Repository: ontodev/robot
Pull request #578 (2019-10-11 21:25:11)

Add a collapse command
# Collapse

Sometimes, a class hierarchy can contain more intermediate classes than necessary, especially when extracting modules. ROBOT includes `collapse` to remove intermediate classes based on a minimal number of subclasses, using the `--threshold` option.

```
robot collapse \
 --input module.owl \
 --threshold 3 \
 --output minimized_module.owl
```

Any intermediate class (has one or more subclasses) that has *less* than the threshold number of subclasses will be removed. Top-level classes (do not have a named superclass) and bottom-level classes (do not have any subclasses) will not be removed. 

For example, given `--threshold 2`:

```
- class:A
    - class:B
    - class:C
        - class:D
           - class:E
    - class:F
        - class:G
        - class:H
```

Becomes:

```
- class:A
    - class:B
    - class:E
    - class:F
            - class:G
            - class:H
```

`class:C` and `class:D` are removed because they each only have one subclass. `class:F` is kept because it has two subclasses, which is the threshold.

If there are any classes that you don't want removed, you can keep them regardless of the number of subclasses using `--precious <IRI/CURIE>` (for a set of terms in a file, use `--precious-terms <term-file>`). 

    robot collapse \
     --input uberon_module.owl \
     --threshold 3 \
     --precious UBERON:0000483 \
     --output results/uberon_minimized.owl

For example, given `--threshold 2` and `--precious class:D`, that same example from above would become:

```
- class:A
    - class:B
    - class:D
        - class:E
    - class:F
            - class:G
            - class:H
```



Patch:
diff --git a/docs/collapse.md b/docs/collapse.md
new file mode 100644
index 00000000..ea422bfb
--- /dev/null
+++ b/docs/collapse.md
@@ -0,0 +1,71 @@
+# Collapse
+
+Sometimes, a class hierarchy can contain more intermediate classes than necessary, especially when extracting modules. ROBOT includes `collapse` to remove intermediate classes based on a minimal number of subclasses, using the `--threshold` option. If `--threshold` is not provided, it will default to `2`. The threshold must be a positive integer greater than or equal to 2.
+
+```
+robot collapse \
+ --input module.owl \
+ --threshold 3 \
+ --output minimized_module.owl
+```
+
+Collapse will not remove root classes or leaf classes (i.e. top-level classes, without a named superclass, and bottom-level classes, without any subclasses). For each intermediate class, if it has *fewer* subclasses than the threshold, then it will be removed.
+
+For example, given `--threshold 2`:
+
+```
+- class:A
+    - class:B
+    - class:C
+        - class:D
+           - class:E
+    - class:F
+        - class:G
+        - class:H
+```
+
+Becomes:
+
+```
+- class:A
+    - class:B
+    - class:E
+    - class:F
+            - class:G
+            - class:H
+```
+
+`class:C` and `class:D` are removed because they each only have one subclass. `class:F` is kept because it has two subclasses, which is the threshold.
+
+If there are any classes that you don't want removed, you can keep them regardless of the number of subclasses using `--precious <IRI/CURIE>` (for a set of terms in a file, use `--precious-terms <term-file>`). 
+
+    robot collapse \
+     --input uberon_module.owl \
+     --threshold 3 \
+     --precious UBERON:0000483 \
+     --output results/uberon_minimized.owl
+
+For example, given `--threshold 2` and `--precious class:D`, that same example from above would become:
+
+```
+- class:A
+    - class:B
+    - class:D
+        - class:E
+    - class:F
+            - class:G
+            - class:H
+```
+ 
+ ---
+ 
+ ## Error Messages
+ 
+ ### Threshold Error
+ 
+ The `--threshold` input must be an integer.
+ 
+ ### Threshold Value Error
+ 
+ The `--threshold` input must be an integer greater than 1.
+
diff --git a/docs/examples/uberon_minimized.owl b/docs/examples/uberon_minimized.owl
new file mode 100644
index 00000000..a6cd9b99
--- /dev/null
+++ b/docs/examples/uberon_minimized.owl
@@ -0,0 +1,2690 @@
+<?xml version="1.0"?>
+<rdf:RDF xmlns="http://purl.obolibrary.org/obo/uberon.owl#"
+     xml:base="http://purl.obolibrary.org/obo/uberon.owl"
+     xmlns:obo="http://purl.obolibrary.org/obo/"
+     xmlns:owl="http://www.w3.org/2002/07/owl#"
+     xmlns:rdf="http://www.w3.org/1999/02/22-rdf-syntax-ns#"
+     xmlns:xml="http://www.w3.org/XML/1998/namespace"
+     xmlns:xsd="http://www.w3.org/2001/XMLSchema#"
+     xmlns:foaf="http://xmlns.com/foaf/0.1/"
+     xmlns:rdfs="http://www.w3.org/2000/01/rdf-schema#"
+     xmlns:oboInOwl="http://www.geneontology.org/formats/oboInOwl#">
+    <owl:Ontology rdf:about="http://purl.obolibrary.org/obo/uberon.owl"/>
+    
+
+
+    <!-- 
+    ///////////////////////////////////////////////////////////////////////////////////////
+    //
+    // Annotation properties
+    //
+    ///////////////////////////////////////////////////////////////////////////////////////
+     -->
+
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/IAO_0000115 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/IAO_0000115"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/IAO_0000116 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/IAO_0000116"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/IAO_0000232 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/IAO_0000232"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/RO_0002174 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/RO_0002174"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/RO_0002175 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/RO_0002175"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBPROP_0000001 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBPROP_0000003 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBPROP_0000007 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/UBPROP_0000007"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBPROP_0000008 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/UBPROP_0000008"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBPROP_0000009 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/UBPROP_0000009"/>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBPROP_0000012 -->
+
+    <owl:AnnotationProperty rdf:about="http://purl.obolibrary.org/obo/UBPROP_0000012"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#date_retrieved -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#date_retrieved"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#external_class -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#external_class"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#external_ontology -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#external_ontology"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasAlternativeId -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasAlternativeId"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasBroadSynonym -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasBroadSynonym"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasDbXref -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasExactSynonym -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasNarrowSynonym -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasNarrowSynonym"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasOBONamespace -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasOBONamespace"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#hasSynonymType -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#hasSynonymType"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#id -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#id"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#inSubset -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#inSubset"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#notes -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#notes"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#ontology -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#ontology"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#shorthand -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#shorthand"/>
+    
+
+
+    <!-- http://www.geneontology.org/formats/oboInOwl#source -->
+
+    <owl:AnnotationProperty rdf:about="http://www.geneontology.org/formats/oboInOwl#source"/>
+    
+
+
+    <!-- http://xmlns.com/foaf/0.1/depicted_by -->
+
+    <owl:AnnotationProperty rdf:about="http://xmlns.com/foaf/0.1/depicted_by"/>
+    
+
+
+    <!-- 
+    ///////////////////////////////////////////////////////////////////////////////////////
+    //
+    // Object Properties
+    //
+    ///////////////////////////////////////////////////////////////////////////////////////
+     -->
+
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/BFO_0000050 -->
+
+    <owl:ObjectProperty rdf:about="http://purl.obolibrary.org/obo/BFO_0000050">
+        <owl:inverseOf rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+        <rdf:type rdf:resource="http://www.w3.org/2002/07/owl#TransitiveProperty"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BFO:0000050</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">part_of</oboInOwl:id>
+        <oboInOwl:shorthand rdf:datatype="http://www.w3.org/2001/XMLSchema#string">part_of</oboInOwl:shorthand>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">part_of</rdfs:label>
+    </owl:ObjectProperty>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/BFO_0000051 -->
+
+    <owl:ObjectProperty rdf:about="http://purl.obolibrary.org/obo/BFO_0000051">
+        <rdf:type rdf:resource="http://www.w3.org/2002/07/owl#TransitiveProperty"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BFO:0000051</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">has_part</oboInOwl:id>
+        <oboInOwl:shorthand rdf:datatype="http://www.w3.org/2001/XMLSchema#string">has_part</oboInOwl:shorthand>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">has_part</rdfs:label>
+    </owl:ObjectProperty>
+    
+
+
+    <!-- 
+    ///////////////////////////////////////////////////////////////////////////////////////
+    //
+    // Classes
+    //
+    ///////////////////////////////////////////////////////////////////////////////////////
+     -->
+
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000062 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000062">
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure that performs a specific function or group of functions [WP].</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organs are commonly observed as visibly distinct structures, but may also exist as loosely associated clusters of cells that work together to perform a specific function or functions.</obo:UBPROP_0000001>
+        <obo:UBPROP_0000012 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO v1 does not include a generic &apos;organ&apos; class, only simple and compound organ. CARO v2 may include organ, see https://github.com/obophenotype/caro/issues/4</obo:UBPROP_0000012>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Organ_(anatomy)"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0178784"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Organ"/>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_4"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/272625005"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000634</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35949</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ENVO:01000162</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:67498</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0003001</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rv5XMb5wpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rwP3iWpwpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0178784</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0003760</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">anatomical unit</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body organ</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">element</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000062</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organ</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure that performs a specific function or group of functions [WP].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Organ_(anatomy)"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organs are commonly observed as visibly distinct structures, but may also exist as loosely associated clusters of cells that work together to perform a specific function or functions.</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO:0048513</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0178784</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Organ</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">element</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:cjm</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000064 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000064">
