus]]* (A.J.Paton) A.J.Paton
**[[Coleus decurrens]]* Gürke
**[[Coleus deflexifolius]]* (Baker) A.J.Paton
**[[Coleus defoliatus]]* (Hochst. ex Benth.) A.J.Paton
**[[Coleus densus]]* (N.E.Br.) A.J.Paton
**[[Coleus descampsii]]* (Briq.) A.J.Paton
**[[Coleus dewildemanianus]]* (Robyns & Lebrun) A.J.Paton
**[[Coleus dichotomus]]* (A.J.Paton) A.J.Paton
**[[Coleus dinteri]]* (Briq.) A.J.Paton
**[[Coleus dissitiflorus]]* Gürke
**[[Coleus divaricatus]]* A.J.Paton
**[[Coleus diversus]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus dolichopodus]]* (Briq.) A.J.Paton
**[[Coleus dumicola]]* (P.I.Forst.) P.I.Forst.
**[[Coleus dysophylloides]]* (Benth.) A.J.Paton
**[[Coleus efoliatus]]* De Wild.
**[[Coleus elliotii]]* (S.Moore) A.J.Paton
**[[Coleus elongatus]]* Trimen
**[[Coleus eminii]]* (Gürke) A.J.Paton
**[[Coleus engleri]]* (Briq.) A.J.Paton
**[[Coleus erici-rosenii]]* (R.E.Fr.) A.J.Paton
**[[Coleus esculentus]]* (N.E.Br.) G.Taylor
**[[Coleus eungellaensis]]* A.J.Paton
**[[Coleus excelsus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus exilis]]* (A.J.Paton) A.J.Paton
**[[Coleus fasciculatus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus ferricola]]* Phillipson, O.Hooper & A.J.Paton
**[[Coleus foetidus]]* (Benth.) A.J.Paton
**[[Coleus foliatus]]* (A.J.Paton) A.J.Paton
**[[Coleus forsteri]]* (Benth.) A.J.Paton
**[[Coleus fragrantissimus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus fredericii]]* G.Taylor
**[[Coleus fruticosus]]* Wight ex Benth.
**[[Coleus galeatus]]* (Vahl) Benth.
**[[Coleus gamblei]]* (Smitha & Sunojk.) Smitha
**[[Coleus garckeanus]]* Vatke
**[[Coleus geminatus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus gibbosus]]* A.J.Paton
**[[Coleus gigantifolius]]* (Suddee) Suddee
**[[Coleus gillettii]]* (J.K.Morton) A.J.Paton
**[[Coleus glabriflorus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus globosus]]* (Ryding) A.J.Paton
**[[Coleus goetzenii]]* (Gürke) A.J.Paton
**[[Coleus gossweileri]]* A.J.Paton
**[[Coleus gracilipedicellatum]]* (Robyns & Lebrun) A.J.Paton
**[[Coleus gracilis]]* Gürke
**[[Coleus gracillimus]]* (T.C.E.Fr.) Robyns & Lebrun
**[[Coleus graminifolius]]* (Perkins) A.J.Paton
**[[Coleus grandicalyx]]* E.A.Bruce
**[[Coleus grandidentatus]]* (Gürke) A.J.Paton
**[[Coleus graniticola]]* (A.Chev.) A.J.Paton
**[[Coleus gratus]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus graveolens]]* (R.Br.) A.J.Paton
**[[Coleus guerkei]]* (Briq.) A.J.Paton
**[[Coleus gymnostomus]]* Gürke
**[[Coleus habrophyllus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus hadiensis]]* (Forssk.) A.J.Paton
**[[Coleus hairulii]]* Kiew
**[[Coleus hallii]]* (J.K.Morton) A.J.Paton
**[[Coleus harmandii]]* (Doan ex Suddee & A.J.Paton) A.J.Paton
**[[Coleus helferi]]* (Hook.f.) A.J.Paton
**[[Coleus hereroensis]]* (Engl.) A.J.Paton
**[[Coleus hijazensis]]* (Abdel Khalik) A.J.Paton
**[[Coleus humulopsis]]* (B.J.Pollard) A.J.Paton
**[[Coleus hymalis]]* (J.R.I.Wood) A.J.Paton
**[[Coleus idukkianus]]* (J.Mathew, Yohannan & B.J.Conn) Smitha
**[[Coleus igniarioides]]* (Ryding) A.J.Paton
**[[Coleus igniarius]]* Schweinf.
**[[Coleus ignotus]]* (A.J.Paton) A.J.Paton
**[[Coleus inflatus]]* Benth.
**[[Coleus inselbergi]]* (B.J.Pollard & A.J.Paton) A.J.Paton
**[[Coleus insignis]]* (Hook.f.) A.J.Paton
**[[Coleus insolitus]]* (C.H.Wright) Robyns & Lebrun
**[[Coleus insularis]]* (P.I.Forst.) P.I.Forst.
**[[Coleus intraterraneus]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus kanneliyensis]]* L.H.Cramer & S. Balas.
