Drusilla (bille)

Drusilla er en slekt av biller som hører til den store underfamilien småkortvinger (Aleocharinae) i familien kortvinger (Staphylinidae). De er spesialiserte på å leve sammen med maur.

Utseende  
Små til middelsstore (gjerne 3-5 millimeter), brunlige eller rødlige kortvinger, kroppen lite hårete bortsett fra hårpenslene på bakkroppen. Antennene er forholdsvis lange og kraftige. Hodet er forholdsvis lite og rundt. Pronotum er lengre enn bredt, gjerne med en bred midtfure. Dekkvingene er forholdsvis korte. Beina er forholdsvis lange og tynne.

Levevis 
Drusilla-artene har spesialisert seg på å leve sammen med maur. Disse artene lever rundt maurtuene  der de dels jakter på maurene, dels eter maurenes matrester.

Systematisk inndeling / norske arter 

 Ordenen biller, Coleoptera
 Underordenen Polyphaga
 Overfamilien åtsel- og rovbiller, Staphylinoidea
 Familien kortvinger (Staphylinidae) Latreille, 1802
 Underfamilien småkortvinger (Aleocharinae) Fleming, 1821
 Stammen Lomechusini 
 Slekten Drusilla Leach, 1819 (synonym Astilbus Dillwyn, 1829)
 Drusilla adjacens (Cameron, 1939)
 Drusilla aeneicollis (Fauvel, 1899)
 Drusilla aequatorialis (Bernhauer, 1927)
 Drusilla aerea (Scheerpeltz, 1934)
 Drusilla affinis (Cameron, 1939)
 Drusilla aino Nakane, 1963
 Drusilla alutacea Reitter, 1901
 Drusilla anceps Assing, 2005
 Drusilla andrewesi (Cameron, 1939)
 Drusilla angularis (Fauvel, 1899)
 Drusilla ashei Eldredge, 2012
 Drusilla assamensis (Cameron, 1939)
 Drusilla attenuata Kistner, 1997
 Drusilla baliensis Pace, 1987
 Drusilla bartolozzii Pace, 2003
 Drusilla bernhaueri (Scheerpeltz, 1934)
 Drusilla besucheti Assing, 2005
 Drusilla bicarinicollis Pace, 2008
 Drusilla bicolor (Cameron, 1950)
 Drusilla bifida Assing, 2016
 Drusilla bilobata Pace, 2014
 Drusilla bispinosa Assing, 2019
 Drusilla bituberculata (Motschulsky, 1858)
 Drusilla borneensis (Bernhauer, 1915)
 Drusilla borneoapicalis Pace, 2014
 Drusilla borneoclara Pace, 2014
 Drusilla borneoruficollis Pace, 2014
 Drusilla borneostricta Pace, 2014
 Drusilla brevipennis (Cameron, 1939)
 Drusilla breviuter Assing, 2019
 Drusilla bruneiensis Pace, 2008
 Drusilla bruneiorum Pace, 2008
 Drusilla bryanti (Cameron, 1950)
 Drusilla bulbata Assing, 2008
 Drusilla bulbosa Pace, 2014
 Drusilla burgeoniana (Bernhauer, 1943)
 Drusilla butuanensis (Bernhauer, 1916)
 Drusilla caelata Pace, 1987
 Drusilla caelaticollis Pace, 2008
 Drusilla calicis Pace, 2012
 Drusilla cameroni (Scheerpeltz, 1934)
 Drusilla canaliculata (Fabricius, 1787) - eneste art i Norge
 Drusilla caputserpentis Pace, 2014
 Drusilla carinithorax (Bernhauer, 1915)
 Drusilla celebensis (Cameron, 1942)
 Drusilla cenwanglaomontis Pace, 2012
 Drusilla cernens Assing, 2005
 Drusilla congoensis (Kistner, 2003)
 Drusilla convexifrons (Fairmaire, 1873)
 Drusilla cretica Assing, 2005
 Drusilla cupulifer (Fauvel, 1906)
 Drusilla damingensis Pace, 2012
 Drusilla danumensis Pace, 2014
 Drusilla dendrolasii Maruyama & Kishimoto, 2002
 Drusilla denigrata Assing, 2005
