Zygadenia

Zygadenia er en slekt av utdødde biller som trolig levde i døde eller dødende trær eller deres røtter. Slekten levde i Mesozoicum. Det blir nå plassert ca. 70 fossile arter i slekten, noe som gjør den til ens vært artrsik fossil-slekt.

Utseende 
Ommatidae er generelt mørke, slanke, flate biller, dekkvingene har opphøyde ribber og nettmønster. Kjevene (mandiblene) er fremoverrettede og ganske store. Antennene er mer eller mindre sylindriske eller perlekjede-aktige.

Historie 
Ommatidae er en gammel gruppe som åpenbart må ha vært mye mer artsrik i tidligere tidsperioder. Arten plassert i denne slekten er kjent fra mye av Jordens mellomalder, fra trias, jura og kritt, mellom 232-94 millioner år før nåtid, fra det næværende England, Tyskland, Spania, Russland, Kasakhstan, Mongolia, Kina, Australia og Argentina.

Systematisk inndeling 
Disse artene har tidligere vært plassert i en rekke ulike slekter, se liste i taksoboksen.

 Ordenen biller, Coleoptera
 Underordenen Archostemata
 Familien Ommatidae Sharp & Muir, 1912
 Gruppen Notocupedini Ponomarenko, 1966
 Slekten Zygadenia Handlirsch, 1906 
 †Zygadenia aliena (Tan and Ren 2006) Yixian Formation, China, Aptian
 †Zygadenia angliae (Giebel 1856) Purbeck Group, United Kingdom, Berriasian, Weald Clay, United Kingdom, Barremian
 †Zygadenia brachycephalus (Ponomarenko, 1994) Bakhar Formation, Mongolia, Bajocian, Shar Teeg, Mongolia, Tithonian
 †Zygadenia caducus (Ponomarenko, 1969) Kzyl-Zhar, Kazakhstan, Turonian
 †Zygadenia caudatus (Ponomarenko, 1966) Zaza Formation, Russia, Aptian
 †Zygadenia cellulosus (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
 †Zygadenia crassus (Ponomarenko, 1969) Sulyukta Formation, Kyrgyzstan, Toarcian
 †Zygadenia diazromerali Soriano & Delclòs, 2006 Las Hoyas, Spain, Barremian
 †Zygadenia dischides (Zhang 1986) Jiulongshan Formation, China, Callovian
 †Zygadenia dundulaensis (Ponomarenko, 1994) Dzun-Bain Formation, Mongolia, Aptian
 †Zygadenia elegans (Ponomarenko, 1994) Dzun-Bain Formation, Mongolia, Aptian
 †Zygadenia epicharis (Tan et al. 2005) Yixian Formation, China, Aptian
 †Zygadenia eumeura (Tan et al. 2006) Yixian Formation, China, Aptian
 †Zygadenia excellens (Ponomarenko, 1966)  Zaza Formation, Russia, Aptian
 †Zygadenia exiguus (Ponomarenko, 1994) Bakhar Formation, Mongolia, Bajocian, Shar Teeg, Mongolia, Tithonian
 †Zygadenia floodpagei Jarzembowski et al. 2015 Weald Clay, United Kingdom, Hauterivian-Barremian
 †Zygadenia foersteri (Ponomarenko, 1971) Karabastau Formation, Kazakhstan, Oxfordian
 †Zygadenia giebeli Ponomarenko 2014 Shar Teeg, Mongolia, Tithonian
 †Zygadenia handlirschi Ponomarenko 2014 Shar Teeg, Mongolia, Tithonian
 †Zygadenia homorus (Lin, 1986) Guanyintan Formation, China, Hettangian
 †Zygadenia issykkulensis (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
 †Zygadenia jurassica Hong 1983 Jiulongshan Formation, China, Callovian
 †Zygadenia kezuoensis Hong 1987 Shahai Formation, China, Aptian
 †Zygadenia khasurtyiensis (Strelnikova 2019) Khasurty, Russia, Aptian
 †Zygadenia khetanensis (Ponomarenko in Cromov et al., 1993) Emanra Formation, Russia, Turonian
 †Zygadenia kirghizicus (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
 †Zygadenia laetus (Lin, 1976)  Jianshangou Formation, China, Barremian
 †Zygadenia laiyangensis (Hong, 1990) Laiyang Formation, China, Aptian
