Mackerras

Decription of Genera and Speces
1 the Plotyzosteria Group 
1.1 Genus Polyzosteria 
Diagnostic features.-Large, apterous Polyzosteriini, distinguished from other
apterous genera of the tribe by 
the margins of the pronotum forming low,
rolled edges, not swollen nor angularly raised,
angles of T7 being acute, 
tarsal segments being bare, and
the dorsal surfaces of the tibiae 
usually being flattened with the spines in 2 rows.
1.2 Genus Euzosteria 
Diagnostic features. Usually medium-sized, apterous Polyzosteriini, 
distinguished from other genera of the tribe by
 the margins of the pronotum being angularly raised, giving the tergum
an excavated appearance; 
 the posterior margin of the metanotum being straight or slightly convex in
the midline.
1.3 Genus Zonioploca 
Small to medium-sized, apterous Polyzosteriini, distinguished from other apterous
genera of the tribe by 
the lateral margins of the pronotum being smoothly swollen,
those of the metanotum very slightly or not at all swollen,
the angles of T7 being acutely produced,
the dorsum being tuberculate or pitted,
the tarsal segments being bare.
1.4 Genus Eppeertia 
Medium-sized to large, apterous Polyzosteriini, like Zonioploca, but more 
thickset; with all thoracic margins thickened in furcata group.
1.5 Genus Megazosteria 
Large, apterous Polyzosteriini, distinguished form other genera of the tribe by
having very broadly expanded thoracic margins with slightly upturned edges giving
a shallowly excavated apperance to the pronotum
the posterior margin of the metanotum being concave in the middle third and convex
laterally on each side, obscuring nearly all T1.
1.6 Genus Desmozosteria 
Medium-sized to fairly large, apterous Polyzosteriini, distinguished from other
genera of the tribe by 
the lateral margins of the pronotum being smoothly swollen ( at least to some extent).
the angles of the abdominal terga being rectangular or forming minute projections
the surface being pitted, but not tuberculate
the tarsal segment being bare.
1.7 Genus Anamesia 
Medium-sized to fairly large, apterous Polyzosteriini, distinguished from other
genera of the tribe by
the thoracic margins not being swollen or upturned
the pronotum usually being considerably narrower than the mesonotum, and often
rather flattened
the angles of all abdominal terga being rectangular or obtuse
the dorsum being pitted, but not tuberculate
the sclerites R2a and R2b of the male genitalia being smoothly rounded without
suprlike process.
1.8 Genus Pseudolampra 
Medium-sized, apterous Polyzosteriini, similar to Anamesia, but distinguished
from it by
being soft body, and somewhat egg-shaped;
often speckled like Epilamprinae
having small, dark, spur-like processes on the sclerites R2a and R2b of the 
male genitalia.
1.9 Genus Cosmozosteria 
Medium-sized, oval or ovoid, apterous, slightly convex Polyzosteriini, 
distinguished from other genera of the tribe by having
dorsum of thorax relatively smooth and that of abdomen becoming progressively
ridged and tuberculate posteriorly
angles of T6 and T7 forming short, sharp projections
male genitalia with R1 ending with 2 finger-like processes set at right angles
to each other, and with R2a and R2b armed with strong darks spurs
spines on the dorsal surfaces of the tibiae in 3 rows
tarsal claws decidedly asymmetrical.
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2 the Platyzosteria Group (VIII) 
2.1 Genus Platyzosteria 
Usually medium-sized to large, occasionally small;
oval and convex or flattened; apterous or with vestigial tegmina; no wing 
vestiges;dorsal surface usually smooth and shining to the naked eye,
sometimes obviously rugose or scabrous, always microscopically pitted
or wrinkled, with minute hairs; predominant colours black, brown, deep
reddish brown, occasionally yellow. Distinguished from other genera of
Polyzosteriini by
1 the pronotum usually being smoothly convex, without upturned edges;
2 the dorsal surface being smooth,rugose, or scabrous, 
but not covered with closely set rounded tubercles as in some
species of Zonioploca; 
3 the angles of T2-T7 being acute, especially
those of T6 and T7 which sometimes form long tapering spines;
4 the tibiae being triseriately spined on their dorsal surfaces;
5 the ventral surfaces of the metatarsi being bare( subgenera Platyzosteria
and Leptozosteria) or spined ( most of subgenus Melanozosteria).
subgenus Platyzosteria
The larger species are usually smooth and somewhat flattened (melanaria
group), or scabrous and convex (scabra group), in contrast to the larger species
of the subgenus Melanozosteria which are usually smooth and convex. The smaller
species are usually scabrous and convex.
Thorax.-Usually apterous or with small vestigial tegmina, not usually separated
from mesonotum for more than about one-half of its length.
Legs.-Fore coxae usually fairly smooth; femoral comb often restricted
to distal half of margin, sometimes occupying two-thirds or more of margin; tarsi
short and bare, hind metatarsus not longer than the remaining segments together,
its pulvillus produced for nearly whole length; claws asymmetrical.
Abdomen.-Tergal angles usually all sharply produced, including those of T9;
lateral margins of T7 and TI0 often serrate; posterolateral margins of S7 and S8
in male usually serrate or tuberculate; cerci often short, sometimes long.
subgenus Melanozosteria
The larger species are usually plump and smooth, in contrast to those belonging
to Platyzostevia (s.s.) which are either rather flattened and smooth, or convex
and scabrous. When vestigial tegmina are present they are often separated 
from the mesonotum for almost its whole length. Apterous species are usually
small.
Fore coxae are often strongly pitted or ridged and with hairs or spinules;
femoral comb composed of fairly large spines usually of unequal size, occupying from
about two-thirds to almost whole of margin; tarsi may be short or fairly long, hind
metatarsus may be short with a short row of spines, or few spines, or occasionally no
spines, or it may be long and strongly spined; spines sometimes present on mid
metatarsi and fore metatarsi; claws usually asymmetrical, occasionally symmetrical
or practically so.
Posterolateral angles of T2-T5 are acute, usually short and little produced,
those of T6 and T7 sharply produced, those of T9 usually blunt; lateral margins of
T7 and TI0 smooth or serrate; posterolateral margins of S7 and S8 in male usually
smooth; cerci usually long and stout.
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subgenus Leptpzosteria
Large to very large, broadly oval, flattened, apterous; dorsum smooth, lateral
margins of proximal abdominal terga strongly serrate; sterna scabrous and serrate
laterally.
Head.-Somewhat flattened, usually smooth, shining, sometimes depressed
and slightly rugose.
Thorax.-Very broad, mesonotum and metanotum usually as wide or wider
than abdomen, surface smooth with some shallow depressions, microscopically
rugose; margins slightly thickened; no trace of tegmina but angles of metanotum
usually slightly produced.
Legs.-Coxae smooth, borders usually pale and processes brown; spines of
femoral comb usually small, sometimes reduced in number to 3-8, usually confined
to distal third of margin, spines on posterolateral margin of fore femur sometimes
reduced to 3 and placed near distal end; middle femora usually flattened
proximally,the ventral groove being more or less obliterated in its proximal 
third; spines on middle and hind femora reduced in size and sometimes in 
number; tarsi short and bare as in subgenus Platyzostevia; all pulvilli and 
arolia large, claws usually only slightly asymmetrical.
Abdomen.-Sometimes appears abnormally extended with the smooth anterior
part of each tergum visible; surface and posterior margins of terga fairly smooth,
surface of 77 often beset with short stiff hairs; lateral margins of proximal terga
sometimes uneven or serrate, those of T6 and T7 always serrate; angles of T2-T7
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