エゾフネガイ属

エゾフネガイ属(英:crepidula)、別名スリッパカタツムリ属、スリッパカサガイ属、スリッパシェル属、カリプトレイド属として知られているのは軟体動物門腹足綱に属し、浅海に生息する貝の分類群のことであるBouchet, P.; Gofas, S. (2011). Crepidula Lamarck, 1799. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=137722 on 2011-05-21。本属はに属し、同じ科に属する主なものは、、 、、エゾフネガイ属および属があるCollin, Rachel. 2003. The Utility of Morphological Characters in Gastropod Phylogenetics: An Example From the Calyptraeidae. Biological Journal of the Linnean Society, 4: 541–593。本属名のエゾフネガイは近年Grandicrepidula属に分類されるようになった。

また、種名を省略してCrepidulaと呼ばれることもある。エゾフネガイ属の種の定義における最近の変更は、DNA配列のデータと解剖学的研究の両方に基づいている。 

多くの解剖により、近縁な属との違いが明らかになった。細胞などの組織だけを調べた場合、これらの間の違いは明らかではないが、3つの遺伝子（COI、16S、および28S）のDNA配列によって区別されている。また、右の画像のように、縦に重なって生息している種もいくつかあるBouchet, P.; Gofas, S. (2011). Crepidula Lamarck, 1799. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=137722 on 2011-05-21。

 研究 
エゾフネガイ属は、その中のさまざまな種が発達、生態学、行動の研究で一般的に利用されているHenry, Jonathan J.; Collin, Rachel; Perry, Kimberly J.. 2010. The Slipper Snail, Crepidula: An emerging Lophotrochozoan model system. Biological Bulletin, 218(3): 211-229.。主な研究は、海洋無脊椎動物における大規模な性転換に関するもので、性転換が環境によるものであることを実証するために使用されてきたMérot, Claire; Collin, Rachel. 2012. Effects of food availability on sex change in two species of Crepidula (Gastropoda: Calyptraeidae). Marine Ecology Progress Series, 449: 173-181.Collin, Rachel; McLellan, Michelle; Gruber, Karl F.; Bailey-Jourdain, Catherine. 2005. Effects of Conspecific Associations on Size at Sex Change in Three Species of Calyptraeid Gastropods. Marine Ecology Progress Series, 1: 89-97.Collin, Rachel. 1995. Sex, size, and position: a test of models predicting size at sex change in the protandrous gastropod crepidula fornicata. The American Naturalist, 146(6): 815-831.。エゾフネガイ属のネコゼフネガイ(Crepidula fornicata)とシマメノウフネガイは、海洋生息地における外来種の研究によく利用されたCoe, Wesley R. 1948. Nutrition and sexuality in protandric gastropods of the genus Crepidula. Biol. Bul. 94(3): 158-160.Hendler, G. and D. R. Franz. 1971. Population dynamics and life history of Crepidula convexa Say (Gastropoda: Prosobranchia) in Delaware Bay. Biol. Bul. 141: 514-526.。

エゾフネガイ属の多くの種の単純な殻のために、カリバガサガイ上科の分類は困難である。多くの場合、似たような殻を持つ別々の種は、普遍的または局地的な分布を持つ。この場合、種の同一性か違いを検証するためには、個体の発生またはDNAデータの綿密な調査が不可欠である。

 生態 

 食物 
すべての腹足類と同様に、エゾフネガイ属はろ過摂食動物である。成虫は大きな鰓を使って浮遊物から微細藻類をろ過し食べるが、小さな幼体も歯舌を使って基質から藻類をこすり落とすことができるという証拠がいくつかある。

 生活 
エゾフネガイ属の一部の種は数体が大きい個体が下部に、小さい個体が上部に、縦に重なって生息している。この際、下部にいる個体はメスとなり、上部にいる個体はオスとなる。また、真ん中の方の個体は性が定まっていない。個体の順序や成熟度などが変わった時、性転換がおこる。