**[[Coleus kanyakumariensis]]* (Shinoj & Sunojk.) Smitha
**[[Coleus kapatensis]]* R.E.Fr.
**[[Coleus kirkii]]* (Baker) A.J.Paton
**[[Coleus kivuensis]]* Lebrun & L.Touss.
**[[Coleus klossii]]* (S.Moore) P.I.Forst. & T.C.Wilson
**[[Coleus koualensis]]* A.Chev. ex Hutch. & Dalziel
**[[Coleus koulikoroensis]]* A.J.Paton
**[[Coleus kunstleri]]* (Prain) A.J.Paton
**[[Coleus lactiflorus]]* Vatke
**[[Coleus laetus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus lageniocalyx]]* Briq.
**[[Coleus lanceolatus]]* (Bojer ex Benth.) A.J.Paton & Phillipson
**[[Coleus lancifolius]]* (Bramley) A.J.Paton
**[[Coleus lanuginosus]]* Hochst. ex Benth.
**[[Coleus lasianthus]]* Gürke
**[[Coleus lateriticola]]* (A.Chev.) Phillipson, O.Hooper & A.J.Paton
**[[Coleus laxus]]* (T.C.Wilson & P.I.Forst.) T.C.Wilson & P.I.Forst.
**[[Coleus leiperi]]* (P.I.Forst.) P.I.Forst.
**[[Coleus linearifolius]]* (J.K.Morton) A.J.Paton
**[[Coleus livingstonei]]* A.J.Paton
**[[Coleus longipetiolatus]]* Gürke
**[[Coleus lyratus]]* (A.Chev.) Roberty
**[[Coleus maculosus]]* (Lam.) A.J.Paton
**[[Coleus madagascariensis]]* (Pers.) A.Chev.
**[[Coleus magnificus]]* P.I.Forst. & A.J.Paton
**[[Coleus malabaricus]]* Benth.
**[[Coleus mannii]]* Hook.f.
**[[Coleus marrubatus]]* (J.K.Morton) A.J.Paton
**[[Coleus megacalyx]]* (A.J.Paton) A.J.Paton
**[[Coleus megadontus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus melleri]]* (Baker) A.J.Paton & Phillipson
**[[Coleus meyeri]]* (Gürke) A.J.Paton
**[[Coleus minor]]* (J.K.Morton) A.J.Paton
**[[Coleus minutiflorus]]* (Ryding) A.J.Paton
**[[Coleus minutus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus mirabilis]]* Briq.
**[[Coleus mirus]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus mitis]]* (R.A.Clement) A.J.Paton
**[[Coleus modestus]]* (Baker) Robyns & Lebrun
**[[Coleus mollis]]* Benth.
**[[Coleus monostachyus]]* (P.Beauv.) A.J.Paton
**[[Coleus mutabilis]]* (Codd) A.J.Paton
**[[Coleus myrianthellus]]* Briq.
**[[Coleus namuliensis]]* E.Downes & I.Darbysh.
**[[Coleus neochilus]]* (Schltr.) Codd
**[[Coleus nepetifolius]]* (Baker) A.J.Paton
**[[Coleus niamniamensis]]* (Gürke) A.J.Paton
**[[Coleus nigericus]]* (Alston) A.J.Paton
**[[Coleus nitidus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus niveus]]* (Hiern) A.J.Paton
**[[Coleus nyikensis]]* Baker
**[[Coleus omissus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus orthodontus]]* (Gürke) A.J.Paton
**[[Coleus otostegioides]]* (Schweinf. ex Gürke) A.J.Paton
**[[Coleus pallidus]]* (Wall. ex Benth.) A.J.Paton
**[[Coleus paniculatus]]* Benth.
**[[Coleus parishii]]* (Hook.f.) A.J.Paton
**[[Coleus parvicalyx]]* (A.J.Paton) A.J.Paton
**[[Coleus penicillatus]]* (A.J.Paton) A.J.Paton
**[[Coleus pentheri]]* Gürke
**[[Coleus perrieri]]* (Hedge) A.J.Paton & Phillipson
**[[Coleus persoonii]]* Benth.
**[[Coleus petiolatissimus]]* Briq.
**[[Coleus petraeus]]* (Backer ex Adelb.) A.J.Paton
**[[Coleus petricola]]* (J.Mathew & B.J.Conn) A.J.Paton
**[[Coleus phulangkaensis]]* (Suddee, Suphuntee & Saengrit) Suddee
**[[Coleus plantagineus]]* (Hook.f.) A.J.Paton
**[[Coleus platyphyllus]]* (A.J.Paton) A.J.Paton
**[[Coleus pobeguinii]]* Hutch. & Dalziel
**[[Coleus polystachyus]]* (Benth.) A.J.Paton
**[[Coleus porcatus]]* (van Jaarsv. & P.J.D.Winter) A.J.Paton
**[[Coleus porphyranthus]]* (T.J.Edwards & N.R.Crouch) A.J.Paton
**[[Coleus prittwitzii]]* (Perkins) A.J.Paton
**[[Coleus prostratus]]* (Gürke) A.J.Paton
**[[Coleus psammophilus]]* (Codd) A.J.Paton
**[[Coleus pseudospeciosus]]* (Buscal. & Muschl.) A.J.Paton
**[[Coleus pulchellus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus rafidahiae]]* Kiew
**[[Coleus recurvata]]* (Ryding) A.J.Paton
**[[Coleus repens]]* Gürke
**[[Coleus reticulatus]]* A.Chev.
