මූසා (ගණය)

පද්ධතිමය සහ වර්ගීකරණය

ඉතිහාසය
Classical Latin briefly had a name for bananas, namely ariena, used by Pliny the Elder (Natural History, XII.12). According to Alexander von Humboldt, that term may have been a borrowing from Sanskrit varana via Greek rendition ouarana, following the Indian campaign of Alexander the Great.

Due to the rarity of bananas in the Late Antiquity and early Middle Ages, the term ariena faded out of use. During the late Middle Ages, international trade brought bananas to Europe, which created the need for a name. In the , Medieval Latin innovated a term musa: this was most likelySome sources assert that Musa is named for Antonius Musa, physician to the Emperor Augustus: see . Pp. 2076–9. the latinization of the Arabic name for the fruit, mauz (موز). Thus, the 11th-century Arabic encyclopedia The Canon of Medicine, which was translated to Latin in medieval times and well known in Europe, shows a correspondence between Arabic mauz and Latin musa. Muz is also the Turkish, Persian, and Somali name for the fruit. 

According to linguist Mark Donohue and archaeologist Tim Denham, the ultimate origin of the Latinized form musa is in the Trans–New Guinea languages, where certain cultivars of bananas are known under a form *muku. From there, the term was borrowed into the Austronesian languages of the area, and migrated across Asia, via the Dravidian languages of India, into Persian, Greek, and Arabic as a Wanderwort:

{|class=wikitable
|+Possible transmission of musa from New Guinea to Latin
|Trans-New Guinea || Austronesian || Dravidian || Indic || Persian || Arabic || Greek || Latin
|-
| #mugu || #mugu > muku > muu || mōttai/mōte || mocā || mōč⁠ || mawz/mawza || mozā || musa
|}

The late Latin term musa was later chosen by Carl Linnaeus in 1753, as the name for the genus. 

From the time of Linnaeus until the 1940s, different types of edible bananas and plantains were given Linnaean binomial names, such as Musa cavendishii, as if they were species. In fact, edible bananas have an extremely complicated origin involving hybridization, mutation, and finally selection by humans. Most edible bananas are seedless (parthenocarpic), hence sterile, so they are propagated vegetatively. The giving of species names to what are actually very complex, largely asexual, hybrids (mostly of two species of wild bananas, Musa acuminata and Musa balbisiana) led to endless confusion in banana botany. In the 1940s and 1950s, it became clear to botanists that the cultivated bananas and plantains could not usefully be assigned Linnean binomials, but were better given cultivar names.

As for the word banana, it came to English from Spanish and Portuguese, which had apparently obtained it from a West African language, possibly Wolof (Senegal).Entry Banana, at Dictionary.com. 

අංශ

Musa sections have a history dating back to 1887, when M.P. Sagot published "Sur le genre Bananier", where the genus Musa was first formally classified. In this article, Sagot arranged the Musa species into three groups, although no section names were assigned to them. The grouping was based on morphological traits, establishing the trio as bananas with fleshy fruit, ornamental bananas with upright inflorescences and bracts that were vibrantly colored, and bananas that were giant in size.

Five years after Sagot's article, J.G. Baker made the first formal designation of Musa sections. To do so, he named three subgenera that almost paralleled the sections that had been described by Sagot. These sections were: 
 M. subg. Physocaulis Baker – defined by a bract with many flowers, inedible fruits, and a bottle-shaped stem
 M. subg. Rhodochlamys Baker – defined by brightly colored bracts with few flowers, usually inedible fruits, and cylindrical stems
 M. subg. Eumusa Baker – defined by green, brown, or dull-violet bracts with many flowers, usually edible fruits, and cylindrical stems.

After this classification, in 1947, Cheeseman reclassified the taxa based on morphological features and chromosome number.Cheesman, E.E. 1947 [pub. 1948]. Classification of the bananas. II. The genus Musa L. Kew Bull. 2: 106–117.  This project proposed four sections: 
 M. sect. Eumusa Cheesman (2n = 2x = 22)
 M. sect. Rhodochlamys (Baker) Cheesman (2n = 2x = 22)
 M. sect. Australimusa Cheesman (2n = 2x = 20)
 M. sect. Callimusa Cheesman (2n = 2x = 20)

The addition of another Musa section came in 1976 by G.C.G. Ardent. The added section, M. sect. Ingentimusa, Ardent was based on a single species, Musa ingens.Argent, G.C.G. 1976. The wild bananas of Papua New Guinea. Notes Roy. Bot. Gard. Edinburgh 35: 77–114. This designation put the number of sections in Musa at five: Eumusa, Rhodochlamys, Callimusa, Australimusa, and Ingentimusa.

In the 21st century, genomics have become cheaper, more efficient, and more accurate, and Musa genetic research has increased exponentially. Research was conducted around a diversity of genomic markers (cpDNA, nrDNA, rDNA, introns, various spacers, etc.). The results of many of these studies suggested that the five sections of Musa defined by morphology (and listed above) were not monophyletic.Wong C, Kiew R, Argent GCG, Set O, Lee SK, Gan YY. Assessment of the validity of the sections in Musa (Musaceae) using AFLP. Ann Bot-London. 2002; 90: 231–238.