 外来種 
エゾフネガイ属は各地で外来種として、その土地の生態系を脅かしているものが多い 。

ネコゼフネガイ(Crepidula fornicata)は、北アメリカの東海岸、フロリダ北部からカナダにまでが原産地である。この種は現在世界中に広まっており、スペインとフランスの北海岸、イギリスの海岸の大部分、そして北海沿岸で非常に侵略的な外来種であると考えられている。サンフランシスコ湾、ピュージェット湾、地中海からも生息が報告されている。Crepidula convexaは、北アメリカの東海岸に自生し、サウスカロライナからニューイングランドまでが原産地である。この種は、ピュージェット湾とサンフランシスコ湾から生息が報告されている。

他に、シマメノウフネガイ(Crepidula onyx)は、カリフォルニアの南海岸とメキシコの北太平洋沿岸が原産地である。この種は現在広く拡散しており、アジアで非常に侵略的な外来種だと考えられている。韓国、日本、香港など東アジアから生息が報告されている。

 種 
サムネイル|240x240ピクセル| Crepidula walshi、現在はSiphopatella walshiと呼ばれる種。
サムネイル|261x261ピクセル| Crepidula immersa、現在はMaoricyptaまたはZeacryptaと呼ばれる種。
エゾフネガイ属の種は次のとおりである Crepidula at ITISRoss, Landon T. (1965). "The genus Crepidula in North America: its taxonomy, phylogeny, and distribution. unpublished Florida State Univ. thesis". Retrieved on May 2009. Crepidula at Discover Life  。なお、種名を斜体で示す。

 Crepidula adunca G. B. Sowerby I, 1825
 Crepidula aeola Dall, 1927Rosenberg, G. (2010). Crepidula aeola Dall, 1927. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=532681 on 2011-01-11
 Crepidula aplysioides Reeve, 1859Rosenberg, G. (2010). Crepidula aplysioides Reeve, 1859. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=532682 on 2011-01-11
 Crepidula arenata Broderip, 1834
 Crepidula argentina Simone, Pastorino & Penchaszadeh, 2000Rosenberg, G. (2010). Crepidula argentina Simone, Pastorino & Penchaszadeh, 2000. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=532683 on 2011-01-11
 Crepidula atrasolea Collin, 2000WoRMS (2010). Crepidula atrasolea Collin, 2000. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=419702 on 2011-01-11
 Crepidula badisparsa Collin, 2005
 Crepidula cachimilla Cledon, Simone & Penchaszadeh, 2004WoRMS (2010). Crepidula cachimilla Cledon, Simone & Penchaszadeh, 2004. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=388656 on 2011-01-11
 Crepidula carioca Simone, 2006WoRMS (2010). Crepidula carioca Simone, 2006. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=389784 on 2011-01-11
 Crepidula cerithicola (C.B.Adams, 1852)
 Crepidula complanata Krauss, 1848
 Crepidula convexa Say, 1822Rosenberg, G. (2010). Crepidula convexa Say, 1822. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=160228 on 2011-01-11 - synonym: Crepidula glauca
 Crepidula coquimbensis Brown & Olivares, 1996Bouchet, P. (2010). Crepidula coquimbensis Brown & Olivares, 1996. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=510048 on 2011-01-11
 Crepidula cymbaeformis Conrad, 1844Rosenberg, G. (2010). Crepidula cymbaeformis Conrad, 1844. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=532684 on 2011-01-11
 Crepidula depressa Say, 1822WoRMS (2010). Crepidula depressa Say, 1822. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=419703 on 2011-01-11
 Crepidula derjugini Golikov & Kussakin, 1962
 Crepidula excavata Broderip, 1834
 Crepidula fimbriata Reeve, 1859
 ネコゼフネガイ Crepidula fornicata (Linnaeus, 1758)Gofas, S. (2010). Crepidula fornicata (Linnaeus, 1758). In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138963 on 2011-01-11 - Common slipper shell, common Atlantic slippersnail, boat shell, quarterdeck shell.
 Crepidula glottidiarum Dall, 1905
 Crepidula goreensis Gmelin, 1791
 Crepidula huertae Collin, 2019
 Crepidula incurva (Broderip, 1834)
 Crepidula intratesta Simone, 2006WoRMS (2010). Crepidula intratesta Simone, 2006. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=389785 on 2011-01-12
 Crepidula lessonii (Broderip, 1834)
 Crepidula maculosa Conrad, 1846WoRMS (2010). Crepidula maculosa Conrad, 1846. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=419704 on 2011-01-12
 Crepidula margarita Simone, 2006WoRMS (2010). Crepidula margarita Simone, 2006. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=389786 on 2011-01-12
 Crepidula marginalis  Broderip, 1834
 Crepidula moulinsii Michaud, 1829Gofas, S. (2010). Crepidula moulinsii Michaud, 1829. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=180993 on 2011-01-12
 Crepidula naticarum Williamson, 1905
 Crepidula navicella (Lesson, 1831)
 Crepidula navicula (Mörch, 1877)Rosenberg, G. (2010). Crepidula navicula (Mörch, 1877). In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=532685 on 2011-01-12
 Crepidula nivea C.B. Adams, 1852
 Crepidula norrisiarum Williamson, 1905
 Crepidula nummaria Gould, 1846
 シマメノウフネガイ Crepidula onyx G. B. Sowerby I, 1824WoRMS (2010). Crepidula onyx G. B. Sowerby I, 1824. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=397063 on 2011-01-12 - onyx slippersnail
 Crepidula perforans Valenciennes, 1846
 "Crepidula philippiana" Gallardo, 1977
 Crepidula plana Say, 1822Rosenberg, G. (2010). Crepidula plana Say, 1822. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=160230 on 2011-01-12
 Crepidula porcellana (Linnaeus, 1758)Gofas, S. (2010). Crepidula porcellana (Linnaeus, 1758). In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138965 on 2011-01-12
 Crepidula protea (d'Orbigny, 1841)Rosenberg, G. (2010). Crepidula protea (d'Orbigny, 1841). In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=532686 on 2011-01-12
 Crepidula pyguaia Simone, 2006WoRMS (2010). Crepidula pyguaia Simone, 2006. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=389787 on 2011-01-12
 Crepidula rostrata C. B. Adams, 1852
 Crepidula striolata Menke, 1851
 Crepidula unguiformis Lamarck, 1822Gofas, S. (2010). Crepidula unguiformis Lamarck, 1822. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138966 on 2011-01-12
 Crepidula ustulatulina Collin, 2002WoRMS (2010). Crepidula ustulatulina Collin, 2002. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=387736 on 2011-01-12
 Crepidula williamsi Coe, 1947
 Crepidula wolfae Collin, 2019

