Kumpai (tumbuhan)

Kumpai (dari bahasa Sunda) adalah tumbuhan paku dari divisio Lycopodiophyta yang kebanyakan tumbuh secara epifit. Kumpai adalah anggota suku Lycopodiaceae, sebagai anaksuku Huperzioideae (menurut PPG I (2016)) atau marga Huperzia arti luas (sensu lato). Dalam sumber-sumber lama pada era pra-molekular, kumpai merupakan bagian dari marga Lycopodium. Penjelasan lebih lanjut diberikan di bagian Taksonomi.

Beberapa anggotanya adalah tanaman hias gantung populer, seperti kumpai lubang (kumpai tali atau kumpai ekor tikus, Huperzia carinata), kumpai picisan (Huperzia nummularifolia), kumpai ekor monyet (Huperzia squarrosa) dan kumpai rantai (Huperzia phlegmaria).

Morfologi 
Sporofit dari genus ini memiliki pucuk yang tidak bercabang yang umumnya tegak dan berbentuk bulat pada penampang melintang. Batang horizontal tidak ada. Daunnya tidak memiliki barisan yang berbeda, dan biasanya berbentuk agak lanset. Pada beberapa spesies, ukurannya bervariasi sesuai dengan musim di mana mereka tumbuh. Cabang-cabang yang mengandung gemmae – struktur seperti kuncup di mana tanaman bereproduksi secara aseksual – terjadi di antara daun. Gemmae berbentuk segitiga, dengan delapan daun dalam pola konstan: empat diratakan menjadi bidang dan dua daun lateral yang besar. Sporangia berbentuk ginjal (reniform), terdapat di pangkal daun yang tidak termodifikasi atau tereduksi. Akar diproduksi di dekat puncak tunas, dan bermigrasi ke bawah di dalam korteks batang untuk muncul di permukaan tanah. Gametofit yang tidak bercabang tidak fotosintesis, melainkan di bawah tanah dan mikoriza .

Flora Amerika Utara membedakan Huperzia dari genus Phlegmariurus tropis epifit berdasarkan perbedaan seperti tunas kompleks dan khusus, permata dan cabang kecil tempat mereka dilahirkan, dan gametofit tidak bercabang.

Taksonomi 
Genus Huperzia diperkenalkan oleh Johann Jakob Bernhardi pada tahun 1801. Bernhardi memisahkan Huperzia dari Lycopodium . Spesies tipenya adalah Lycopodium selago yang menjadi Huperzia selago .Pada klasifikasi menurut Pteridophyte Phylogeny Group (PPG I) (2016), Huperzia ditempatkan ke dalam anaksuku Huperzioideae dari suku Lycopodiaceae. Studi filogenetik yang dilakukan Field et al. (2016), yang didasarkan pada data molekular dan morfologi, mencapai kesimpulan bahwa pilihan satu genus (Huperzia s.l.) maupun pembagian dalam tiga genus (Huperzia s.s., Phlegmariurus, dan Phylloglossum) dapat diterima.

Banyak dari jenis yang sebelumnya ditempatkan dalam Huperzia arti luas dimasukkan dalam Phlegmariurus. Sebelumnya, Flora of North America juga telah memisahkan Huperzia dari Phegmariurus.  Namun, Phlegmariurus sebenarnya secara morfologi sulit untuk dibedakan dari Huperzia sehingga ada penulis yang lebih menyukai pengelompokan dalam satu marga Huperzia.

Jenis 
Klasifikasi PPG I menyebutkan ada 25 spesies dalam genus Huperzia .  , Ferns of The World mendaftarkan 56 jenis, dengan catatan bahwa "banyak spesies masih perlu dipindahkan ke genera terpisah lainnya".

Huperzia appalachiana Beitel & Mickel
Huperzia archboldiana (Nessel) Holub
Huperzia arunachalensis (D.D.Pant & P.S.Pandey) Fraser-Jenk.
Huperzia asiatica (Ching) N.Shrestha & X.C.Zhang
Huperzia australiana (Herter) Holub
Huperzia beccarii (Alderw.) Holub
Huperzia bucahwangensis Ching
Huperzia campestris (Alderw.) Holub
Huperzia chinensis (Christ) Ching
Huperzia chishuiensis X.Y.Wang & P.S.Wang
Huperzia continentalis Testo, A.Haines & A.V.Gilman
Huperzia crispata (Ching) Ching
Huperzia emeiensis (Ching & H.S.Kung) Ching & H.S.Kung
Huperzia erubescens (Brack.) Holub
Huperzia everettii (Herter) Holub
Huperzia fuegiana (Roiv.) Holub
Huperzia gedeana (Alderw.) Holub
Huperzia goliathensis (Alderw.) Holub
Huperzia haleakalae (Brack.) Holub
Huperzia herteriana (Kümmerle) T.Sen & U.Sen
Huperzia javanica (Sw.) Fraser-Jenk.
Huperzia jejuensis B.Y.Sun & J.Lim
Huperzia kangdingensis (Ching) Ching
Huperzia kunmingensis Ching
Huperzia laipoensis Ching
Huperzia lajouensis Ching
Huperzia leishanensis X.Y.Wang
Huperzia liangshanica (H.S.Kung) Ching & H.S.Kung
Huperzia lucidula (Michx.) Trevis.
Huperzia medogensis Ching & Y.X.Lin
Huperzia meghalaica Fraser-Jenk.
Huperzia miniata (Spring) Trevis.
Huperzia minima (Herter) Holub
Huperzia miyoshiana (Makino) Ching
Huperzia muscicola W.M.Chu
Huperzia nanchuanensis (Ching & H.S.Kung) Ching & H.S.Kung
Huperzia nanlingensis Shrestha, F.W.Xing, X.P.Qi, Y.H.Yan & X.C.Zhang
Huperzia occidentalis (Clute) Kartesz & Gandhi
Huperzia petrovii Sipliv.
Huperzia porophila (F.E.Lloyd & Underw.) Holub
Huperzia quasipolytrichoides (Hayata) Ching
Huperzia rubicaulis S.K.Wu & X.Cheng
Huperzia rubricaulis (Alderw.) Holub
Huperzia saururoides (Bory & D'Urv.) Rothm.
Huperzia selago (L.) Bernh. (including H. selago subsp. appressa - Appalachian firmoss)
Huperzia serrata (Thunb.) Trevis.
