Agriotes

Agriotes er en slekt av biller som hører til familien smellere (Elateridae).

Utseende 
Middelsstore, litt brede, brunlige til svarte smellere. Antennene er gjerne sagtakkede, pronotum hvelvet med rundede sider, hårete.

Larvene er sylindriske, glatte med en voksaktige overflate og spredte hår, gjerne rød- eller gulaktige. Hodet er noe flatt med fremoverstrakte kjever. Det bakerste leddet har vanligvis et par kraftige utvekster av varierende form. Beina er ganske korte.

Levevis 
De voksne billene flyr godt og treffes ofte flygende, på blomster eller i vegetasjonen. Agriotini har larver som lever i jorda og noen kan gjøre betydelig skade på planterøtter.

Utbredelse 
Slekten er utbredt i Nord-, Mellom- og Sør-Amerika, Asia og Europa.

Systematisk inndeling 

 

 Ordenen biller, Coleoptera
 Underordenen Polyphaga
 Overfamilien smellere, Elateroidea
 Familien smellere, Elateridae
 Underfamilien storsmellere, Elaterinae Leach, 1815
 Stammen Agriotini Laporte de Castelnau, 1840 - noen ganger regnet til underfamilien Melanotinae
 Understammen Agriotina
 Slekten Agriotes Eschscholtz, 1829 - ca. 400 arter, i Norge:
 strandsmeller Agriotes sputator (Linnaeus, 1758)
 stripesmeller Agriotes lineatus (Linnaeus, 1767)
 åkersmeller Agriotes obscurus (Linnaeus, 1758)