 同じ種名が命名された種

 Crepidula aculeata (Gmelin, 1791): synonym of Bostrycapulus aculeatus (Gmelin, 1791)
 Crepidula aculeata (Gmelin, 1791) sensu Zenetos et al.2004: synonym of Bostrycapulus odites Collin, 2005
 Crepidula acuta H. C. Lea, 1842: synonym of Crepidula convexa Say, 1822
 Crepidula adspersa Dunker, 1846: synonym of Crepidula porcellana Lamarck, 1801
 Crepidula bilobata Reeve, 1859: synonym of Crepidula lingulata Gould, 1846: synonym of Crepipatella lingulata (Gould, 1846)
 Crepidula calceolina Deshayes, 1830: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula californica Tryon, 1886: synonym of Crepidula aculeata (Gmelin, 1791): synonym of Bostrycapulus aculeatus (Gmelin, 1791)
 Crepidula calyptraeiformis Deshayes, 1830: synonym of Bostrycapulus calyptraeiformis (Deshayes, 1830)
 Crepidula candida Risso, 1826: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula capensis Quoy & Gaimard, 1835: synonym of Crepipatella capensis (Quoy & Gaimard, 1835)
 Crepidula cataldi Parenzan, 1970: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula coei Berry, 1950: synonym of Crepidula naticarum Williamson, 1905
 Crepidula contorta Quoy & Gaimard, 1832: synonym of Maoricrypta monoxyla (Lesson, 1830)
 Crepidula costata G. B. Sowerby I, 1824: synonym of Maoricrypta costata (G. B. Sowerby I, 1824)
 Crepidula costata Deshayes, 1830: synonym of Maoricrypta costata (G. B. Sowerby I, 1824)
 Crepidula costulata Dunker, 1852: synonym of Crepidula incurva (Broderip, 1834)
 Crepidula crepidula (Linnaeus, 1767): synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula depressa Deshayes, 1830: synonym of Crepidula dilatata Lamarck, 1822: synonym of Crepipatella dilatata (Lamarck, 1822)
 Crepidula deshayesii de Folin, 1867: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula desmoulinsi Locard, 1886: synonym of Crepidula moulinsii Michaud, 1829
 Crepidula dilatata Lamarck, 1822: synonym of Crepipatella dilatata (Lamarck, 1822)
 Crepidula dorsata (Broderip, 1834): synonym of Crepipatella dorsata (Broderip, 1834)
 Crepidula excisa Philippi, 1849: synonym of Ergaea walshi (Reeve, 1859)
 Crepidula explanata Gould, 1853: synonym of Crepidula perforans (Valenciennes, 1846)
 Crepidula exuviata Reeve, 1859: synonym of Crepidula perforans (Valenciennes, 1846)
 Crepidula fasciata de Roissy, 1805: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula fecunda Gallardo, 1979: synonym of Crepipatella peruviana (Lamarck, 1822)
 Crepidula fissurata G. B. Sowerby II, 1883: synonym of Crepipatella lingulata (Gould, 1846)
 Crepidula foliacea Broderip, 1834: synonym of Crepidula aculeata (Gmelin, 1791): synonym of Bostrycapulus aculeatus (Gmelin, 1791)
 Crepidula gibbosa Defrance, 1818: synonym of Crepidula moulinsii Michaud, 1829
 Crepidula glauca Say, 1822: synonym of Crepidula convexa Say, 1822
 Crepidula grandis Middendorff, 1849: synonym of Grandicrepidula grandis (Middendorff, 1849)
 Crepidula gravispinosa Kuroda & Habe, 1950: synonym of Crepidula aculeata (Gmelin, 1791): synonym of Bostrycapulus aculeatus (Gmelin, 1791)
 Crepidula hochstetteriana Wilkens, 1922 : synonym of † Maoricrypta radiata (Hutton, 1873)
 Crepidula immersa Angas, 1865: synonym of Maoricrypta immersa (Angas, 1865)
 Crepidula incurva Zittel, 1864: synonym of † Crepidula wilckensi Finlay, 1924 : synonym of Maoricrypta radiata (Hutton, 1873) †