Huperzia shresthae Fraser-Jenk.
Huperzia somae (Hayata) Ching
Huperzia sprengeri (Nessel) Holub
Huperzia suberecta (Lowe) Tardieu
Huperzia sumatrana (Alderw.) Holub
Huperzia sutchueniana (Herter) Ching
Huperzia tibetica (Ching) Ching
Huperzia yakusimensis (Herter) Holub
Huperzia zollingeri (Herter) Holub

Jenis yang dialihkan karena perbedaan sistem klasifikasi
Dalam versi satu marga, Phylloglossum drummondii menjadi sinonim bagi Huperzia drummondii. Sementara, hingga Oktober 2019, jenis-jenis yang dialihkan dari Huperzia ke Phlegmariurus menurut Checklist of Ferns dan Lycophytes of the World (versi tiga marga) adalah sebagai berikut.

Huperzia acerosa (Sw.) B.Øllg. = Phlegmariurus acerosus (Sw.) B.Øllg.
Huperzia acifolia (Rolleri) B.Øllg. = Phlegmariurus acifolius (Rolleri) B.Øllg.
Huperzia acuta (Rolleri) B.Øllg. = Phlegmariurus acutus (Rolleri) B.Øllg.
Huperzia affinis (Trevis.) B.Øllg. = Phlegmariurus affinis (Trevis.) B.Øllg.
Huperzia afromontana Pic. Serm. = Phlegmariurus afromontanus (Pic. Serm.) A.R.Field & Bostock comb. nov.
 Huperzia aloifolia (Hook. & Grev.) Trevis. = Phlegmariurus aloifolius (Hook. & Grev.) A.R.Field & Bostock comb. nov.
Huperzia ambrensis Rakotondr. = Phlegmariurus ambrensis (Rakotondr.) A.R.Field & Bauret
Huperzia amentacea B.Øllg. = Phlegmariurus amentaceus (B.Øllg.) B.Øllg.
Huperzia andina Rosenst. = Phlegmariurus andinus (Rosenst.) B.Øllg.
Huperzia apolinari-mariae Nessel = Phlegmariurus apolinari-mariae (Nessel) comb. ined.
Huperzia aqualupiana Spring = Phlegmariurus aqualupianus (Spring) B.Øllg.
Huperzia arcturi Herter = Phlegmariurus arcturi (Herter) B.Øllg.
Huperzia arcuata B.Øllg. = Phlegmariurus arcuatus (B.Øllg.) B.Øllg.
Huperzia aristei Nessel = Phlegmariurus aristei (Nessel) comb. ined.
Huperzia ascendens Nessel = Phlegmariurus ascendens (Nessel) B.Øllg.
Huperzia attenuata Spring = Phlegmariurus attenuatus (Spring) B.Øllg.
Huperzia australis Willd. = Phlegmariurus australis (Willd.) A.R.Field
Huperzia austroecuadorica B.Øllg. = Phlegmariurus austroecuadoricus (B.Øllg.) B.Øllg.
Huperzia austrosinica Ching) = Phlegmariurus austrosinicus (Ching) Li Bing Zhang
Huperzia axillaris Roxb. = Phlegmariurus axillaris (Roxb.) comb. ined.
Huperzia badiniana B.Øllg. & P.G.Windisch = Phlegmariurus badinianus (B.Øllg. & P.G.Windisch) B.Øllg.
Huperzia balansae (Herter) Holub = Phlegmariurus balansae (Herter) A.R.Field & Bostock
Huperzia bampsiana Pic.Serm. = Phlegmariurus bampsianus (Pic. Serm.) A.R.Field & Bostock
Huperzia banayanica (Herter) Holub = Phlegmariurus banayanicus (Herter) A.R.Field & Bostock
Huperzia beiteliana Mickel = Phlegmariurus beitelianus (Mickel) comb. ined.
Huperzia beitelii B.Øllg. = Phlegmariurus beitelii (B.Øllg.) B.Øllg.
Huperzia biformis Hook. = Phlegmariurus biformis (Hook.) B.Øllg.
Huperzia binervia (Herter)  = Phlegmariurus binervius (Herter) B.Øllg.
Huperzia bolanica (Rosenst.) Holub = Phlegmariurus bolanicus (Rosenst.) A.R.Field & Bostock
Huperzia brachiata Maxon = Phlegmariurus brachiatus (Maxon) B.Øllg.
Huperzia brachystachys (Baker) Pic.Serm. = Phlegmariurus brachystachys (Baker) A.R.Field & Bostock
Huperzia bradeorum Christ = Phlegmariurus bradeorum (Christ) B.Øllg.
Huperzia brassii (Copel.) Holub = Phlegmariurus brassii (Copel.) A.R.Field & Bostock
Huperzia brevifolia (Hook. & Grev.)  = Phlegmariurus brevifolius (Hook. & Grev.) B.Øllg.
Huperzia brongniartii (Spring)  = Phlegmariurus brongniartii (Spring) B.Øllg.
Huperzia buesii (Herter) . = Phlegmariurus buesii (Herter) B.Øllg.
Huperzia callitrichifolia (Mett.)  = Phlegmariurus callitrichifolius (Mett.) B.Øllg.
Huperzia campiana B.Øllg. = Phlegmariurus campianus (B.Øllg.) B.Øllg.
Huperzia cancellata (Spring) Ching = Phlegmariurus cancellatus (Spring) Ching
Huperzia capellae (Herter) = Phlegmariurus capellae (Herter) B.Øllg.
Huperzia capillaris (Sodiro) B.Øllg. = Phlegmariurus capillaris (Sodiro) B.Øllg.
Huperzia carinata (Desv. ex Poir.) Trevis. = Phlegmariurus carinatus (Desv. ex Poir.) Ching
Huperzia catacachiensis (Nessel) B.Øllg. = Phlegmariurus catacachiensis (Nessel) B.Øllg.
Huperzia catharinae (Christ) comb. ined. = Phlegmariurus catharinae (Christ) comb. ined.
Huperzia cavifolia (C.Chr.) A.R.Field & Bostock = Phlegmariurus cavifolius (C.Chr.) A.R.Field & Bostock
Huperzia ceylanica (Spring) comb. ined. = Phlegmariurus ceylanicus (Spring) comb. ined.
Huperzia chamaeleon (Herter) B.Øllg. = Phlegmariurus chamaeleon (Herter) B.Øllg.