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure which has as its direct parts two or more types of tissue and is continuous with one or more anatomical structures likewise constituted by two or more portions of tissues distinct from those of their complement. Examples: osteon, cortical bone, neck of femur, bronchopulmonary segment, left lobe of liver, anterior right side of heart, interventricular branch of left coronary artery, right atrium, mitral valve, head of pancreas[FMA].</obo:IAO_0000115>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_16"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/113343008"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/91717005"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0011124</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000635</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:82472</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cardinal organ part</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">regional part of organ</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000064</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#disjointness_axiom_removed"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organ part</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000064"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure which has as its direct parts two or more types of tissue and is continuous with one or more anatomical structures likewise constituted by two or more portions of tissues distinct from those of their complement. Examples: osteon, cortical bone, neck of femur, bronchopulmonary segment, left lobe of liver, anterior right side of heart, interventricular branch of left coronary artery, right atrium, mitral valve, head of pancreas[FMA].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:82472</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000064"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">regional part of organ</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">NIFSTD:birnlex_16</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000383 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000383">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002204"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of all the muscles of the body[VSAO, modified].</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of the muscles of the body.[VSAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000012 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">we place the MA term musculature here, rather than under uberon:musculature, as this seems more appropriate given the structure of MA</obo:UBPROP_0000012>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000307</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000088</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001369</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001485</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000801</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35578</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00005069</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:72954</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0002888</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000025</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000025</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000033</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0004042</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscle system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscle system of body</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscular system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculature system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all muscles</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of muscles of body</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">vertebrate muscular system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muskelsystem</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000383</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculature of body</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of all the muscles of the body[VSAO, modified].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000033</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of the muscles of the body.[VSAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-08-14</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000033</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:curator</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000012"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">we place the MA term musculature here, rather than under uberon:musculature, as this seems more appropriate given the structure of MA</owl:annotatedTarget>
+        <oboInOwl:external_ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA</oboInOwl:external_ontology>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscular system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:72954</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculature system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000088</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all muscles</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:72954</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of muscles of body</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:72954</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">vertebrate muscular system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001369</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muskelsystem</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001485</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000467 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000467">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000480"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical group that is has as its parts distinct anatomical structures interconnected by anatomical structures at a lower level of granularity[CARO]. A group of organs that work together to perform a certain task [Wikipedia].</obo:IAO_0000115>
+        <oboInOwl:hasBroadSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">system</oboInOwl:hasBroadSynonym>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Organ_system"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0460002"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Organ_System"/>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_14"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/278195005"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000007</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000011</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000011</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BSA:0000049</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-2088</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000011</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003011</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA:392</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:16103</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100000</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00004856</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:7149</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000011</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000003</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rCWM0QCtDEdyAAADggVbxzQ</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001439</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TGMA:0001831</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0460002</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001725</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0005746</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0005763</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003002</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001439</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:AnatomicalSystem</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organ system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000467</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">anatomical system</rdfs:label>
+        <rdfs:seeAlso rdf:resource="http://purl.obolibrary.org/obo/EHDAA2_0001330"/>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical group that is has as its parts distinct anatomical structures interconnected by anatomical structures at a lower level of granularity[CARO]. A group of organs that work together to perform a certain task [Wikipedia].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Organ_system"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000011</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasBroadSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO:0048731</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0460002</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Organ_System</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">NIFSTD:birnlex_14</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000468 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000468">
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure that is an individual member of a species and consists of more than one cell.</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TODO - split body and mc organism? body continues after death stage</obo:IAO_0000116>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organismal</obo:UBPROP_0000007>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Multi-cellular_organism"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Whole_Organism"/>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_18"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/243928005"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010026</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000191</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000012</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BSA:0000038</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000042</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000012</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0002906</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003103</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003191</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA:1</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA:9178</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:25765</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100016</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00000001</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:256135</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000012</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TADS:0000001</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001094</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TGMA:0001832</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000671</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0007833</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003004</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001094</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:Organism</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">multi-cellular organism</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organism</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">whole organism</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasNarrowSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">animal</oboInOwl:hasNarrowSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Koerper</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">whole body</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000468</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">multicellular organism</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure that is an individual member of a species and consists of more than one cell.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Multi-cellular_organism"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000012</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">multi-cellular organism</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000012</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasNarrowSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">animal</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000042</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Koerper</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001489</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">whole body</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001489</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000475 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000475">
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom>
+                    <owl:Restriction>
+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001009"/>
+                    </owl:Restriction>
+                </owl:someValuesFrom>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom>
+                    <owl:Restriction>
+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001016"/>
+                    </owl:Restriction>
+                </owl:someValuesFrom>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom>
+                    <owl:Restriction>
+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+                    </owl:Restriction>
+                </owl:someValuesFrom>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom>
+                    <owl:Restriction>
+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+                    </owl:Restriction>