Based on the incorrect section grouping, Markku Häkkinen proposed another reclassification of the Musa sections in 2013. Using a multitude of genetic evidence and markers from other studies, Häkkinen suggested the reduction of five Musa sections into two: Musa and Callimusa. Unlike sectional classifications of the past, this hypothesis was based on genetic markers rather than morphological features or chromosome number. The two groups were generally formed by the clustering of the previously defined groups: 
Musa sect. Rhotochlamys and M. sect. Eumusa  became M. sect. Musa
M. sect. Ingetimusa, M. sect. Callimusa and M. sect. Australimusa became M. sect. Callimusa

The advance of genomic analysis technologies and further data on the relatedness of Musa species, formulated Häkkinen's two sections and later corroborated them as correct subcategories for the genus.Feng, Huimin, et al. "Molecular Phylogeny of Genus Musa Determined by Simple Sequence Repeat Markers." Plant Genetic Resources, vol. 14, no. 3, 2015, pp. 192–199.,  The history of Musa sections provides an example of genomics superseding morphological evidence and thus classifications.

විශේෂ
The World Checklist of Selected Plant Families accepts 68 species and two primary hybrids, , which are listed below. The assignment to sections is based on GRIN (where this gives the species), regrouped according to Wong et al.

Section Callimusa (incorporating Australimusa)
[A] and [C] indicate known placement in the former sections Australimusa and Callimusa, respectively.
M. × alinsanaya R.V.Valmayor [A]
M. azizii Häkkinen
M. barioensis Häkkinen
M. bauensis Häkkinen & Meekiong [C]
M. beccarii N.W.Simmonds [A]
M. boman Argent [A]
M. borneensis Becc. [C]
M. bukensis Argent [A] 
M. campestris Becc. [C]
M. coccinea Andrews [C] – scarlet banana
M. exotica R.V.Valmayor [C]
M. fitzalanii F.Muell. [A] – extinct
M. gracilis Holttum [C]
M. hirta Becc. [A]
M. insularimontana Hayata [A]
M. jackeyi W.Hill [A]
M. johnsii Argent [A]
M. lawitiensis Nasution & Supard. [C]
M. lokok Geri & Ng
M. lolodensis Cheesman [A]
M. maclayi F.Muell. ex Mikl.-Maclay [A]
M. monticola M.Hotta ex Argent [A]
M. muluensis M.Hotta [A]
M. paracoccinea A.Z.Liu & D.Z.Li [C]
M. peekelii Lauterb. [A]
M. salaccensis Zoll. ex Backer [A]
M. textilis Née [A] – Abacá
M. × troglodytarum L. [A] – the cultivated Fe'i bananas
M. tuberculata M.Hotta [A]
M. violascens Ridl. [C]
M. viridis R.V.Valmayor et al.
M. voonii Häkkinen

Section Ingentimusa
M. ingens N.W.Simmonds

මූසා කොටස (රොඩොක්ලැමයිස් ඇතුලත් කිරීම)
M. acuminata Colla – wild seeded banana, one of the two main ancestors of modern edible banana cultivars
M. acuminata subsp. zebrina [= M. sumatrana] – blood banana
M. aurantiaca G.Mann ex Baker
M. balbisiana Colla – wild seeded banana, one of the two main ancestors of modern edible banana cultivars
M. banksii F.Muell.
M. basjoo Siebold & Zucc. ex Iinuma – Japanese fiber banana, hardy banana
M. cheesmanii N.W.Simmonds
M. chunii Häkkinen
M. griersonii Noltie
M. itinerans Cheesman
M. mannii H.Wendl. ex Baker
M. nagensium Prain
M. ochracea K.Sheph.
M. ornata Roxb.
Musa × paradisiaca L. = M. acuminata × M. balbisiana – many of the cultivated edible bananas
M. rosea Baker
M. rubinea Häkkinen & C.H.Teo
M. rubra Wall. ex Kurz (syn. Musa laterita Cheesman)
M. sanguinea Hook.f.
M. schizocarpa N.W.Simmonds
M. siamensis Häkkinen & Rich.H.Wallace
M. sikkimensis Kurz
M. thomsonii (King ex Baker) A.M.Cowan & Cowan
M. velutina H.Wendl. & Drude – pink banana
M. yunnanensis Häkkinen & H.Wang – Yunnan banana, wild forest banana
M. zaifui Häkkinen & H.Wang

Section undetermined or unknown

M. arfakiana Argent
M. arunachalensis A.Joe, Sreejith & M.Sabu
M. celebica Warb. ex K.Schum.
M. juwiniana Meekiong
M. kattuvazhana K.C.Jacob
M. lanceolata Warb. ex K.Schum.
M. lutea R.V.Valmayor et al.
M. sakaiana Meekiong et al.
M. splendida A.Chev.
M. tonkinensis R.V.Valmayor et al.
M. yamiensis C.L.Yeh & J.H.Chen

Formerly placed here
Ensete davyae (Stapf) Cheesman (as M. davyae Stapf)
Ensete gilletii (De Wild.) Cheesman (as M. gilletii De Wild. or M. martretiana A.Chev.)
Ensete glaucum (Roxb.) Cheesman (as M. glauca Roxb.)
Ensete lasiocarpum (Franch.) Cheesman (as M. lasiocarpa Franch.) – also placed in a separate genus as Musella lasiocarpa (Franch.) C.Y.Wu ex H.W.Li
Ensete livingstoniana (J. Kirk) Cheesman (as M. livingstoniana J.Kirk)
Ensete perrieri (Stapf) Cheesman (as M. perrieri Claverie)
Ensete superbum (Roxb.) Cheesman (as M. superba Roxb.)
Ensete ventricosum (Welw.) Cheesman (as M. arnoldiana De Wild., M. ensete J.F.Gmel. or M. ventricosum (Welw.) Cheesman)
Heliconia bihai (L.) L. (as M. bihai L.)