Huperzia changii T.Y.Hsieh = Phlegmariurus changii T.Y.Hsieh
Huperzia chiricana (Maxon) B.Øllg. = Phlegmariurus chiricanus (Maxon) B.Øllg.
Huperzia christii (Silveira) B.Øllg. = Phlegmariurus christii (Silveira) B.Øllg.
Huperzia ciliatospiculata B.Øllg. = Phlegmariurus ciliatospiculatus B.Øllg.
Huperzia ciliolata (B.Øllg.) B.Øllg. = Phlegmariurus ciliolatus (B.Øllg.) B.Øllg.
Huperzia clarkiana (Nessel) comb. ined. = Phlegmariurus clarkianus (Nessel) comb. ined.
Huperzia cleefiana (B.Øllg.) B.Øllg. = Phlegmariurus cleefianus (B.Øllg.) B.Øllg.
Huperzia colanensis (B.Øllg.) B.Øllg. = Phlegmariurus colanensis (B.Øllg.) B.Øllg.
Huperzia columnaris (B.Øllg.) B.Øllg. = Phlegmariurus columnaris (B.Øllg.) B.Øllg.
Huperzia comans (Herter ex Nessel) comb. ined. = Phlegmariurus comans (Herter ex Nessel) comb. ined.
Huperzia compacta (Hook.) B.Øllg. = Phlegmariurus compactus (Hook.) B.Øllg.
Huperzia copelandiana (R.C.Y.Chou & Bartlett) A.R.Field = Phlegmariurus copelandianus (R.C.Y.Chou & Bartlett) A.R.Field
Huperzia coralium (Spring) A.R.Field & Bostock = Phlegmariurus coralium (Spring) A.R.Field & Bostock
Huperzia coriacea (Rolleri) comb. ined. = Phlegmariurus coriacea (Rolleri) comb. ined.
Huperzia costaricensis (Herter) comb. ined. = Phlegmariurus costaricensis (Herter) comb. ined.
Huperzia crassa (Humb. & Bonpl. ex Willd.) B.Øllg. = Phlegmariurus crassus (Humb. & Bonpl. ex Willd.) B.Øllg.
Huperzia creber (Alderw.) A.R.Field & Bostock = Phlegmariurus creber (Alderw.) A.R.Field & Bostock
Huperzia crucis-australis (Herter) B.Øllg. = Phlegmariurus crucis-australis (Herter) B.Øllg.
Huperzia cruenta (Spring) B.Øllg. = Phlegmariurus cruentus (Spring) B.Øllg.
Huperzia cryptomerina (Maxim.) Satou = Phlegmariurus cryptomerinus (Maxim.) Satou
Huperzia cuernavacensis (Underw. & F.E.Lloyd) B.Øllg. = Phlegmariurus cuernavacensis (Underw. & F.E.Lloyd) B.Øllg.
Huperzia cumingii (Nessel) B.Øllg. = Phlegmariurus cumingii (Nessel) B.Øllg.
Huperzia cuneifolia (Hieron.) B.Øllg. = Phlegmariurus cuneifolius (Hieron.) B.Øllg.
Huperzia cunninghamioides (Hayata) Ching = Phlegmariurus cunninghamioides (Hayata) Ching
Huperzia curiosa (Herter) A.R.Field & Bostock = Phlegmariurus curiosus (Herter) A.R.Field & Bostock
Huperzia curvifolia (Kunze) B.Øllg. = Phlegmariurus curvifolius (Kunze) B.Øllg.
Huperzia dacrydioides (Baker) A.R.Field & Bostock = Phlegmariurus dacrydioides (Baker) A.R.Field & Bostock
Huperzia dalhousieana (Spring) A.R.Field & Bostock = Phlegmariurus dalhousieanus (Spring) A.R.Field & Bostock
Huperzia darwiniana (Herter ex Nessel) B.Øllg. = Phlegmariurus darwinianus (Herter ex Nessel) B.Øllg.
Huperzia delavayi (Christ & Herter) comb. ined. = Phlegmariurus delavayi (Christ & Herter) comb. ined.
Huperzia delbrueckii (Herter) A.R.Field & Bostock = Phlegmariurus delbrueckii (Herter) A.R.Field & Bostock
Huperzia deminuens (Herter) B.Øllg. = Phlegmariurus deminuens (Herter) B.Øllg.
Huperzia dentata (Herter) Arana = Phlegmariurus dentatus (Herter) Arana
Huperzia dianae (Herter) B.Øllg. = Phlegmariurus dianae (Herter) B.Øllg.
Huperzia dichaeoides (Maxon) B.Øllg. = Phlegmariurus dichaeoides (Maxon) B.Øllg.
Huperzia dichotoma (Jacq.) W.H.Wagner = Phlegmariurus dichotomus (Jacq.) W.H.Wagner
Huperzia dielsii (Herter) A.R.Field & Bostock = Phlegmariurus dielsii (Herter) A.R.Field & Bostock
Huperzia divergens (Alderw.) A.R.Field & Testo = Phlegmariurus divergens (Alderw.) A.R.Field & Testo
Huperzia dura (Pic. Serm.) A.R.Field & Bostock = Phlegmariurus durus (Pic. Serm.) A.R.Field & Bostock
Huperzia echinata (Spring) B.Øllg. = Phlegmariurus echinatus (Spring) B.Øllg.
Huperzia ellenbeckii (Nessel) A.R.Field & Bostock = Phlegmariurus ellenbeckii (Nessel) A.R.Field & Bostock
Huperzia elmeri (Herter) A.R.Field & Bostock = Phlegmariurus elmeri (Herter) A.R.Field & Bostock
Huperzia engleri (Hieron. & Herter) B.Øllg. = Phlegmariurus engleri (Hieron. & Herter) B.Øllg.
Huperzia eremorum (Rolleri) B.Øllg. = Phlegmariurus eremorum (Rolleri) B.Øllg.
Huperzia ericifolia (C.Presl) B.Øllg. = Phlegmariurus ericifolius (C.Presl) B.Øllg.
Huperzia erosa (Beitel & W.H.Wagner) comb. ined. = Phlegmariurus erosus (Beitel & W.H.Wagner) comb. ined.
Huperzia erythrocaulon (Fée) comb. ined. = Phlegmariurus erythrocaulon (Fée) comb. ined.
Huperzia espinosana (B.Øllg.) B.Øllg. = Phlegmariurus espinosanus (B.Øllg.) B.Øllg.