+                </owl:someValuesFrom>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure which is a subdivision of a whole organism, consisting of components of multiple anatomical systems, largely surrounded by a contiguous region of integument.</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Reflects CARO2. todo - check the inclusion of FMA &apos;cardinal body part here&apos;, and check child terms for consistency</obo:IAO_0000116>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Body_part"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0229962"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Body_Part"/>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_7"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010053</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-2084</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000808</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:36031</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00007009</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:7153</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0002433</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000293</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A01</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000293</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rvViAHJwpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001308</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TGMA:0001840</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0229962</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001758</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003013</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001308</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:BodyPart</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">anatomic region</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body part</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body region</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cardinal body part</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000475</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organism subdivision</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure which is a subdivision of a whole organism, consisting of components of multiple anatomical systems, largely surrounded by a contiguous region of integument.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:DOS</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0229962</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Body_Part</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">anatomic region</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0002433</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body region</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:7153</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cardinal body part</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:7153</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000479 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000479">
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Multicellular anatomical structure that consists of many cells of one or a few types, arranged in an extracellular matrix such that their long-range organisation is at least partly a repetition of their short-range organisation.</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">changed label and definition to reflect CARO2</obo:IAO_0000116>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0040300"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Tissue"/>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_19"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000607</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000043</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000043</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-2090</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000043</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003043</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35868</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00007003</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9637</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000043</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0003002</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A10</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001477</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TGMA:0001844</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0040300</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001757</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0005729</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003040</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001477</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:Tissue</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">portion of tissue</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">tissue portion</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasNarrowSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">simple tissue</oboInOwl:hasNarrowSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000479</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">tissue</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000479"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Multicellular anatomical structure that consists of many cells of one or a few types, arranged in an extracellular matrix such that their long-range organisation is at least partly a repetition of their short-range organisation.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000043</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000479"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0040300</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Tissue</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000479"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">portion of tissue</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000043</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000479"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasNarrowSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">simple tissue</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000043</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000480 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000480">
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000468"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure consisting of at least two non-overlapping organs, multi-tissue aggregates or portion of tissues or cells of different types that does not constitute an organism, organ, multi-tissue aggregate, or portion of tissue.</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Will be obsoleted in v2 of CARO [https://github.com/obophenotype/caro/issues/2]</obo:IAO_0000116>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010008</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001512</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TGMA:0001846</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001724</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003001</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001512</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000480</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">anatomical group</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000480"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure consisting of at least two non-overlapping organs, multi-tissue aggregates or portion of tissues or cells of different types that does not constitute an organism, organ, multi-tissue aggregate, or portion of tissue.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000483 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000483">
+        <owl:equivalentClass>
+            <owl:Class>
+                <owl:unionOf rdf:parseType="Collection">
+                    <rdf:Description rdf:about="http://purl.obolibrary.org/obo/UBERON_0000486"/>
+                    <rdf:Description rdf:about="http://purl.obolibrary.org/obo/UBERON_0000490"/>
+                </owl:unionOf>
+            </owl:Class>
+        </owl:equivalentClass>
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000479"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0005769"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Portion of tissue, that consists of one or more layers of epithelial cells connected to each other by cell junctions and which is underlain by a basal lamina. Examples: simple squamous epithelium, glandular cuboidal epithelium, transitional epithelium, myoepithelium[CARO].</obo:IAO_0000115>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The two basic types of metazoan tissue are epithelial and connective. The simplest metazoans, and developmental stages of many primitive invertebrates, consist solely of these two layers. Thus, epithelial and connective tissues may be the primary (original) tissues of metazoans, and both are important in the functional organization of animals.[well established][VHOG]</obo:UBPROP_0000003>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epithelial</obo:UBPROP_0000007>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Epithelium"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0014609"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Epithelium"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/31610004"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000144</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010055</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000066</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000066</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000416</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-0288</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000066</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003066</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:32738</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00007005</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9639</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:402</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000066</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0003060</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A10.272</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001486</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0014609</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000387</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003045</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001486</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epithelial tissue</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">portion of epithelium</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000483</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epithelium</rdfs:label>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/c/c3/Tkanka_nablonkowa.png"/>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Portion of tissue, that consists of one or more layers of epithelial cells connected to each other by cell junctions and which is underlain by a basal lamina. Examples: simple squamous epithelium, glandular cuboidal epithelium, transitional epithelium, myoepithelium[CARO].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Epithelium"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The two basic types of metazoan tissue are epithelial and connective. The simplest metazoans, and developmental stages of many primitive invertebrates, consist solely of these two layers. Thus, epithelial and connective tissues may be the primary (original) tissues of metazoans, and both are important in the functional organization of animals.[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000387</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ISBN:978-0030259821 Ruppert EE, Fox RS, Barnes RD, Invertebrate zoology: a functional evolutionary approach (2003) p.59</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0014609</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Epithelium</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">portion of epithelium</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9639</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000486 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000486">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium which consists of more than one layer of epithelial cells that may or may not be in contact with a basement membrane. Examples: keratinized stratified squamous epithelium, ciliated stratified columnar epithelium.[FMA]</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium that consists of more than one layer of epithelial cells.