 Crepidula italica Defrance, 1818: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula lamina H. C. Lea, 1843: synonym of Crepidula plana Say, 1822
 Crepidula lingulata Gould, 1846: synonym of Crepipatella lingulata (Gould, 1846)
 Crepidula lirata Reeve, 1859: synonym of Crepidula incurva (Broderip, 1834)
 Crepidula maculata Quoy & Gaimard, 1832:  synonym for Sigapatella novaezelandiae.
 Crepidula minuta Middendorff, 1849: synonym of Crepidula nummaria Gould, 1846
 Crepidula monoxyla (Lesson, 1831): synonym of Maoricrypta monoxyla (Lesson, 1830)
 Crepidula nautiloides Lesson, 1832: synonym of Crepipatella dilatata (Lamarck, 1822)
 Crepidula navicelloides Carpenter, 1864: synonym of Crepidula nummaria Gould, 1846
 Crepidula nebulata Mabille, 1895: synonym of Crepidula nivea C. B. Adams, 1852
 Crepidula orbella Yokoyama, 1920: synonym of Ergaea walshi (Reeve, 1859)
 Crepidula orbiculata Dall, 1919: synonym of Crepipatella orbiculata (Dall, 1919)
 Crepidula osculans C. B. Adams, 1852: synonym of Phenacolepas osculans (C. B. Adams, 1852)
 Crepidula pallida (Broderip, 1834): synonym of Calyptraea pallida Broderip, 1834
 Crepidula patagonica d'Orbigny, 1841: synonym of Crepipatella patagonica (d'Orbigny, 1841)
 Crepidula patula Deshayes, 1830: synonym of Crepipatella peruviana (Lamarck, 1822)
 Crepidula peruviana Lamarck, 1822: synonym of Crepipatella peruviana (Lamarck, 1822)
 Crepidula plana (A. Adams & Reeve, 1850): synonym of Ergaea walshi (Reeve, 1859)
 Crepidula pulchella Aradas, 1846: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula rhyssema Olsson & Harbison, 1953: synonym of Crepidula plana Say, 1822
 Crepidula riisei Dunker, 1852: synonym of Crepidula fornicata (Linnaeus, 1758)
 Crepidula rostriformis Gould, 1846: synonym of Crepidula adunca G. B. Sowerby I, 1825
 Crepidula rugosa Carpenter, 1856: synonym of Crepidula onyx G. B. Sowerby I, 1824
 Crepidula rugulosa Dunker, 1846: synonym of Crepipatella capensis (Quoy & Gaimard, 1835)
 Crepidula sandaliformis de Serres, 1830: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula sandalina Deshayes, 1833: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula scabies Reeve, 1859: synonym of Ergaea walshi (Reeve, 1859)
 Crepidula sinuosa Turton, 1825: synonym of Crepidula unguiformis Lamarck, 1822
 Crepidula solida Hinds, 1844: synonym of Crepidula adunca G. B. Sowerby I, 1825
 Crepidula squama (Broderip, 1834): synonym of Crepidula striolata Menke, 1851
 Crepidula strigellata Dunker, 1853: synonym of Crepidula striolata Menke, 1851
 Crepidula sulin Dautzenberg, 1912: synonym of Crepidula porcellana Lamarck, 1801
 Crepidula tomentosa Quoy & Gaimard, 1832: synonym of Crepidula aculeata (Gmelin, 1791): synonym of Bostrycapulus aculeatus (Gmelin, 1791)
 Crepidula uncata Menke, 1847: synonym of Crepidula adunca G. B. Sowerby I, 1825
 Crepidula uncinata Philippi, 1887: synonym of Crepidula dilatata Lamarck, 1822: synonym of Crepipatella dilatata (Lamarck, 1822)
 Crepidula virginica Conrad, 1871: synonym of Crepidula fornicata (Linnaeus, 1758)
 Crepidula walshi Reeve, 1859: synonym of Ergaea walshi (Reeve, 1859)
 † Crepidula wilckensi Finlay, 1924: synonym of  † Maoricrypta radiata (Hutton, 1873)
 Crepidula youngi (Powell, 1940): synonym of Maoricrypta youngi Powell, 1940