Huperzia eversa (Poir.) B.Øllg. = Phlegmariurus eversus (Poir.) B.Øllg.
Huperzia fargesii (Herter) Ching = Phlegmariurus fargesii (Herter) Ching
Huperzia filicaulon (Copel.) A.R.Field & Testo = Phlegmariurus filicaulon (Copel.) A.R.Field & Testo
Huperzia filiformis (Sw.) W.H.Wagner = Phlegmariurus filiformis (Sw.) W.H.Wagner
Huperzia firma (Mett.) B.Øllg. = Phlegmariurus firmus (Mett.) B.Øllg.
Huperzia flagellacea (Kuhn) A.R.Field & Bostock = Phlegmariurus flagellaceus (Kuhn) A.R.Field & Bostock
Huperzia flexibilis (Fée) B.Øllg. = Phlegmariurus flexibilis (Fée) B.Øllg.
Huperzia foliacea (Maxon) B.Øllg. = Phlegmariurus foliaceus (Maxon) B.Øllg.
Huperzia foliosa (Copel.) A.R.Field & Bostock = Phlegmariurus foliosus (Copel.) A.R.Field & Bostock
Huperzia fontinaloides (Spring) B.Øllg. = Phlegmariurus fontinaloides (Spring) B.Øllg.
Huperzia fordii (Baker) Ching = Phlegmariurus fordii (Baker) Ching
Huperzia friburgensis (Herter ex Nessel & Hoehne) B.Øllg. = Phlegmariurus friburgensis (Herter ex Nessel & Hoehne) B.Øllg.
Huperzia funicula (Herter) comb. ined. = Phlegmariurus funiculus (Herter) comb. ined.
Huperzia funiformis (Cham. ex Spring) B.Øllg. = Phlegmariurus funiformis (Cham. ex Spring) B.Øllg.
Huperzia gagnepainiana (Herter) A.R.Field & Bostock = Phlegmariurus gagnepainianus (Herter) A.R.Field & Bostock
Huperzia galapagensis (O.J.Hamann) B.Øllg. = Phlegmariurus galapagensis (O.J.Hamann) B.Øllg.
Huperzia gigantea (Herter) A.R.Field & Bostock = Phlegmariurus giganteus (Herter) A.R.Field & Bostock
Huperzia gnidioides (L. fil.) A.R.Field & Bostock = Phlegmariurus gnidioides (L. fil.) A.R.Field & Bostock
Huperzia goebelii (Nessel) A.R.Field & Bostock = Phlegmariurus goebelii (Nessel) A.R.Field & Bostock
Huperzia goudotii (Herter) B.Øllg. = Phlegmariurus goudotii (Herter) B.Øllg.
Huperzia guandongensis Ching = Phlegmariurus guandongensis Ching
Huperzia gunturensis (Alderw.) A.R.Field & Bostock = Phlegmariurus gunturensis (Alderw.) A.R.Field & Bostock
Huperzia haeckelii (Herter) A.R.Field & Testo = Phlegmariurus haeckelii (Herter) A.R.Field & Testo
Huperzia haitensis (Herter) comb. ined. = Phlegmariurus haitensis (Herter) comb. ined.
Huperzia hamiltonii (Spreng.) Á.Löve & D.Löve = Phlegmariurus hamiltonii (Spreng.) Á.Löve & D.Löve
Huperzia harmsii (Nessel) A.R.Field & Bostock = Phlegmariurus harmsii (Nessel) A.R.Field & Bostock
Huperzia hartwegiana (Spring) B.Øllg. = Phlegmariurus hartwegianus (Spring) B.Øllg.
Huperzia hastata (B.Øllg.) B.Øllg. = Phlegmariurus hastatus (B.Øllg.) B.Øllg.
Huperzia hellwigii (Warb.) A.R.Field & Bostock = Phlegmariurus hellwigii (Warb.) A.R.Field & Bostock
Huperzia hemleri (Nessel) B.Øllg. = Phlegmariurus hemleri (Nessel) B.Øllg.
Huperzia henryi (Baker) Ching = Phlegmariurus henryi (Baker) Ching
Huperzia heterocarpa (Fée) B.Øllg. = Phlegmariurus heterocarpus (Fée) B.Øllg.
Huperzia heteroclita (Desv. ex Poir.) B.Øllg. = Phlegmariurus heteroclitus (Desv. ex Poir.) B.Øllg.
Huperzia hexasticha (B.Øllg. & P.G.Windisch) B.Øllg. = Phlegmariurus hexastichus (B.Øllg. & P.G.Windisch) B.Øllg.
Huperzia hildebrandtii (Herter) A.R.Field & Bauret = Phlegmariurus hildebrandtii (Herter) A.R.Field & Bauret
Huperzia hippuridea (Christ) B.Øllg. = Phlegmariurus hippurideus (Christ) B.Øllg.
Huperzia hippuris (Desv. ex Poir.) A.R.Field & Testo = Phlegmariurus hippuris (Desv. ex Poir.) A.R.Field & Testo
Huperzia hoffmannii (Maxon) B.Øllg. = Phlegmariurus hoffmannii (Maxon) B.Øllg.
Huperzia hohenackeri (Herter) B.Øllg. = Phlegmariurus hohenackeri (Herter) B.Øllg.
Huperzia holstii (Hieron.) A.R.Field & Bostock = Phlegmariurus holstii (Hieron.) A.R.Field & Bostock
Huperzia homocarpa (Herter) B.Øllg. = Phlegmariurus homocarpus (Herter) B.Øllg.
Huperzia horizontalis (Nessel) A.R.Field & Bostock = Phlegmariurus horizontalis (Nessel) A.R.Field & Bostock
Huperzia huberi (B.Øllg.) B.Øllg. = Phlegmariurus huberi (B.Øllg.) B.Øllg.
Huperzia hugonis B.Øllg. = Phlegmariurus hugonis B.Øllg.
Huperzia humbertii (Nessel) A.R.Field & Bostock = Phlegmariurus humbertii (Nessel) A.R.Field & Bostock
Huperzia humbertii-henrici (Herter) A.R.Field & Bostock = Phlegmariurus humbertii-henrici (Herter) A.R.Field & Bostock
Huperzia hypogaea (B.Øllg.) B.Øllg. = Phlegmariurus hypogaeus (B.Øllg.) B.Øllg.