[CARO]</obo:UBPROP_0000001>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Stratified_epithelium"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0682575"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Stratified_Epithelium"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/309044005"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010059</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000069</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000069</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0002074</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000069</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003069</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:45562</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000069</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001494</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0682575</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0004006</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001494</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">stratified epithelium</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">laminated epithelium</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000486</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">multilaminar epithelium</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000486"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium which consists of more than one layer of epithelial cells that may or may not be in contact with a basement membrane. Examples: keratinized stratified squamous epithelium, ciliated stratified columnar epithelium.[FMA]</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Stratified_epithelium"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:FMA</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000486"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium that consists of more than one layer of epithelial cells.[CARO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000069</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:source>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:45562</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000486"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0682575</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Stratified_Epithelium</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000486"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">laminated epithelium</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0002074</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000490 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000490">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium which consists of a single layer of epithelial cells. Examples: endothelium, mesothelium, glandular squamous epithelium.[FMA]</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">consider adding disjointness axiom between unilaminar and multilaminar - but note that this will render EHDAA2:0003244 (chorionic trophoblast) unsatisfiable</obo:IAO_0000116>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium that consists of a single layer of epithelial cells.[CARO]</obo:UBPROP_0000001>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0682574"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Simple_Epithelium"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/309043004"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010062</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000073</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000073</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0002073</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000073</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003073</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:45561</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000073</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001495</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0682574</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0004007</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001495</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">simple epithelium</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000490</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">unilaminar epithelium</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000490"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium which consists of a single layer of epithelial cells. Examples: endothelium, mesothelium, glandular squamous epithelium.[FMA]</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:45561</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000490"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Epithelium that consists of a single layer of epithelial cells.[CARO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0000073</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:MAH</oboInOwl:source>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:45561</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000490"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0682574</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Simple_Epithelium</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000490"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">simple epithelium</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:45561</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000916 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000916">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002417"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The subdivision of the vertebrate body between the thorax and pelvis. The ventral part of the abdomen contains the abdominal cavity and visceral organs. The dorsal part includes the abdominal section of the vertebral column.</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Vertebrate specific. In arthropods &apos;abdomen&apos; is the most distal section of the body which lies behind the thorax or cephalothorax. If need be we can introduce some grouping class</obo:IAO_0000116>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdominal</obo:UBPROP_0000007>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">celiac</obo:UBPROP_0000007>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Abdomen"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/302553009"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000020</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-0001</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000968</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35102</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100011</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9577</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:16</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000029</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000298</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A01.047</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000298</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rvVjgyZwpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:Abdomen</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdominopelvic region</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdominopelvis</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adult abdomen</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">belly</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">celiac region</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000916</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdomen</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000916"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The subdivision of the vertebrate body between the thorax and pelvis. The ventral part of the abdomen contains the abdominal cavity and visceral organs. The dorsal part includes the abdominal section of the vertebral column.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:cjm</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000916"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdominopelvic region</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9577</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000916"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdominopelvis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9577</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0000949 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0000949">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of the glands and parts of glands that produce endocrine secretions and help to integrate and control bodily metabolic activity.</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system containing glands which regulates bodily functions though the secretion of hormones.[AAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Multicellular organisms have complex endocrine systems, allowing responses to environmental stimuli, regulation of development, reproduction, and homeostasis. Nuclear receptors (NRs), a metazoan-specific family of ligand-activated transcription factors, play central roles in endocrine responses, as intermediates between signaling molecules and target genes. The NR family includes ligand-bound and orphan receptors, that is, receptors with no known ligand or for which there is no ligand Pocket. Understanding NR evolution has been further improved by comparison of several completed genomes, particularly those of deuterostomes and ecdysozoans. In contrast, evolution of NR ligands is still much debated. One hypothesis proposes that several independent gains and losses of ligand-binding ability in NRs occurred in protostomes and deuterostomes. A second hypothesis, pertaining to the NR3 subfamily (vertebrate steroid hormone receptors and estrogen related receptor), proposes that before the divergence of protostomes and deuterostomes, there was an ancestral steroid receptor (AncSR) that was ligand-activated and that orphan receptors secondarily lost the ability to bind a ligand. (...) Our analysis reveals that steroidogenesis has been independently elaborated in the 3 main bilaterian lineages (...).[well established][VHOG]</obo:UBPROP_0000003>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">endocrine</obo:UBPROP_0000007>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Endocrine_system"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0014136"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Endocrine_System"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/278876000"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010279</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-1301</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0002969</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0002224</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35306</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100128</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00005068</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9668</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:439</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000012</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A06</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001158</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0014136</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000098</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000158</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001158</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">endocrine glandular system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">endocrine system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">systema endocrinum</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0000949</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">endocrine system</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000949"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of the glands and parts of glands that produce endocrine secretions and help to integrate and control bodily metabolic activity.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Endocrine_system"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">NLM:endocrine+system</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000949"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system containing glands which regulates bodily functions though the secretion of hormones.[AAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010279</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:BJB</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000949"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Multicellular organisms have complex endocrine systems, allowing responses to environmental stimuli, regulation of development, reproduction, and homeostasis. Nuclear receptors (NRs), a metazoan-specific family of ligand-activated transcription factors, play central roles in endocrine responses, as intermediates between signaling molecules and target genes. The NR family includes ligand-bound and orphan receptors, that is, receptors with no known ligand or for which there is no ligand Pocket. Understanding NR evolution has been further improved by comparison of several completed genomes, particularly those of deuterostomes and ecdysozoans. In contrast, evolution of NR ligands is still much debated. One hypothesis proposes that several independent gains and losses of ligand-binding ability in NRs occurred in protostomes and deuterostomes. A second hypothesis, pertaining to the NR3 subfamily (vertebrate steroid hormone receptors and estrogen related receptor), proposes that before the divergence of protostomes and deuterostomes, there was an ancestral steroid receptor (AncSR) that was ligand-activated and that orphan receptors secondarily lost the ability to bind a ligand. (...) Our analysis reveals that steroidogenesis has been independently elaborated in the 3 main bilaterian lineages (...).[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000098</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">DOI:10.1073/pnas.0812138106 Markov GV, Tavares R, Dauphin-Villemant C, Demeneix BA, Baker ME, Laudet V, Independent elaboration of steroid hormone signaling pathways in metazoans. PNAS (2009)</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000949"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0014136</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Endocrine_System</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000949"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">endocrine glandular system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0002224</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0001009 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0001009">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organ system that passes nutrients (such as amino acids and electrolytes), gases, hormones, blood cells, etc. to and from cells in the body to help fight diseases and help stabilize body temperature and pH to maintain homeostasis[WP].</obo:IAO_0000115>