以前はエゾフネガイ属に分類されていたが、その後、属から除外あるいは削除された種には、次のものがある。

 Crepidula costata is a synonym for Maoricrypta costata - ribbed slipper shell
 Crepidula youngi Powell, 1940
 Crepidula wilckensi Finlay, 1924
 Crepidula immersa G. F. Angas, 1867

 Crepidula grandis Middendorff, Middendorff, 1849Gofas, S. (2010). Crepidula grandis Middendorff, 1849. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=254469 on 2011-01-12
 Grandicrepidula collinae Marshall
 Crepidula walshi L. A. Reeve, 1859
 Crepidula maculata Quoy & Gaimard, Gmelin, 1791

 出典 

 参考文献 
 Collin, R. 2003. The Utility of Morphological Characters in Gastropod Phylogenetics: An Example From the Calyptraeidae. Biological Journal of the Linnean Society, 4: 541–593
 Collin, R. 2003. Phylogenetic Relationships Among Calyptraeid Gastropods and Their Implications for the Biogeography of Marine Speciation. Systematic Biology, 5: 618-640.
 Collin, R. 2002. Another Last Word on Crepidula Convexa With a Description of C. Ustulatulina N. Sp. (Gastropoda: Calyptraeidae) From the Gulf of Mexico and Southern Florida. Bulletin of Marine Science, 1: 177-184.
 Collin, R. 2000. Phylogeny of the Crepidula plana (Gastropoda: Calyptraeidae) cryptic species complex in North America. Canadian Journal of Zoology. 78: 1500-1514.
 Hoagland, K.E. 1977. Systematic review of fossil and recent Crepidula and discussion of the evolution of the Calyptraeidae. Malacologia 16(2): 353-420
 Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). . XII, 195 pp. (
 Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213

 関連項目 
エゾフネガイ
 外部リンク 
 Hoagland KE1983。パリの国立自然史博物館にあるCrepidula（前鰓類：Calyptraeidae）のタイプ標本に関する注記。フィラデルフィア自然科学アカデミーの議事録135：1–8 。
 calyptraeid分類学のpfds

Category:カリバガサガイ科