Huperzia hystrix (Herter) B.Øllg. = Phlegmariurus hystrix (Herter) B.Øllg.
Huperzia ignambiensis (Compton) A.R.Field & Testo = Phlegmariurus ignambiensis (Compton) A.R.Field & Testo
Huperzia iminii Kiew = Phlegmariurus iminii Kiew
Huperzia innocentia (Herter) comb. ined. = Phlegmariurus innocentius (Herter) comb. ined.
Huperzia insularis (Carmich.) comb. ined. = Phlegmariurus insularis (Carmich.) comb. ined.
Huperzia intermedia (Trevis.) B.Øllg. = Phlegmariurus intermedius (Trevis.) B.Øllg.
Huperzia itambensis (B.Øllg. & P.G.Windisch) B.Øllg. = Phlegmariurus itambensis (B.Øllg. & P.G.Windisch) B.Øllg.
Huperzia jaegeri (Herter) A.R.Field & Bostock = Phlegmariurus jaegeri (Herter) A.R.Field & Bostock
Huperzia kajewskii (Copel.) A.R.Field & Testo = Phlegmariurus kajewskii (Copel.) A.R.Field & Testo
Huperzia keysseri (Herter & Schltr. ex Diels) comb. ined. = Phlegmariurus keysseri (Herter & Schltr. ex Diels) comb. ined.
Huperzia killipii (Herter) B.Øllg. = Phlegmariurus killipii (Herter) B.Øllg.
Huperzia × koolauensis W.H.Wagner*Phlegmariurus × koolauensis W.H.Wagner
Huperzia kuesteri (Nessel) B.Øllg. = Phlegmariurus kuesteri (Nessel) B.Øllg.
Huperzia lancifolia (Maxon) B.Øllg. = Phlegmariurus lancifolius (Maxon) B.Øllg.
Huperzia lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) A.R.Field & Bostock = Phlegmariurus lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) A.R.Field & Bostock
Huperzia lecomteana (Nessel) A.R.Field & Bostock = Phlegmariurus lecomteanus (Nessel) A.R.Field & Bostock
Huperzia ledermannii (Herter) A.R.Field & Bostock = Phlegmariurus ledermannii (Herter) A.R.Field & Bostock
Huperzia lehnertii Testo = Phlegmariurus lehnertii Testo
Huperzia leptodon (Herter) comb. ined. = Phlegmariurus leptodon (Herter) comb. ined.
Huperzia lignosa (Herter) B.Øllg. = Phlegmariurus lignosus (Herter) B.Øllg.
Huperzia lindenii (Spring) B.Øllg. = Phlegmariurus lindenii (Spring) B.Øllg.
Huperzia linifolia (L.) B.Øllg. = Phlegmariurus linifolius (L.) B.Øllg.
Huperzia llanganatensis (B.Øllg.) B.Øllg. = Phlegmariurus llanganatensis (B.Øllg.) B.Øllg.
Huperzia lockyeri (D.Jones & B.Gray) A.R.Field & Bostock = Phlegmariurus lockyeri (D.Jones & B.Gray) A.R.Field & Bostock
Huperzia loefgreniana (Silveira) B.Øllg. = Phlegmariurus loefgrenianus (Silveira) B.Øllg.
Huperzia longa (Copel.) A.R.Field & Bostock = Phlegmariurus longus (Copel.) A.R.Field & Bostock
Huperzia loxensis (B.Øllg.) B.Øllg. = Phlegmariurus loxensis (B.Øllg.) B.Øllg.
Huperzia luederwaldtii (Nessel) comb. ined. = Phlegmariurus luederwaldtii (Nessel) comb. ined.
Huperzia luteynii B.Øllg. = Phlegmariurus luteynii B.Øllg.
Huperzia macbridei (Herter) B.Øllg. = Phlegmariurus macbridei (Herter) B.Øllg.
Huperzia macgregorii (Baker) A.R.Field & Bostock = Phlegmariurus macgregorii (Baker) A.R.Field & Bostock
Huperzia magnusiana (Herter) A.R.Field & Testo = Phlegmariurus magnusianus (Herter) A.R.Field & Testo
Huperzia mandiocana (Raddi) B.Øllg. = Phlegmariurus mandiocanus (Raddi) B.Øllg.
Huperzia mannii (Hillebr.) W.H.Wagner = Phlegmariurus mannii (Hillebr.) W.H.Wagner
Huperzia marsupiiformis (D.Jones & B.Gray) A.R.Field & Bostock = Phlegmariurus marsupiiformis (D.Jones & B.Gray) A.R.Field & Bostock
Huperzia martii (Wawra) B.Øllg. = Phlegmariurus martii (Wawra) B.Øllg.
Huperzia megastachya (Baker) A.R.Field & Bostock = Phlegmariurus megastachyus (Baker) A.R.Field & Bostock
Huperzia melanesica (Brownlie) A.R.Field & Testo = Phlegmariurus melanesicus (Brownlie) A.R.Field & Testo
Huperzia merrillii (Herter) A.R.Field & Bostock = Phlegmariurus merrillii (Herter) A.R.Field & Bostock
Huperzia mesoamericana (B.Øllg.) B.Øllg. = Phlegmariurus mesoamericanus (B.Øllg.) B.Øllg.
Huperzia mexicana (Herter) B.Øllg. = Phlegmariurus mexicanus (Herter) B.Øllg.
Huperzia mildbraedii (Herter) A.R.Field & Bostock = Phlegmariurus mildbraedii (Herter) A.R.Field & Bostock
Huperzia mingjoui X.C.Zhang = Phlegmariurus mingjoui X.C.Zhang
Huperzia minutifolia (Alderw.) A.R.Field & Bostock = Phlegmariurus minutifolius (Alderw.) A.R.Field & Bostock
Huperzia mirabilis (Willd.) A.R.Field & Testo = Phlegmariurus mirabilis (Willd.) A.R.Field & Testo
Huperzia mollicoma (Spring) B.Øllg. = Phlegmariurus mollicomus (Spring) B.Øllg.
Huperzia molongensis (Herter) B.Øllg. = Phlegmariurus molongensis (Herter) B.Øllg.
Huperzia monticola Kiew = Phlegmariurus monticola Kiew
Huperzia mooreana (Sander ex Baker) B.Øllg. = Phlegmariurus mooreanus (Sander ex Baker) B.Øllg.