+        <obo:IAO_0000232 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">the cardiovascular system and the lymphatic system are parts of the circulatory system</obo:IAO_0000232>
+        <obo:RO_0002174 rdf:resource="http://purl.obolibrary.org/obo/NCBITaxon_6231"/>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system of ion binding, a pumping mechanism, and an efficient vascular system; consisting of the blood, heart, and blood and lymph vessels, respectively.[AAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">We should divest ourselves of the view that earlier vertebrate groups were &apos;on their way&apos; to becoming mammals, as clearly they were not such visionaries. Neither were their systems &apos;imperfect&apos; as earlier anatomists thought. Instead, their circulatory systems served them well to address the ecological demands arising from their lifestyles.[well established][VHOG]</obo:UBPROP_0000003>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Circulatory_system"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000959</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-2103</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00005057</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rvVjzG5wpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001248</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">systema cardiovasculare</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0001009</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">circulatory system</rdfs:label>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/2/29/Circulatory_System_en.svg"/>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/thumb/2/29/Circulatory_System_en.svg/200px-Circulatory_System_en.svg.png"/>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001009"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organ system that passes nutrients (such as amino acids and electrolytes), gases, hormones, blood cells, etc. to and from cells in the body to help fight diseases and help stabilize body temperature and pH to maintain homeostasis[WP].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Circulatory_system"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001009"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/RO_0002174"/>
+        <owl:annotatedTarget rdf:resource="http://purl.obolibrary.org/obo/NCBITaxon_6231"/>
+        <oboInOwl:notes rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt coelomocyte currently classified as circulating cell</oboInOwl:notes>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001009"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system of ion binding, a pumping mechanism, and an efficient vascular system; consisting of the blood, heart, and blood and lymph vessels, respectively.[AAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000959</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:BJB</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001009"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">We should divest ourselves of the view that earlier vertebrate groups were &apos;on their way&apos; to becoming mammals, as clearly they were not such visionaries. Neither were their systems &apos;imperfect&apos; as earlier anatomists thought. Instead, their circulatory systems served them well to address the ecological demands arising from their lifestyles.[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001248</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ISBN:978-0072528305 Kardong KV, Vertebrates: Comparative Anatomy, Function, Evolution (2006) p.493</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001009"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">systema cardiovasculare</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Circulatory_system"/>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#LATIN"/>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0001016 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0001016">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The nervous system is an organ system containing predominantly neuron and glial cells. In bilaterally symmetrical organism, it is arranged in a network of tree-like structures connected to a central body. The main functions of the nervous system are to regulate and control body functions, and to receive sensory input, process this information, and generate behavior [CUMBO].</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">A regulatory system of the body that consists of neurons and neuroglial cells. The nervous system is divided into two parts, the central nervous system (CNS) and the peripheral nervous system (PNS). (Source: BioGlossary, www.Biology-Text.com)[TAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system consisting of nerve bodies and nerve fibers which regulate the response of the body to external and internal stimuli.[AAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Nervous systems evolved in the ancestor of Eumetazoa.[well established][VHOG]</obo:UBPROP_0000003>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">nervous</obo:UBPROP_0000007>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">neural</obo:UBPROP_0000007>
+        <oboInOwl:hasDbXref rdf:resource="http://braininfo.rprc.washington.edu/centraldirectory.aspx?ID=3236"/>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Nervous_system"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0027763"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Nervous_System"/>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_844"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/278196006"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000324</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000079</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001484</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-1313</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000802</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0001246</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA:826</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:16469</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100162</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00005093</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:7157</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:466</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000016</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000026</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A08</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000026</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rvViT_pwpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0000396</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0027763</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000402</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0005735</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000177</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0000396</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">neurological system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasNarrowSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">nerve net</oboInOwl:hasNarrowSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">systema nervosum</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0001016</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#cumbo"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">nervous system</rdfs:label>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/b/b2/TE-Nervous_system_diagram.svg"/>
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+    <owl:Axiom>
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+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
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+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Nervous_system"/>
+        <oboInOwl:hasDbXref rdf:resource="http://uri.neuinfo.org/nif/nifstd/birnlex_844"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ISBN:0-14-051288-8</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ISBN:3110148986</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">NLM:nervous+system</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WB:rynl</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFIN:curator</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001016"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">A regulatory system of the body that consists of neurons and neuroglial cells. The nervous system is divided into two parts, the central nervous system (CNS) and the peripheral nervous system (PNS). (Source: BioGlossary, www.Biology-Text.com)[TAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-08-14</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0000396</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFIN:curator</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001016"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system consisting of nerve bodies and nerve fibers which regulate the response of the body to external and internal stimuli.[AAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000324</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:BJB</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001016"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Nervous systems evolved in the ancestor of Eumetazoa.[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000402</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ISBN:978-0198566694 Schmidt-Rhaesa A, The evolution of organ systems (2007) p.117</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001016"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0027763</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Nervous_System</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001016"/>
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+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">neurological system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO:0050877</oboInOwl:hasDbXref>
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+    <owl:Axiom>
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+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasNarrowSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">nerve net</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Nerve_net"/>
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+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">systema nervosum</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Nervous_system"/>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#LATIN"/>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0001235 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0001235">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000064"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">the thick outer layer of the adrenal gland that produces and secretes steroid hormones such as corticosterone, estrone and aldosterone</obo:IAO_0000115>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">All craniates have groups of cells homologous to the mammalian adrenocortical and chromaffin tissues (medulla), but they are scattered in and near the kidneys in fishes. (...) The cortical and chromaffin tissues come together to form adrenal glands in tetrapods.[well established][VHOG]</obo:UBPROP_0000003>
+        <obo:UBPROP_0000008 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Kardong states that mammals are the first to have distinct cortext and medulla, but this contradicts XAO</obo:UBPROP_0000008>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_cortex"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0001613"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Adrenal_Cortex"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/362584002"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0011009</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000045</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-0015</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000237</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:18427</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100136</oboInOwl:hasDbXref>