 Drusilla denticollis (Bernhauer, 1933)
 Drusilla dharmapuriensis Pace, 2001
 Drusilla dimidiata Pace, 1987
 Drusilla divergens Pace, 2014
 Drusilla dorylophila (Bernhauer, 1929)
 Drusilla drescheri (Cameron, 1939)
 Drusilla dumogaicola Pace, 2004
 Drusilla dusunorum Pace, 1993
 Drusilla endopilosella Pace, 2000
 Drusilla endorica (Saulcy, 1865)
 Drusilla erichsoni (Peyron, 1857)
 Drusilla feae (Cameron, 1939)
 Drusilla fissata (Bernhauer, 1915)
 Drusilla flagellata Assing, 2015
 Drusilla flava (Cameron, 1939)
 Drusilla flavicornis (Motschulsky, 1858)
 Drusilla foeda Pace, 2008
 Drusilla fontis Pace, 2008
 Drusilla foreli (Wasmann, 1890)
 Drusilla fossulicollis Pace, 2008
 Drusilla fossulifera Pace, 1987
 Drusilla frater (Cameron, 1950)
 Drusilla gabonensis Pace, 2012
 Drusilla ganglbaueri (Bernhauer, 1906)
 Drusilla gibberella Pace, 1998
 Drusilla gilva Pace, 2012
 Drusilla gracilicornis (Bernhauer, 1902)
 Drusilla gracilis (Hochhuth, 1849)
 Drusilla granulata (Bernhauer, 1927)
 Drusilla granulicollis (Cameron, 1939)
 Drusilla guineensis (Bernhauer, 1932)
 Drusilla haematica (Fauvel, 1899)
 Drusilla heydeni (Eppelsheim, 1887)
 Drusilla hipponensis (Fauvel, 1898)
 Drusilla impressicollis (Kraatz, 1857)
 Drusilla imurai Maruyama & Kishimoto, 2002
 Drusilla incola (Fauvel, 1904)
 Drusilla inflatae Maruyama, Yek, Hashim & Ito, 2003
 Drusilla intermedia (Cameron, 1933)
 Drusilla italica (Bernhauer, 1903)
 Drusilla jacobsoni (Cameron, 1936)
 Drusilla javana (Cameron, 1939)
 Drusilla kabyliana (Fagel, 1958)
 Drusilla kadooriorum Pace, 1998
 Drusilla kamerunensis (Bernhauer, 1912)
 Drusilla kathmanduensis Pace, 1992
 Drusilla katiki Pace, 1993
 Drusilla keiichii Maruyama, 2004
 Drusilla khamhengi Pace, 1986
 Drusilla kinabaluensis Pace, 1987
 Drusilla kisumuensis Pace, 1996
 Drusilla laevicauda (Bernhauer, 1903)
 Drusilla lateremaculata Pace, 2016
 Drusilla lativentris Assing, 2017
 Drusilla lethierryi (Fauvel, 1898)
 Drusilla liangicola Pace, 1993
 Drusilla limata Assing, 2005
 Drusilla lineatocollis (Kraatz, 1859)
 Drusilla longatheca Pace, 2004
 Drusilla lucens (Cameron, 1950)
 Drusilla lucida (Cameron, 1950)
 Drusilla luzonica (Bernhauer, 1915)
 Drusilla lydica Assing, 2007
 Drusilla magniceps (Cameron, 1939)
 Drusilla mahnerti Pace, 1996
 Drusilla malaisei (Scheerpeltz, 1965)
 Drusilla malindensis Pace, 1991
 Drusilla manenekarum Pace, 1993
 Drusilla manubrifera Pace, 2010
 Drusilla memnonia (Märkel, 1844)
 Drusilla meridiana (Fauvel, 1900)
 Drusilla mixta (Cameron, 1939)
 Drusilla montanella (Bernhauer, 1936)
 Drusilla monticola (Cameron, 1943)
 Drusilla nearctica Eldredge, 2012
 Drusilla necaerea Pace, 2008
 Drusilla nepalensis Pace, 1992
 Drusilla nigra (Cameron, 1925)
 Drusilla nigrescens (Motschulsky, 1858)
 Drusilla nomurai Maruyama, 2000
 Drusilla obliqua (Bernhauer, 1916)
 Drusilla obliquoides Pace, 2003
 Drusilla occipitalis Pace, 1987
 Drusilla ocularis (Fenyes, 1914)
 Drusilla opacicollis (Cameron, 1920)
 Drusilla opacipennis (Bernhauer, 1902)