Huperzia multifaria (Alderw.) A.R.Field & Bostock = Phlegmariurus multifarius (Alderw.) A.R.Field & Bostock
Huperzia myrsinites (Lam.) B.Øllg. = Phlegmariurus myrsinites (Lam.) B.Øllg.
Huperzia myrtifolia (G.Forst.) A.R.Field & Bostock = Phlegmariurus myrtifolius (G.Forst.) A.R.Field & Bostock
Huperzia myrtuosa (Spring) B.Øllg. = Phlegmariurus myrtuosus (Spring) B.Øllg.
Huperzia neocaledonica (Nessel) A.R.Field & Bostock = Phlegmariurus neocaledonicus (Nessel) A.R.Field & Bostock
Huperzia nesselii (Brause ex Nessel) B.Øllg. = Phlegmariurus nesselii (Brause ex Nessel) B.Øllg.
Huperzia niligarica (Spring) A.R.Field & Bostock = Phlegmariurus niligaricus (Spring) A.R.Field & Bostock
Huperzia nuda (Christ ex Nessel & Hoehne) B.Øllg. = Phlegmariurus nudus (Christ ex Nessel & Hoehne) B.Øllg.
Huperzia nummulariifolia (Blume) Jermy = Phlegmariurus nummulariifolius (Blume) Ching
Huperzia nutans (Brack.) W.H.Wagner = Phlegmariurus nutans (Brack.) W.H.Wagner
Huperzia nyalamensis (Ching & S.K.Wu) H.S.Kung & L.B.Zhang = Phlegmariurus nyalamensis (Ching & S.K.Wu) H.S.Kung & L.B.Zhang
Huperzia obovalifolia (Bonap.) A.R.Field & Testo = Phlegmariurus obovalifolius (Bonap.) A.R.Field & Testo
Huperzia obtusifolia (P.Beauv.) A.R.Field & Bostock = Phlegmariurus obtusifolius (P.Beauv.) A.R.Field & Bostock
Huperzia ocanana (Herter) B.Øllg. = Phlegmariurus ocananus (Herter) B.Øllg.
Huperzia oceaniana (Herter) A.R.Field & Bostock = Phlegmariurus oceanianus (Herter) A.R.Field & Bostock
Huperzia oellgaardii (A.Rojas) B.Øllg. = Phlegmariurus oellgaardii (A.Rojas) B.Øllg.
Huperzia oltmannsii (Herter ex Nessel) A.R.Field & Bostock = Phlegmariurus oltmannsii (Herter ex Nessel) A.R.Field & Bostock
Huperzia ophioglossoides (Lam.) A.R.Field & Bostock = Phlegmariurus ophioglossoides (Lam.) A.R.Field & Bostock
Huperzia orizabae (Underw. & F.E.Lloyd) B.Øllg. = Phlegmariurus orizabae (Underw. & F.E.Lloyd) B.Øllg.
Huperzia ovatifolia (Ching) W.M.Chu ex H.S.Kung & Li Bing Zhang = Phlegmariurus ovatifolius (Ching) W.M.Chu ex H.S.Kung & Li Bing Zhang
Huperzia pachyphylla (Kuhn ex Herter) A.R.Field & Testo = Phlegmariurus pachyphyllus (Kuhn ex Herter) A.R.Field & Testo
Huperzia pachyskelos B.Øllg. = Phlegmariurus pachyskelos B.Øllg.
Huperzia parksii (Copel.) A.R.Field & Bostock = Phlegmariurus parksii (Copel.) A.R.Field & Bostock
Huperzia patentissima (Alderw.) A.R.Field & Bostock = Phlegmariurus patentissimus (Alderw.) A.R.Field & Bostock
Huperzia pearcei (Baker) B.Øllg. = Phlegmariurus pearcei (Baker) B.Øllg.
Huperzia pecten (Baker) A.R.Field & Bostock = Phlegmariurus pecten (Baker) A.R.Field & Bostock
Huperzia perrieriana (Tardieu) A.R.Field & Bostock = Phlegmariurus perrierianus (Tardieu) A.R.Field & Bostock
Huperzia pflanzii (Nessel) B.Øllg. = Phlegmariurus pflanzii (Nessel) B.Øllg.
Huperzia phlegmaria (L.) Rothm. = Phlegmariurus phlegmaria (L.) Holub
Huperzia phlegmarioides (Gaudich.) A.R.Field & Bostock = Phlegmariurus phlegmarioides (Gaudich.) A.R.Field & Bostock
Huperzia phylicifolia (Desv. ex Poir.) B.Øllg. = Phlegmariurus phylicifolius (Desv. ex Poir.) B.Øllg.
Huperzia phyllantha (Hook. & Arn.) R.D.Dixit = Phlegmariurus phyllanthus (Hook. & Arn.) R.D.Dixit
Huperzia picardae (Krug) B.Øllg. = Phlegmariurus picardae (Krug) B.Øllg.
Huperzia pichiana (Tardieu) A.R.Field & Bostock = Phlegmariurus pichianus (Tardieu) A.R.Field & Bostock
Huperzia pinifolia (Trevis.) Kiew = Phlegmariurus pinifolius (Trevis.) Kiew
Huperzia pithyoides (Schltdl. & Cham.) B.Øllg. = Phlegmariurus pithyoides (Schltdl. & Cham.) B.Øllg.
Huperzia pittieri (Christ) B.Øllg. = Phlegmariurus pittieri (Christ) B.Øllg.
Huperzia podocarpensis B.Øllg. = Phlegmariurus podocarpensis B.Øllg.
Huperzia polaris (Nessel) comb. ined. = Phlegmariurus polaris (Nessel) comb. ined.
Huperzia polycarpos (Kunze) B.Øllg. = Phlegmariurus polycarpos (Kunze) B.Øllg.
Huperzia polyclada (Sodiro) comb. ined. = Phlegmariurus polycladus (Sodiro) comb. ined.
Huperzia polydactyla (B.Øllg.) B.Øllg. = Phlegmariurus polydactylus (B.Øllg.) B.Øllg.
Huperzia polylepidetorum (B.Øllg.) B.Øllg. = Phlegmariurus polylepidetorum (B.Øllg.) B.Øllg.
Huperzia pringlei (Underw. & F.E.Lloyd) B.Øllg. = Phlegmariurus pringlei (Underw. & F.E.Lloyd) B.Øllg.