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+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:447</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000118</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000494</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A06.407.071.140</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0001613</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001481</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000165</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal gland cortex</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cortex (glandula suprarenalis)</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cortex of adrenal gland</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cortex of suprarenal gland</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">suprarenal cortex</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cortex glandulae suprarenalis</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">suprarenal</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0001235</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal cortex</rdfs:label>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/5/5c/Gray1185.png"/>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/thumb/5/5c/Gray1185.png/200px-Gray1185.png"/>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001235"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">the thick outer layer of the adrenal gland that produces and secretes steroid hormones such as corticosterone, estrone and aldosterone</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MP:0008288,ISBN:0-683-40008-8,MGI:llw2</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001235"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">All craniates have groups of cells homologous to the mammalian adrenocortical and chromaffin tissues (medulla), but they are scattered in and near the kidneys in fishes. (...) The cortical and chromaffin tissues come together to form adrenal glands in tetrapods.[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001481</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ISBN:978-0030223693 Liem KF, Bemis WE, Walker WF, Grande L, Functional Anatomy of the Vertebrates: An Evolutionary Perspective (2001) p.518 and Figure 15-9</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001235"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0001613</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Adrenal_Cortex</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001235"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cortex glandulae suprarenalis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_cortex"/>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#LATIN"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001235"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">cortex glandulae suprarenalis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000045</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0001434 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0001434">
+        <owl:equivalentClass>
+            <owl:Class>
+                <owl:intersectionOf rdf:parseType="Collection">
+                    <rdf:Description rdf:about="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+                    <owl:Restriction>
+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004288"/>
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+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004770"/>
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+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002204"/>
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+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004288"/>
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+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
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+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system consisting of multiple elements and tissues that provides physical support.[TAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that is a multi-element, multi-tissue anatomical cluster that consists of the skeleton and the articular system.[VSAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">System that provides physical support to the organism.[AAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">By taking a holistic approach, integration of the evidence from molecular and developmental features of model organisms, the phylogenetic distribution in the &apos;new animal phylogeny&apos; and the earliest fossilized remains of mineralized animal skeletons suggests independent origins of the skeleton at the phylum level.[debated][VHOG]</obo:UBPROP_0000003>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">skeletal</obo:UBPROP_0000007>
+        <obo:UBPROP_0000012 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO defines skeletal system very generically: The skeleton is the bony framework of the body in vertebrates (endoskeleton) or the hard outer envelope of insects (exoskeleton or dermoskeleton) GO:0001501; however, all annotations are to vertebrates</obo:UBPROP_0000012>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0037253"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Skeletal_System"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000566</oboInOwl:hasDbXref>
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+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-1320</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000806</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003168</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35773</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:23881</oboInOwl:hasDbXref>
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+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0000434</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">skeleton system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasNarrowSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all bones and joints</oboInOwl:hasNarrowSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Skelettsystem</oboInOwl:hasRelatedSynonym>
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+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">skeletal system</rdfs:label>
+        <rdfs:seeAlso rdf:datatype="http://www.w3.org/2001/XMLSchema#anyURI">https://github.com/obophenotype/uberon/wiki/The-skeletal-system</rdfs:seeAlso>
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+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
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+        <oboInOwl:hasDbXref rdf:resource="http://dx.plos.org/10.1371/journal.pone.0051070"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO_REF:0000034</oboInOwl:hasDbXref>
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+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system consisting of multiple elements and tissues that provides physical support.[TAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-08-14</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0000434</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:wd</oboInOwl:source>
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+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
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+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-08-14</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000027</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO_REF:0000034, http://dx.plos.org/10.1371/journal.pone.0051070</oboInOwl:source>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">PSPUB:0000170</oboInOwl:source>
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+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">System that provides physical support to the organism.[AAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0000566</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:LAP</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
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+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001254</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">DOI:10.1159/000324245 Donoghue PCJ, Sansom IJ, Origin and early evolution of vertebrate skeletonization. Microscopy research and technique (2002)</oboInOwl:source>
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+        <oboInOwl:external_ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO</oboInOwl:external_ontology>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0037253</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Skeletal_System</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasNarrowSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all bones and joints</owl:annotatedTarget>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#SENSU"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Skelettsystem</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001486</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0002100 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0002100">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013701"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013702"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision which is the part of the body posterior to the cervical region (or head, when cervical region not present) and anterior to the caudal region. Includes the sacrum when present.</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision that is the part of the body posterior to the head and anterior to the tail.[AAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision which is the part of the body posterior to the head and anterior to the tail.[TAO]</obo:UBPROP_0000001>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Torso"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0460005"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Trunk"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/262225004"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010339</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000116</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001493</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-1071</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000966</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:31857</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:7181</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000004</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000296</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000296</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rvVkJjpwpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001115</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0460005</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000054</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003025</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001115</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:Trunk</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">thoracolumbar region</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">torso</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">trunk region</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Rumpf</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0002100</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">trunk</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision which is the part of the body posterior to the cervical region (or head, when cervical region not present) and anterior to the caudal region. Includes the sacrum when present.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Torso"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001115</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERONREF:0000006</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision that is the part of the body posterior to the head and anterior to the tail.[AAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010339</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:BJB</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision which is the part of the body posterior to the head and anterior to the tail.[TAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-08-14</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001115</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFIN:curator</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0460005</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Trunk</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">trunk region</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000054</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Rumpf</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001493</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0002204 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0002204">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of the muscular and skeletal systems.</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that provides locomotion and physical support to the organism.[AAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">There are more than 50,000 extant vertebrate species, representing over 500 million years of evolution. During that time, the vertebrate musculoskeletal systems have adapted to aquatic, terrestrial, fossorial, and arboreal lifestyles, while simultaneously retaining functionally integrated axial and appendicular skeletal systems.