**[[Coleus rhodesianum]]* (N.E.Br.) A.J.Paton
**[[Coleus robustus]]* (Hook.f.) A.J.Paton
**[[Coleus rotundifolius]]* (Poir.) A.Chev. & Perrot
**[[Coleus ruandensis]]* (De Wild.) A.J.Paton
**[[Coleus rutenbergianus]]* (Vatke) A.J.Paton & Phillipson
**[[Coleus sahyadricus]]* (Smitha & Sunojk.) Smitha
**[[Coleus sallyae]]* (A.J.Paton) A.J.Paton
**[[Coleus sanguineus]]* (Britten) A.J.Paton
**[[Coleus saxorum]]* (J.Mathew, Yohannan & B.J.Conn) Smitha
**[[Coleus scaber]]* (Benth.) A.J.Paton
**[[Coleus scandens]]* Gürke
**[[Coleus scebeli]]* Chiov.
**[[Coleus schizophyllus]]* (Baker) A.J.Paton
**[[Coleus schliebenii]]* (Mildbr.) A.J.Paton
**[[Coleus scruposus]]* A.J.Paton
**[[Coleus scutellarioides]]* (L.) Benth.
**[[Coleus seretii]]* De Wild.
**[[Coleus sessilifolius]]* (A.J.Paton) A.J.Paton
**[[Coleus shirensis]]* Gürke
**[[Coleus shoolamudianus]]* (Sunil & Naveen Kum.) Smitha & A.J.Paton
**[[Coleus sigmoideus]]* (A.J.Paton) A.J.Paton
**[[Coleus socotranus]]* (Radcl.-Sm.) A.J.Paton
**[[Coleus sphaerocephalus]]* (Baker) A.J.Paton
**[[Coleus splendens]]* (P.I.Forst.) P.I.Forst.
**[[Coleus splendidus]]* A.Chev.
**[[Coleus stachyoides]]* (Oliv.) E.A.Bruce
**[[Coleus steenisii]]* (H.Keng) A.J.Paton
**[[Coleus stenostachys]]* (Baker) A.J.Paton & Phillipson
**[[Coleus strictipes]]* (G.Taylor) A.J.Paton
**[[Coleus strobilifer]]* (Roxb.) A.J.Paton
**[[Coleus stuhlmannii]]* (Gürke) A.J.Paton
**[[Coleus suaveolens]]* (S.T.Blake) P.I.Forst. & T.C.Wilson
**[[Coleus subspicatus]]* (Hochst.) Walp.
**[[Coleus succulentus]]* Pax
**[[Coleus suffruticosus]]* (Wight) A.J.Paton
**[[Coleus sylvestris]]* (Gürke) A.J.Paton & Phillipson
**[[Coleus tenuicaulis]]* Hook.f.
**[[Coleus tetradenifolius]]* (A.J.Paton) A.J.Paton
**[[Coleus tetragonus]]* (Gürke) Robyns & Lebrun
**[[Coleus thalassoscopicus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus thyrsoideus]]* Baker
**[[Coleus togoensis]]* (Perkins) A.J.Paton
**[[Coleus tomentifolius]]* (Suddee) Suddee
**[[Coleus torrenticola]]* (P.I.Forst.) P.I.Forst.
**[[Coleus triangularis]]* (A.J.Paton) A.J.Paton
**[[Coleus trullatus]]* (A.J.Paton) A.J.Paton
**[[Coleus umbrosus]]* Vatke
**[[Coleus unguentarius]]* (Codd) A.J.Paton
**[[Coleus urticifolius]]* Benth.
**[[Coleus velutinus]]* (Trimen) A.J.Paton
**[[Coleus venteri]]* (van Jaarsv. & Hankey) A.J.Paton
**[[Coleus ventosus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus venustus]]* (P.I.Forst.) P.I.Forst.
**[[Coleus verticillatus]]* (Baker) A.J.Paton
**[[Coleus vettiveroides]]* K.C.Jacob
**[[Coleus veyretiae]]* (Guillaumet & A.Cornet) A.J.Paton & Phillipson
**[[Coleus villosus]]* (Forssk.) A.J.Paton
**[[Coleus wallamanensis]]* (T.C.Wilson & P.I.Forst.) T.C.Wilson & P.I.Forst.
**[[Coleus welwitschii]]* Briq.
**[[Coleus xanthanthus]]* C.Y.Wu & Y.C.Huang
**[[Coleus xerophilus]]* (Codd) A.J.Paton
**[[Coleus xylopodus]]* (Lukhoba & A.J.Paton) A.J.Paton
**[[Coleus yemenensis]]* A.J.Paton
**[[Coleus zombensis]]* (Baker) Mwany.

## Rujukan