Huperzia prolifera (Blume) A.R.Field & Bostock = Phlegmariurus proliferus (Blume) A.R.Field & Bostock
Huperzia pruinosa (Hieron. & Herter) B.Øllg. = Phlegmariurus pruinosus (Hieron. & Herter) B.Øllg.
Huperzia pseudovaria (Brownlie) comb. ined. = Phlegmariurus pseudovaria (Brownlie) comb. ined.
Huperzia pulcherrima (Hook. & Grev.) Á. & D.Löve = Phlegmariurus pulcherrimus (Hook. & Grev.) Á. & D.Löve
Huperzia pungentifolia (Silveira) B.Øllg. = Phlegmariurus pungentifolius (Silveira) B.Øllg.
Huperzia quadrifariata (Bory) B.Øllg. = Phlegmariurus quadrifariatus (Bory) B.Øllg.
Huperzia recurvifolia (Rolleri) B.Øllg. = Phlegmariurus recurvifolius (Rolleri) B.Øllg.
Huperzia reflexa (Lam.) B.Øllg. = Phlegmariurus reflexus (Lam.) B.Øllg.
Huperzia regnellii (Maxon) B.Øllg. = Phlegmariurus regnellii (Maxon) B.Øllg.
Huperzia ribourtii (Herter) A.R.Field & Bostock = Phlegmariurus ribourtii (Herter) A.R.Field & Bostock
Huperzia riobambensis (Nessel) B.Øllg. = Phlegmariurus riobambensis (Nessel) B.Øllg.
Huperzia robusta (Klotzsch) B.Øllg. = Phlegmariurus robustus (Klotzsch) B.Øllg.
Huperzia rosenstockiana (Herter) B.Øllg. = Phlegmariurus rosenstockianus (Herter) B.Øllg.
Huperzia rostrifolia (Silveira) B.Øllg. = Phlegmariurus rostrifolius (Silveira) B.Øllg.
Huperzia ruber (Cham. & Schltdl.) B.Øllg. = Phlegmariurus ruber (Cham. & Schltdl.) B.Øllg.
Huperzia rubrica (Herter) A.R.Field & Bostock = Phlegmariurus rubricus (Herter) A.R.Field & Bostock
Huperzia rufescens (Hook.) B.Øllg. = Phlegmariurus rufescens (Hook.) B.Øllg.
Huperzia rupicola (Alderw.) A.R.Field & Bostock = Phlegmariurus rupicola (Alderw.) A.R.Field & Bostock
Huperzia sagasteguiana (B.Øllg.) B.Øllg. = Phlegmariurus sagasteguianus (B.Øllg.) B.Øllg.
Huperzia salvinioides (Herter) Ching = Phlegmariurus salvinioides (Herter) Ching
Huperzia samoana (Herter ex Nessel) A.R.Field & Bostock = Phlegmariurus samoanus (Herter ex Nessel) A.R.Field & Bostock
Huperzia sarmentosa (Spring) B.Øllg. = Phlegmariurus sarmentosus (Spring) B.Øllg.
Huperzia saurura (Lam.) B.Øllg. = Phlegmariurus saururus (Lam.) B.Øllg.
Huperzia scabrida (B.Øllg.) B.Øllg. = Phlegmariurus scabridus (B.Øllg.) B.Øllg.
Huperzia schlechteri (E.Pritz.) A.R.Field & Bostock = Phlegmariurus schlechteri (E.Pritz.) A.R.Field & Bostock
Huperzia schlimii (Herter) B.Øllg. = Phlegmariurus schlimii (Herter) B.Øllg.
Huperzia schmidtchenii (Hieron.) B.Øllg. = Phlegmariurus schmidtchenii (Hieron.) B.Øllg.
Huperzia sellifolia (B.Øllg.) B.Øllg. = Phlegmariurus sellifolius (B.Øllg.) B.Øllg.
Huperzia sellowiana (Herter) B.Øllg. = Phlegmariurus sellowianus (Herter) B.Øllg.
Huperzia serpentiformis (Herter) B.Øllg. = Phlegmariurus serpentiformis (Herter) B.Øllg.
Huperzia setifolia (Alderw.) A.R.Field & Bostock = Phlegmariurus setifolius (Alderw.) A.R.Field & Bostock
Huperzia shangsiensis C.Y.Yang = Phlegmariurus shangsiensis C.Y.Yang
Huperzia sieberiana (Spring) B.Øllg. = Phlegmariurus sieberianus (Spring) B.Øllg.
Huperzia sieboldii (Miq.) Ching = Phlegmariurus sieboldii (Miq.) Ching
Huperzia silveirae (Herter ex Nessel & Hoehne) B.Øllg. = Phlegmariurus silveirae (Herter ex Nessel & Hoehne) B.Øllg.
Huperzia silverstonei B.Øllg. = Phlegmariurus silverstonei B.Øllg.
Huperzia sintenisii (Herter) B.Øllg. = Phlegmariurus sintenisii (Herter) B.Øllg.
Huperzia societensis (J.W.Moore) A.R.Field = Phlegmariurus societensis (J.W.Moore) A.R.Field
Huperzia socratis (Herter) comb. ined. = Phlegmariurus socratis (Herter) comb. ined.
Huperzia sooiana Lawalrée = Phlegmariurus sooianus Lawalrée
Huperzia sotae (Rolleri) B.Øllg. = Phlegmariurus sotae (Rolleri) B.Øllg.
Huperzia sphagnicola B.Øllg. = Phlegmariurus sphagnicola B.Øllg.
Huperzia squarrosa (G.Forst.) Trevis. = Phlegmariurus squarrosus (G.Forst.) Á. & D.Löve
Huperzia staudtii (Nessel)  = Phlegmariurus staudtii (Nessel) A.R.Field & Bostock
Huperzia stemmermanniae A.C.Medeiros & Wagner = Phlegmariurus stemmermanniae A.C.Medeiros & Wagner
Huperzia stephani B.Øllg. = Phlegmariurus stephani B.Øllg.
Huperzia stricta (Baker) A.R.Field & Bostock = Phlegmariurus strictus (Baker) A.R.Field & Bostock
Huperzia subfalciformis (Alderw.) A.R.Field & Bostock = Phlegmariurus subfalciformis (Alderw.) A.R.Field & Bostock
Huperzia subintegra (Hillebr.) comb. ined. = Phlegmariurus subintegrus (Hillebr.) comb. ined.
