两眼猪笼草

两眼猪笼草（学名：）是婆罗洲和苏门答腊特有的热带食虫植物McPherson, S.R. 2009. Pitcher Plants of the Old World. 2 volumes. Redfern Natural History Productions, Poole.，少量分布于周围的许多岛屿中，包括邦加島、大納土納島 Mansur, M. 2012. Keanekaragaman jenis tumbuhan pemakan serangga dan laju fotosintesisnya di Pulau Natuna. [Diversity on insectivorous plants and its photosynthetic rate in Natuna Island.] Berita Biologi 11(1): 33–42. Abstract 、尼亞斯島及西比路島。Clarke, C.M. 2001. Appendix C: Distribution Maps. In: Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu. pp. 299–307.虽然部分文献显示其分布于西马来西亚Macfarlane, J.M. 1908. Nepenthaceae. In: A. Engler. Das Pflanzenreich IV, III, Heft 36: 1–91.Danser, B.H. 1928. The Nepenthaceae of the Netherlands Indies. Bulletin du Jardin Botanique de Buitenzorg, Série III, 9(3–4): 249–438.Holttum, R.E. 1940. Malayan pitcher-plants. Malayan Nature Journal 1: 35–44.Shivas, R.G. 1984. Pitcher Plants of Peninsular Malaysia & Singapore. Maruzen Asia, Kuala Lumpur.和新加坡Green, S. 1967. Notes on the distribution of Nepenthes species in Singapore. The Gardens' Bulletin Singapore 22: 53–65.，但这些记录似乎是错误的Adam, J.H., C.C. Wilcock & M.D. Swaine 1992.  Journal of Tropical Forest Science 5(1): 13–25.Clarke, C.M. 2006. Introduction. In: Danser, B.H. The Nepenthaceae of the Netherlands Indies. Natural History Publications (Borneo), Kota Kinabalu. pp. 1–15.。其种加词“reinwardtiana”来源于卡斯帕·格奥尔格·卡尔·赖因瓦特，所以其也称为瑞华德猪笼草。Phillipps, A. & A. Lamb 1996. Pitcher-Plants of Borneo. Natural History Publications (Borneo), Kota Kinabalu.

两眼猪笼草分布于海拔0至2200米处。高海拔地区与低海拔地区两眼猪笼草的捕虫笼大小和颜色存在差异，低海拔变型的捕虫笼更细长，高海拔地区的捕虫笼更矮胖。两眼猪笼草的捕虫笼色彩丰富，可为绿色至深红色。两眼猪笼草捕虫笼的内表面可存在两个“眼点”，其随植株生长状态，或存在或消失。

 分类关系 
 纳奎友丁猪笼草 
2006年，朱马特·哈吉·亚当和哈菲查·A·哈米德描述了纳奎友丁猪笼草。Adam, J.H. & Hafiza A. Hamid 2006.  International Journal of Botany 2(4): 431-436. ISSN 1811-9700 Fong, L.F. 2007. New pitcher species . The Star, May 11, 2007.纳奎友丁猪笼草存在于婆罗洲沙巴由根地咬去往金马尼士的道路及特鲁斯马迪山山脚，海拔1400米至1425米的开阔次生植被中。

虽然两眼猪笼草和纳奎友丁猪笼草之间存在密切的近缘关系，但朱马特·哈吉·亚当和哈菲查·A·哈米德认为“因为两眼猪笼草和纳奎友丁猪笼草存在许多形态上的差异，所以他们不能归为同一个物种”。但是另一些学者认为，两眼猪笼草和纳奎友丁猪笼草之间的差异太小了，不足以将它们区分成两个物种，并认为纳奎友丁猪笼草是两眼猪笼草的同物异名。Rice, B.A. 2006. Do you want to tell me about a species I missed?  The Carnivorous Plant FAQ.Schlauer, J. N.d. Nepenthes naquiyuddinii . Carnivorous Plant Database.此外，这些学者认为纳奎友丁猪笼草可能是暗色猪笼草和两眼猪笼草的自然杂交种。

 进化分支 
2001年，查尔斯·克拉克对来自苏门答腊岛和西马来西亚的猪笼草进行了分支系统学分析，共利用了70个形态特征。以下为猪笼草进化树的“第6分支”。Clarke, C.M. 2001. Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu.

 种下类群 
Nepenthes reinwardtiana var. samarindaensis J.H.Adam & Wilcock (1993)Adam, J.H. & C.C. Wilcock 1993. A new variety of Nepenthes reinwardtiana Miquel from Kalimantan, Borneo. Edinburgh Journal of Botany 50(1): 99–104.

 自然杂交种 

N. albomarginata × N. reinwardtiana [=N. × ferrugineomarginata]Clarke, C.M. 1997. Nepenthes of Borneo. Natural History Publications (Borneo), Kota Kinabalu.
N. ampullaria × N. reinwardtiana
N. clipeata × N. reinwardtiana
N. fusca × N. reinwardtiana[=?N. naquiyuddinii]
N. gracilis × N. reinwardtiana
N. gymnamphora × N. reinwardtiana
N. hispida × N. reinwardtiana
N. macrovulgaris × N. reinwardtiana
N. mirabilis × N. reinwardtianaPhillipps, A., A. Lamb & C.C. Lee 2008. Pitcher Plants of Borneo. Second Edition. Natural History Publications (Borneo), Kota Kinabalu.
N. reinwardtiana × N. spathulata
N. reinwardtiana × N. stenophylla
N. reinwardtiana × N. tentaculata[=?N. murudensis]
N. reinwardtiana × N. tobaica Dariana 2010. Keanekaragaman Nepenthes dan pohon inang di Taman Wisata Alam Sicikeh-Cikeh Kabupaten Dairi Sumatera Utara . M.Sc. thesis, University of North Sumatra, Medan.

 注释 

 参考文献 

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 扩展阅读 

 食虫植物照片搜寻引擎中两眼猪笼草的照片 
 食虫植物照片搜寻引擎中苏门答腊两眼猪笼草的照片 

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reinwardtiana
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