 †Zygadenia lapidarius (Ponomarenko, 1968) Karabastau Formation, Kazakhstan, Oxfordian
 †Zygadenia laticella (Ponomarenko, 1969) Madygen Formation, Kyrgyzstan, Carnian
 †Zygadenia latus (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
 †Zygadenia lentus (Ren, 1995) Dabeigou Formation, China, Hauterivian
 †Zygadenia lini (Ponomarenko et al. 2012) Guanyintan Formation, China Toarcian/Aalenian
 †Zygadenia liui Jarzembowski et al. 2015 Yixian Formation, China, Aptian
 †Zygadenia longicollis (Ponomarenko, 1994) Bakhar Formation, Mongolia, Bajocian
 †Zygadenia longicoxa Soriano & Delclòs, 2006  Las Hoyas, Spain, Barremian
 †Zygadenia ludongensis (Wang and Liu 1996) Laiyang Formation, China, Aptian
 †Zygadenia martinae Jarzembowski et al. 2015 Cattamarra Coal Measures, Australia, Toarcian - bør kanskje plasseres i en egen slekt
 †Zygadenia martinclosas Soriano & Delclòs, 2006 La Pedrera de Rúbies Formation, Las Hoyas, Spain, Barremian
 †Zygadenia miniscula Tan et al. 2006, Yixian Formation, China, Aptian
 †Zygadenia mongolicus (Ponomarenko, 1994) Ulan-Argalant Formation, Mongolia, Barremian
 †Zygadenia nigromonticola (Ponomarenko, 1968) Karabastau Formation, Kazakhstan, Oxfordian
 †Zygadenia oculata Soriano & Delclòs, 2006 La Pedrera de Rúbies Formation, Spain, Barremian
 †Zygadenia oxypygus (Ponomarenko, 1969) Koldzat Formation, Kazakhstan, Carnian
 †Zygadenia patulus Ponomarenko, 1985 Tyumen Formation, Russian, Callovian
 †Zygadenia picturatus (Ponomarenko, 1964) Karabastau Formation
 †Zygadenia pingi (Ponomarenko and Ren 2010) Daohugou, China, Callovian
 †Zygadenia porrecta Tan et al. 2006 Yixian Formation, China, Aptian
 †Zygadenia protensa Tan et al. 2006 Yixian Formation, China, Aptian
 †Zygadenia psilata Tan et al. 2005 Yixian Formation, China, Aptian
 †Zygadenia pulcher (Ponomarenko, 1968) Karabastau Formation, Kazakhstan, Oxfordian
 †Zygadenia reticulatus (Oppenheim, 1888) Solnhofen, Germany, Tithonian
 †Zygadenia rostratus (Ponomarenko, 1969) Madygen Formation, Kyrgyzstan, Carnian
 †Zygadenia rudis Tan et al. 2005 Yixian Formation, China, Aptian
 †Zygadenia semen (Ponomarenko, 2000) Argun Formation, Russia, Aptian
 †Zygadenia shiluoensis Hong 1984 Tongchuan Formation, China, Ladinian
 †Zygadenia sibirica Ponomarenko, 2000 Argun Formation, Russia, Aptian
 †Zygadenia simpsoni Jarzembowski et al. 2015 Weald Clay, United Kingdom, Hauterivian-Barremian
 †Zygadenia siniestri Soriano & Delclòs, 2006 Las Hoyas, Spain, Barremian
 †Zygadenia sinitzae Ponomarenko, 2000 Argun Formation, Russia, Aptian
 †Zygadenia sogutensis (Ponomarenko, 1969) Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
 †Zygadenia stabilis Tan et al. 2005 Yixian Formation, China, Aptian
 †Zygadenia tenuis (Ponomarenko, 1969) Madygen Formation, Kyrgyzstan, Carnian
 †Zygadenia tripartitus (Oppenheim, 1888) Solnhofen, Germany, Tithonian
 †Zygadenia tuanwangensis (Hong, 1990) Laiyang Formation, China, Aptian
 †Zygadenia tuberculatus (Giebel, 1856) Purbeck Group, United Kingdom, Berriasian
 †Zygadenia valida Lin 1976 Jehol Group, China, Aptian
 †Zygadenia viridis Soriano & Delclòs, 2006 Las Hoyas, Spain, Barremian
 †Zygadenia vitimensis (Ponomarenko, 1966) Zaza Formation, Russia, Aptian
 †Zygadenia westraliensis Riek 1968 Cattamarra Coal Measures, Australia, Toarcian - bør kanskje plasseres i en egen slekt

Referanser

Eksterne lenker 
 

Biller formelt beskrevet i 1826
Archostemata
Fossile insekter