Huperzia subtrifoliata (Brownlie) A.R.Field & Bostock = Phlegmariurus subtrifoliatus (Brownlie) A.R.Field & Bostock
Huperzia subtubulosa (Herter) comb. ined. = Phlegmariurus subtubulosus (Herter) comb. ined.
Huperzia subulata (Desv. ex Poir.) B.Øllg. = Phlegmariurus subulatus (Desv. ex Poir.) B.Øllg.
Huperzia tafanensis (Nessel) comb. ined. = Phlegmariurus tafanensis (Nessel) comb. ined.
Huperzia talamancana (B.Øllg.) B.Øllg. = Phlegmariurus talamancanus (B.Øllg.) B.Øllg.
Huperzia talamauana (Alderw.) A.R.Field & Bostock = Phlegmariurus talamauanus (Alderw.) A.R.Field & Bostock
Huperzia talpiphila (B.Øllg.) B.Øllg. = Phlegmariurus talpiphilus (B.Øllg.) B.Øllg.
Huperzia tardieuae (Herter) A.R.Field & Testo = Phlegmariurus tardieuae (Herter) A.R.Field & Testo
Huperzia tauri (Herter) A.R.Field & Testo = Phlegmariurus tauri (Herter) A.R.Field & Testo
Huperzia taxifolia (Sm.) Á. & D.Löve = Phlegmariurus taxifolius (Sm.) Á. & D.Löve
Huperzia tenuicaulis (Underw. & F.E.Lloyd) B.Øllg. = Phlegmariurus tenuicaulis (Underw. & F.E.Lloyd) B.Øllg.
Huperzia tenuis (Humb. & Bonpl. ex Willd.) B.Øllg. = Phlegmariurus tenuis (Humb. & Bonpl. ex Willd.) B.Øllg.
Huperzia teretirigida (Rakotondr.) comb. ined. = Phlegmariurus teretirigidus (Rakotondr.) comb. ined.
Huperzia terrae-guilelmii (Herter) A.R.Field & Bostock = Phlegmariurus terrae-guilelmii (Herter) A.R.Field & Bostock
Huperzia tetragona (Hook. & Grev.) B.Øllg. = Phlegmariurus tetragonus (Hook. & Grev.) B.Øllg.
Huperzia tetrastichoides (A.R.Field & Bostock) A.R.Field & Bostock = Phlegmariurus tetrastichoides (A.R.Field & Bostock) A.R.Field & Bostock
Huperzia tetrasticha (Kunze) A.R.Field & Bostock = Phlegmariurus tetrastichus (Kunze) A.R.Field & Bostock
Huperzia toppingii (Herter) A.R.Field & Bostock = Phlegmariurus toppingii (Herter) A.R.Field & Bostock
Huperzia tournayana Lawalrée = Phlegmariurus tournayanus Lawalrée
Huperzia transilla (Sodiro ex Baker) B.Øllg. = Phlegmariurus transilla (Sodiro ex Baker) B.Øllg.
Huperzia treitubensis (Silveira) B.Øllg. = Phlegmariurus treitubensis (Silveira) B.Øllg.
Huperzia trifoliata (Copel.) A.R.Field & Bostock = Phlegmariurus trifoliatus (Copel.) A.R.Field & Bostock
Huperzia trigona (C.Chr.) A.R.Field & Bostock = Phlegmariurus trigonus (C.Chr.) A.R.Field & Bostock
Huperzia tryonorum B.Øllg. = Phlegmariurus tryonorum B.Øllg.
Huperzia tubulosa (Maxon) B.Øllg. = Phlegmariurus tubulosus (Maxon) B.Øllg.
Huperzia ulei (Herter) comb. ined. = Phlegmariurus ulei (Herter) comb. ined.
Huperzia ulixis (Herter) B.Øllg. = Phlegmariurus ulixis (Herter) B.Øllg.
Huperzia unguiculata (B.Øllg.) B.Øllg. = Phlegmariurus unguiculatus (B.Øllg.) B.Øllg.
Huperzia urbani (Herter) B.Øllg. = Phlegmariurus urbani (Herter) B.Øllg.
Huperzia vanuatuensis A.R.Field = Phlegmariurus vanuatuensis A.R.Field
Huperzia varia (R.Br.) A.R.Field & Bostock = Phlegmariurus varius (R.Br.) A.R.Field & Bostock
Huperzia venezuelanica (Herter) B.Øllg. = Phlegmariurus venezuelanicus (Herter) B.Øllg.
Huperzia vernicosa (Hook. & Grev.) Á. & D.Löve = Phlegmariurus vernicosus (Hook. & Grev.) Á. & D.Löve
Huperzia villonacensis (B.Øllg.) B.Øllg. = Phlegmariurus villonacensis (B.Øllg.) B.Øllg.
Huperzia warneckei (Herter ex Nessel) A.R.Field & Testo = Phlegmariurus warneckei (Herter ex Nessel) A.R.Field & Testo
Huperzia watsoniana (Maxon) B.Øllg. = Phlegmariurus watsonianus (Maxon) B.Øllg.
Huperzia weberbaueri (Nessel) B.Øllg. = Phlegmariurus weberbaueri (Nessel) B.Øllg.
Huperzia weddellii (Herter) B.Øllg. = Phlegmariurus weddellii (Herter) B.Øllg.
Huperzia wilsonii (Underw. & F.E.Lloyd) B.Øllg. = Phlegmariurus wilsonii (Underw. & F.E.Lloyd) B.Øllg.
Huperzia xiphophylla (Baker) A.R.Field & Bostock = Phlegmariurus xiphophyllus (Baker) A.R.Field & Bostock
Huperzia yunnanensis Ching = Phlegmariurus yunnanensis Ching

Distribusi dan habitat 
Sesuai batasan dalam klasifikasi PPG I, Huperzia didistribusikan di habitat beriklim sedang, Arktik dan alpine, termasuk juga yang ada di pegunungan di Asia tropis.  Spesiesnya terestrial atau epifit.  Phlegmariurus adalah epifit, dan memiliki distribusi tropis di seluruh dunia,  sehingga ketika Huperzia didefinisikan secara luas untuk mencakup ketiga genera dari subfamili Huperzioideae, ia memiliki distribusi hampir di seluruh dunia, tidak ada terutama di Afrika Utara, Semenanjung Arab dan Asia Barat.

Rujukan 

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Lycopodiophyta