 Drusilla operosa Pace, 1987
 Drusilla orientalis (Cameron, 1939)
 Drusilla orissaensis Pace, 2011
 Drusilla pallidicornis Assing, 2005
 Drusilla papuana (Cameron, 1937)
 Drusilla pendleburyi (Cameron, 1950)
 Drusilla penicillata  Assing, 2019
 Drusilla perakensis Pace, 2013
 Drusilla perdensa Pace, 2004
 Drusilla perforans Assing, 2015
 Drusilla persica Assing, 2005
 Drusilla persulcata Pace, 2004
 Drusilla pervisa Pace, 1987
 Drusilla philippina (Bernhauer, 1915)
 Drusilla planaticollis (Kraatz, 1857)
 Drusilla plena Pace, 2004
 Drusilla plicipennis (Bernhauer, 1915)
 Drusilla profunda Pace, 2014
 Drusilla proxima (Cameron, 1939)
 Drusilla pudukkottaiensis Pace, 2001
 Drusilla puetzi Assing, 2011
 Drusilla punctatissima (Kraatz, 1857)
 Drusilla raffrayi (Fairmaire, 1871)
 Drusilla rantepaoicola Pace, 1993
 Drusilla recta Assing, 2005
 Drusilla refugita Pace, 2000
 Drusilla rougemonti Pace, 1987
 Drusilla rougemontiana Pace, 2014
 Drusilla ruficornis (Cameron, 1939)
 Drusilla rufiventris (Cameron, 1950)
 Drusilla rufobrunnea (Cameron, 1939)
 Drusilla rufocaelata Pace, 2008
 Drusilla rufoflava (Cameron, 1950)
 Drusilla sabahensis Pace, 2014
 Drusilla sabahorum Pace, 2014
 Drusilla schawalleri Kistner, 1994
 Drusilla sculpticollides Hlavàč, Newton & Maruyama, 2011
 Drusilla semicircularis (Bernhauer, 1929)
 Drusilla semicyanea (Cameron, 1930)
 Drusilla semimonticola Pace, 2008
 Drusilla semiobliqua Pace, 2005
 Drusilla semipunctata (Bernhauer, 1929)
 Drusilla semirubra (Cameron, 1950)
 Drusilla serrulae Pace, 2014
 Drusilla shavrini  Assing, 2019
 Drusilla singalorum Pace, 1988
 Drusilla sinicola Pace, 2012
 Drusilla sinuosa Assing, 2005
 Drusilla smetanai Pace, 1992
 Drusilla sparsa (Sharp, 1874)
 Drusilla spiniventris  Assing, 2019
 Drusilla spissatheca Pace, 2008
 Drusilla stevensi (Cameron, 1939)
 Drusilla strigicollis (Cameron, 1928)
 Drusilla subsplendens Assing, 2011
 Drusilla suturalis (Erichson, 1843)
 Drusilla takashii Maruyama, 2004
 Drusilla taygetana Assing, 2005
 Drusilla terrestris Pace, 2008
 Drusilla thai Pace, 2012
 Drusilla thaifuscicollis Pace, 2012
 Drusilla theryi (Fauvel, 1898)
 Drusilla tigrata Pace, 1992
 Drusilla todolarum Pace, 1993
 Drusilla trigonocephala (Cameron, 1939)
 Drusilla trina Pace, 2014
 Drusilla tronqueti Pace, 1988
 Drusilla tuberculata (Bernhauer, 1916)
 Drusilla ulusegamicola Pace, 2015
 Drusilla umranicola Pace, 2005
 Drusilla umtaliensis Pace, 1996
 Drusilla unicolor (Cameron, 1950)
 Drusilla veluticollis (Cameron, 1928)
 Drusilla villosa (Peyerimhoff, 1909)
 Drusilla wallacei (Kistner & McNairn, 1991)
 Drusilla watanabei Maruyama & Kishimoto, 2002
 Drusilla yunnanensis Pace, 1993
 Drusilla zhejiangensis Pace, 1998
 Drusilla zimbabwensis Pace, 1996
 Drusilla zyrasoides Dvořàk, 1988

Referanser

 Silfverberg, H. 1992. Enumeratio Coleopterorum Fennoscandiae, Daniae et Baltiae (Liste over Nordens biller). Helsinki.

Eksterne lenker 
 

Småkortvinger
Biller formelt beskrevet i 1819
Dyr formelt beskrevet av William Elford Leach