[well established][VHOG]</obo:UBPROP_0000003>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculoskeletal</obo:UBPROP_0000007>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Musculoskeletal_system"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0026860"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Musculoskeletal_System"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/278858007"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010546</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-1311</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:32714</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100139</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:7482</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:98</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0002418</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A02</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rQRpVNgAKEdyHxgDggVfs8g</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0026860</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001275</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000031</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000168</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculo-skeletal system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0002204</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculoskeletal system</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002204"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of the muscular and skeletal systems.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000031</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:curator</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002204"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that provides locomotion and physical support to the organism.[AAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010546</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:EJS</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002204"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">There are more than 50,000 extant vertebrate species, representing over 500 million years of evolution. During that time, the vertebrate musculoskeletal systems have adapted to aquatic, terrestrial, fossorial, and arboreal lifestyles, while simultaneously retaining functionally integrated axial and appendicular skeletal systems.[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001275</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">DOI:10.1002/jez.b.21246 Shearman RM, Burke AC, The lateral somitic frontier in ontogeny and phylogeny. Journal of Experimental Zoology (2009)</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002204"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0026860</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Musculoskeletal_System</oboInOwl:source>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0002343 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0002343">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000480"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000383"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000916"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Set of all muscles in abdomen.</obo:IAO_0000115>
+        <obo:IAO_0000232 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">distinction between abdomen muscle and abdomen musculature</obo:IAO_0000232>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:71294</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:86917</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0001327</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0001327</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdominal musculature</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscle group of abdomen</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscles of abdomen</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculature of abdomen</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculature of abdominal wall</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of muscles of abdomen</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0002343</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#vertebrate_core"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdomen musculature</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002343"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Set of all muscles in abdomen.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OBOL:automatic</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002343"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscle group of abdomen</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:71294</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002343"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">muscles of abdomen</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:71294</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002343"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">musculature of abdominal wall</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:86917</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002343"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of muscles of abdomen</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:71294</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0002369 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0002369">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000916"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000949"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002417"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Either of a pair of complex endocrine organs near the anterior medial border of the kidney consisting of a mesodermal cortex that produces glucocorticoid, mineralocorticoid, and androgenic hormones and an ectodermal medulla that produces epinephrine and norepinephrine[BTO].</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure which is found on the surface of the kidney and secretes various hormones including epinephrine, norephinephrine, aldosterone, corticosterone, and cortisol.[AAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">All craniates have groups of cells homologous to the mammalian adrenocortical and chromaffin tissues, but they are scattered in and near the kidneys in fishes. (...) The cortical and chromaffin tissues come together to form adrenal glands in tetrapods.[well established][VHOG]</obo:UBPROP_0000003>
+        <obo:UBPROP_0000008 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The origin of the adrenal gland is still controversial. It is thought to share the same origin as the kidney and gonads, derived from coelomic epithelium of the urogenital ridge and/or the underlying mesenchyme (Keegan and Hammer, 2002; Morohashi, 1997). We follow Kardong and state homology at the level of the cortex and medulla rather than gland as a whole</obo:UBPROP_0000008>
+        <obo:UBPROP_0000009 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">suprarenal cortex manufactures corticosteroids; suprarenal medulla manufactures epinephrine and norepinephrine; suprarenal medulla receives preganglionic sympathetic innervation from the greater thoracic splanchnic n.</obo:UBPROP_0000009>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_gland"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0001625"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Adrenal_Gland"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/181127006"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010551</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-0016</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000238</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:18426</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100135</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:9604</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:446</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000116</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000071</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A06.407.071</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000071</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rvXYiz5wpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0001625</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001141</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000164</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:AdrenalGland</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">glandula adrenalis</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">glandula suprarenalis</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal capsule</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal medulla cell</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">atrabiliary capsule</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epinephric gland</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epinephros</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">glandula suprarenalis</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">interrenal gland</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">suprarenal capsule</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">suprarenal gland</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0002369</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#organ_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal gland</rdfs:label>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/9/9d/Illu_endocrine_system_New.png"/>
+        <foaf:depicted_by rdf:resource="http://upload.wikimedia.org/wikipedia/commons/thumb/c/c6/Illu_endocrine_system.jpg/200px-Illu_endocrine_system.jpg"/>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Either of a pair of complex endocrine organs near the anterior medial border of the kidney consisting of a mesodermal cortex that produces glucocorticoid, mineralocorticoid, and androgenic hormones and an ectodermal medulla that produces epinephrine and norepinephrine[BTO].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_gland"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure which is found on the surface of the kidney and secretes various hormones including epinephrine, norephinephrine, aldosterone, corticosterone, and cortisol.[AAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-06-20</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010551</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:BJB</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">All craniates have groups of cells homologous to the mammalian adrenocortical and chromaffin tissues, but they are scattered in and near the kidneys in fishes. (...) The cortical and chromaffin tissues come together to form adrenal glands in tetrapods.[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0001141</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ISBN:978-0030223693 Liem KF, Bemis WE, Walker WF, Grande L, Functional Anatomy of the Vertebrates: An Evolutionary Perspective (2001) p.518 and Figure 15-9</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0001625</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Adrenal_Gland</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">glandula adrenalis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_gland"/>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#LATIN"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">glandula suprarenalis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_gland"/>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#LATIN"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal capsule</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">atrabiliary capsule</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epinephric gland</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_gland"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epinephros</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">glandula suprarenalis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">interrenal gland</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO:0030325</oboInOwl:hasDbXref>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#HOMOLOGY"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">suprarenal capsule</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000047</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">suprarenal gland</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Adrenal_gland"/>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0002416 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0002416">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0007376"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">An organ system that protects the body from damage, forming the bounding layer of the organism and comprising the outer epithelial layer and any associated adnexa. In vertebrates, the integumental system consists of the epidermis, dermis plus associated glands and adnexa such as hair and scales. In invertebrates, the integumental system may include cuticle.</obo:IAO_0000115>
+        <obo:UBPROP_0000003 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">(...) the integument of many tetrapods is reinforced by a morphologically and structurally diverse assemblage of skeletal elements. These elements are widely understood to be derivatives of the once all-encompassing dermal skeleton of stem-gnathostomes (...).[well established][VHOG]</obo:UBPROP_0000003>