Liste over øvrige arter 
 Agriotes acuminatus (Stephens, 1830)
 Agriotes acutus Champion, 1896
 Agriotes adanensis Pic, 1910
 Agriotes adlbaueri Platia & Schimmel, 1992
 Agriotes aequalis Schwarz, 1891
 Agriotes agonischioides Fleutiaux, 1918
 Agriotes alternus Candèze, 1863
 Agriotes amabilis Candèze, 1863
 Agriotes anachoretus (Ménétriés, 1832)
 Agriotes anatolicus Platia, 2003
 Agriotes andalusiacus Franz, 1967
 Agriotes angustatus Miwa, 1928
 Agriotes angustatus Champion, 1896
 Agriotes angustus Fleutiaux, 1939
 Agriotes apicalis LeConte, 1884
 Agriotes apicalis (Candèze, 1857)
 Agriotes aquilus Platia, 2003
 Agriotes arcanus Brown, 1933
 Agriotes asaokai Ôhira, 1994
 Agriotes audisioi Platia & Schimmel, 1992
 Agriotes australasiae Blanchard, 1853
 Agriotes australis Fairmaire, 1883
 Agriotes avulsus (LeConte, 1853)
 Agriotes babanus Kishii, 1989
 Agriotes bagherii Platia, Furlan & Gudenzi, 2002
 Agriotes barkulensis Jagemann, 1942
 Agriotes barriesi Cate & Platia, 1997
 Agriotes belfragei Becker, 1956
 Agriotes bicolor Candèze, 1863
 Agriotes bimaculatus (Candèze, 1857)
 Agriotes binhus Fleutiaux, 1939
 Agriotes binotatus Champion, 1896
 Agriotes biplagiatus Klug
 Agriotes birmanicus Fleutiaux, 1942
 Agriotes bivittatus Van Dyke, 1932
 Agriotes bogatschevi Dolin, 1969
 Agriotes bonnairei Buysson, 1890
 Agriotes bonvouloiri (Candèze, 1874)
 Agriotes brevis Candèze, 1863
 Agriotes breviusculus Candèze, 1863
 Agriotes brunneipennis (Fleutiaux, 1902)
 Agriotes brunneus Schaeffer, 1916
 Agriotes bulgaricus Platia & Gudenzi, 2007
 Agriotes caecus Knull, 1959
 Agriotes caspicus Heyden, 1883
 Agriotes castaneipennis Champion, 1896
 Agriotes chilensis Schwarz, 1902
 Agriotes cilicensis Platia & Gudenzi, 1998
 Agriotes cinereiventris Champion, 1896
 Agriotes colchicus Gurjeva, 1975
 Agriotes collaris (LeConte, 1853)
 Agriotes colonnellii Guglielmi & Platia, 1985
 Agriotes concolor (Fleutiaux, 1925)
 Agriotes confusus (Fleutiaux, 1902)
 Agriotes connexivus Schwarz, 1891
 Agriotes conspicuus Schwarz, 1891
 Agriotes constrictus Reitter, 1900
 Agriotes coomani Fleutiaux, 1939
 Agriotes corsicus Candèze, 1863
 Agriotes crassiusculus Platia & Gudenzi, 1998
 Agriotes cribrithorax Pic, 1910
 Agriotes criddlei Van Dyke, 1932
 Agriotes cruciatus Champion, 1896
 Agriotes curticollis Champion, 1896
 Agriotes curtus Candèze, 1878
 Agriotes cylindricollis Schwarz, 1902
 Agriotes cylindricus Van Dyke, 1932
 Agriotes defreinai Platia & Gudenzi, 1998
 Agriotes dehiscans Gistel, 1857
 Agriotes deletus Fleutiaux, 1939
 Agriotes depressus Schwarz, 1891
 Agriotes desbrochersi Buysson, 1890
 Agriotes dilataticoxis Buysson, 1904
 Agriotes dualis Fleutiaux, 1939
 Agriotes dubius Fleutiaux, 1907
 Agriotes duporti Fleutiaux, 1939
 Agriotes dusaneki Platia & Gudenzi, 1998
 Agriotes elegantissimus (Candèze, 1881)
 Agriotes elegantulus Lewis, 1894
 Agriotes ellenicus Cate & Platia, 1997
 Agriotes emaciatus Platia & Gudenzi, 1997
 Agriotes espinosus Becker, 1956
 Agriotes fairmairei (Candèze, 1857)
 Agriotes ferrugineipennis (LeConte, 1861)
 Agriotes flavobasalis Heyden, 1889
 Agriotes francki (Fleutiaux, 1934)
 Agriotes fucosus (LeConte, 1853)
 Agriotes fulgens Ôhira, 1966
 Agriotes fulvescens Candèze, 1863
 Agriotes fulvus Fleutiaux, 1939
 Agriotes furlani Platia, 2003
 Agriotes fusiformis Candèze, 1876
 Agriotes gallicus Lacordaire, 1835
 Agriotes ganglbaueri Schwarz, 1891
 Agriotes germaini Fleutiaux, 1907
 Agriotes goksunensis Platia, 2004
