Nyra (books)

Nyra is the secondary antagonist of the Guardians of Gahoole book series, serving as the secondary antagonist of the first half and became the main antagonist in The Burning. 
She is the mate of Kludd and mother of Coryn, 
After her husband's death at the hands of Twilight, Nyra becomes he leader of the Pure Ones and wants to make the Guardians pay for i.