 Agriotes gracilis (Miwa, 1931)
 Agriotes graecus Franz, 1967
 Agriotes grandicollis Fleutiaux, 1939
 Agriotes grandinii Candèze, 1863
 Agriotes granulosus Platia & Schimmel, 1992
 Agriotes gratiosus (Fleutiaux, 1925)
 Agriotes gratiosus Fleutiaux, 1925
 Agriotes guadulpensis Candèze, 1863
 Agriotes guatemalensis Champion, 1896
 Agriotes guatemalensis (Fleutiaux, 1940)
 Agriotes gurgistanus (Faldermann, 1835)
 Agriotes hedini Fleutiaux, 1936
 Agriotes heydeni Schwarz, 1891
 Agriotes hilaris Candèze, 1863
 Agriotes hirayamai Miwa, 1934
 Agriotes hirsutus Champion, 1896
 Agriotes hoodi Becker, 1956
 Agriotes imperfectus LeConte, 1884
 Agriotes incallidus Champion, 1896
 Agriotes incognitus Schwarz, 1891
 Agriotes informis Schwarz, 1891
 Agriotes infuscatus Desbrochers des Loges, 1870
 Agriotes insanus Candèze, 1863
 Agriotes insolitus Champion, 1896
 Agriotes integricollis Reitter, 1911
 Agriotes intermedius (Golbach, 1976)
 Agriotes interstitialis Fleutiaux, 1939
 Agriotes iranicus Platia, Furlan & Gudenzi, 2002
 Agriotes iraqensis Platia & Gudenzi, 1997
 Agriotes isabellinus (Melsheimer, 1845)
 Agriotes izmirensis Cate & Platia, 1997
 Agriotes jeanvoinei (Fleutiaux, 1939)
 Agriotes kambaitinus Fleutiaux, 1942
 Agriotes kamus Fleutiaux, 1939
 Agriotes karsantianus Pic, 1910
 Agriotes kinzelbachi Platia & Schimmel, 1994
 Agriotes kirghisicus Yablokov-Khnzoryan, 1970
 Agriotes koltzei Reitter, 1890
 Agriotes kraatzi Schwarz, 1891
 Agriotes kubani Platia & Gudenzi, 1997
 Agriotes laevicarinatus Platia & Gudenzi, 1999
 Agriotes laoticus Fleutiaux, 1939
 Agriotes lapicida Faldermann, 1835
 Agriotes lateralis Candèze, 1878
 Agriotes leinfesti Platia & Gudenzi, 1998
 Agriotes leucophaeatus Candèze, 1873
 Agriotes leuthneri Platia & Gudenzi, 1997
 Agriotes ligatus Candèze, 1878
 Agriotes limosus (LeConte, 1853)
 Agriotes linearis Fleutiaux, 1939
 stripesmeller Agriotes lineatus (Olivier, 1790)
 Agriotes lineipennis Candèze, 1863
 Agriotes litigiosus (P. Rossi, 1792)
 Agriotes liukuiensis Kishii, 1989
 Agriotes lizleri Platia, 2003
 Agriotes longipennis Candèze, 1863
 Agriotes longithorax Becker, 1956
 Agriotes lundbergi Platia, 1989
 Agriotes luteonotatus Pic, 1913
 Agriotes luteus Fleutiaux, 1940
 Agriotes maceki Platia & Gudenzi, 1997
 Agriotes magnanii Platia & Gudenzi, 1996
 Agriotes malaisei Fleutiaux, 1942
 Agriotes mancus (Say, 1823)
 Agriotes medvedevi Dolin, 1960
 Agriotes melanurus (Fleutiaux, 1925)
 Agriotes mertliki Platia, 2003
 Agriotes meticulosus Candèze, 1863
 Agriotes mexicanus Champion, 1896
 Agriotes miniatocollis Chevrolat, 1835
 Agriotes mixtus Champion, 1896
 Agriotes modestus Kiesenwetter, 1858
 Agriotes mollardi Fleutiaux, 1918
 Agriotes monticolus Champion, 1896
 Agriotes nadezhdae Cherepanov, 1965
 Agriotes nigricans Platia & Gudenzi, 1997
 Agriotes nigropubens Reitter, 1904
 Agriotes nigror Platia, 2003
 Agriotes notatus Candèze, 1863
 Agriotes novus Fleutiaux, 1939
 Agriotes nuceus Fairmaire, 1866
 Agriotes oblongicollis (Melsheimer, 1845)
 Agriotes ochraceipilosus (Fleutiaux, 1942)
 Agriotes olivieri Desbrochers des Loges, 1875
 Agriotes opacicollis Champion, 1896
 Agriotes opaculus (LeConte, 1859)
 Agriotes oreas Gistel, 1857
 Agriotes oregonensis Becker, 1956
 Agriotes orientalis Fleutiaux, 1939
 Agriotes oxianus Iablokoff-Khnzorian, 1970
 Agriotes pallidulus (Illiger, 1807)
 Agriotes paludum Kiesenwetter, 1859
 Agriotes pamirensis Gurjeva, 1978
 Agriotes passosi Giuseppe & Serrano, 2002
 Agriotes pauxillus Champion, 1896
 Agriotes pavesii Platia & Gudenzi, 2001
 Agriotes pectoralis Champion, 1896
 Agriotes petterssoni Platia & Gudenzi, 2000
 Agriotes pexus Candèze, 1863
 Agriotes pilosellus (Schönherr, 1817)
 Agriotes propinquus Fleutiaux, 1939
 Agriotes propleuralis Platia & Gudenzi, 1998
 Agriotes proximoides Platia, Furlan & Gudenzi, 2002
 Agriotes proximus Schwarz, 1891
 Agriotes pubescens Melsheimer, 1845
 Agriotes pulcherrimus Candèze, 1863
 Agriotes quadraticollis Champion, 1896
 Agriotes quadrilineatus Champion, 1896
 Agriotes quadrivittatus Candèze, 1863
 Agriotes quebecensis Brown, 1933
 Agriotes radula Desbrochers des Loges, 1875
 Agriotes reitteri Schwarz, 1891
 Agriotes riesei Platia, 2004
 Agriotes rojasi (Candèze, 1857)
 Agriotes rotundicollis Becker, 1956
 Agriotes rufipalpis Brullé, 1832
 Agriotes rufohumeralis Knull, 1959
 Agriotes rufus (Fleutiaux, 1918)
 Agriotes rugatus Fleutiaux, 1939
 Agriotes rugipennis Schwarz, 1891
 Agriotes sagittus Becker, 1956
 Agriotes samai Platia, Furlan & Gudenzi, 2002
 Agriotes sameki Platia, 2003
 Agriotes sareptanus Desbrochers des Loges, 1869
 Agriotes scapularis Candèze, 1863
 Agriotes scapularis Champion, 1896
 Agriotes schuberti Platia, 2004
 Agriotes schurmanni Platia & Gudenzi, 1998
 Agriotes sericatus Schwarz, 1891
 Agriotes sexualis Platia, 2004
 Agriotes shirozui Kishii, 1991
 Agriotes shykshanus Kishii, 1989
 Agriotes siciliensis Pic, 1912
 Agriotes sikorae Schwarz
 Agriotes simplex (Candèze, 1891)
 Agriotes singularius Fleutiaux, 1940
 Agriotes solai Platia, 2004
 Agriotes solitarius Fleutiaux, 1939
 Agriotes sordidus (Illiger, 1807)
 Agriotes sparsus LeConte, 1884
 Agriotes squalidus Schwarz, 1891
 Agriotes stabilis (LeConte, 1853)
 Agriotes starcki Schwarz, 1891
 Agriotes stepanovorum Orlov, 1997
 Agriotes strigosus Kiesenwetter, 1858
 Agriotes sublineatus Champion, 1896
 Agriotes subsulcatus Pic, 1913
 Agriotes subvittatus Motschulsky, 1860
 Agriotes succiniferus Becker, 1963
 Agriotes sylviae Cate & Platia, 1997
 Agriotes syriacus Platia & Gudenzi, 1997
 Agriotes tadzhikistanicus Gurjeva, 1967
 Agriotes taeniatus Candèze, 1863
 Agriotes takasago Kishii, 1989
 Agriotes tamdaoensis Fleutiaux, 1939
 Agriotes tardus Brown, 1933
 Agriotes tauricus Heyden, 1882
 Agriotes thevenetii Horn, 1872
 Agriotes tomentosus Fleutiaux, 1939
 Agriotes tonkinensis (Fleutiaux, 1894)
 Agriotes torquatus LeConte, 1884
 Agriotes tranninhus Fleutiaux, 1939
 Agriotes transcaucasicus Dolin, 2003
 Agriotes tres Fleutiaux, 1939
 Agriotes trilineatus Champion, 1896
 Agriotes tristiculus Reitter, 1906
 Agriotes trivittatus Champion, 1896
 Agriotes turcicus Candèze, 1863
 Agriotes turkmenicus Platia & Gudenzi, 1999
 Agriotes unicolor Koenig, 1889
 Agriotes ustulatus (Schaller, 1783)
 Agriotes vaccinus Candèze, 1863
 Agriotes vandykei Becker, 1956
 Agriotes vastus Gurjeva, 1972
 Agriotes vicinus Fleutiaux, 1907
 Agriotes villosus (Fleutiaux, 1918)
 Agriotes virgatus Candèze, 1863
 Agriotes werneri Platia, 2003
 Agriotes witzgalli Platia & Schimmel, 1993
 Agriotes yushihanus Kishii, 1989
 Agriotes zinovjevi Gurjeva, 1967

Kilder 
 Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (Liste over Nordens biller). Helsinki. 
 Zahradník, J. og Chvála, M. 1991. Teknologisk forlags store bok om insekter. N.W. Damm. Om smellere, side 282-287.

Eksterne lenker 
 
 

Smellere
Biller formelt beskrevet i 1829
Dyr formelt beskrevet av Johann Friedrich von Eschscholtz