+        <oboInOwl:hasAlternativeId rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0007029</oboInOwl:hasAlternativeId>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Integumentary_system"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0037267"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Integumentary_System"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/361692004"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000154</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000118</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CALOHA:TS-1299</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">CARO:0002001</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EFO:0000807</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0000836</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2_RETIRED:0003154</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA:6520</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:17524</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EV:0100151</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00004969</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:72979</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000421</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000014</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000033</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A17</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000033</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TADS:0000108</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0037267</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000403</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0000176</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:Surface</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumentary system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body surface</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">dermal system</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">external covering of organism</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumentum commune</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organism surface</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">surface</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0002416</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#grouping_class"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumental system</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">An organ system that protects the body from damage, forming the bounding layer of the organism and comprising the outer epithelial layer and any associated adnexa. In vertebrates, the integumental system consists of the epidermis, dermis plus associated glands and adnexa such as hair and scales. In invertebrates, the integumental system may include cuticle.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://orcid.org/0000-0002-6601-2165"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000003"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">(...) the integument of many tetrapods is reinforced by a morphologically and structurally diverse assemblage of skeletal elements. These elements are widely understood to be derivatives of the once all-encompassing dermal skeleton of stem-gnathostomes (...).[well established][VHOG]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-09-17</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000403</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG</oboInOwl:ontology>
+        <oboInOwl:source rdf:resource="http://bgee.unil.ch/"/>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">DOI:10.1111/j.1469-7580.2008.01043.x Vickaryous MK, Sire JY, The integumentary skeleton of tetrapods: origin, evolution, and development. J Anat (2009)</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0037267</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Integumentary_System</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumentary system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:72979</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">dermal system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000033</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumentum commune</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Integumentary_system"/>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#LATIN"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">organism surface</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003154</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">surface</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BILA:0000118</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0002417 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0002417">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002100"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013701"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The abdominal segment of the torso.</obo:IAO_0000115>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Lumbar"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/362875007"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35104</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:259211</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000021</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdomen/pelvis/perineum</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">lower body</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">lumbar region</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0002417</oboInOwl:id>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdominal segment of trunk</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002417"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The abdominal segment of the torso.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Lumbar"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002417"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">abdomen/pelvis/perineum</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000021</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002417"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">lower body</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000021</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002417"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">lumbar region</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="https://orcid.org/0000-0002-6601-2165"/>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0003102 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0003102">
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000051"/>
+                <owl:someValuesFrom>
+                    <owl:Restriction>
+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0007376"/>
+                    </owl:Restriction>
+                </owl:someValuesFrom>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure that overlaps the outer epithelial layer and is adjacent to the space surrounding the organism.</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision which is the collection of anatomical structures on the body surface.[ZFA]</obo:UBPROP_0000001>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Surface_structure"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010337</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO_RETIRED:0000010</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2_RETIRED:0003010</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0000292</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000001</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0003028</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFA:0000292</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:SurfaceRegion</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">anatomical surface feature</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">surface feature</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">surface region</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0003102</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#upper_level"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">surface structure</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003102"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical structure that overlaps the outer epithelial layer and is adjacent to the space surrounding the organism.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://orcid.org/0000-0002-6601-2165"/>
+        <oboInOwl:hasDbXref rdf:resource="http://purl.obolibrary.org/obo/uberon/tracker/24"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003102"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Organism subdivision which is the collection of anatomical structures on the body surface.[ZFA]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-08-14</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO:0000292</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ZFIN:curator</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003102"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">anatomical surface feature</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0003010</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0003297 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0003297">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002416"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">A gland that is part of a integumental system [Automatically generated definition].</obo:IAO_0000115>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0000837</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA:6522</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:17758</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000144</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VHOG:0000654</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumental gland</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumental system gland</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumentary gland</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0003297</oboInOwl:id>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">gland of integumental system</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003297"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">A gland that is part of a integumental system [Automatically generated definition].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OBOL:automatic</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003297"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumental gland</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:cjm</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003297"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumental system gland</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:cjm</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003297"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">integumentary gland</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:cjm</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0004288 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0004288">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000480"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical cluster that consists of all the skeletal elements (eg., bone, cartilage, and teeth) of the body.</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical cluster that consists of all the skeletal elements (eg., bone, cartilage, and teeth) of the body.[VSAO]</obo:UBPROP_0000001>
+        <obo:UBPROP_0000007 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">skeletal</obo:UBPROP_0000007>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Skeleton"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/361378004"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000168</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA2:0001843</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EHDAA:5047</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:17213</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:23875</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:177</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0003006</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MAT:0000032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MESH:A02.835</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MIAA:0000032</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">OpenCyc:Mx4rvVi1rpwpEbGdrcN5Y29ycA</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000026</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">XAO:0004053</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">galen:Skeleton</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all bones</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of bones of body</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0004288</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#efo_slim"/>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#uberon_slim"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">skeleton</rdfs:label>
+        <rdfs:seeAlso rdf:datatype="http://www.w3.org/2001/XMLSchema#anyURI">https://github.com/obophenotype/uberon/wiki/The-skeletal-system</rdfs:seeAlso>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004288"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical cluster that consists of all the skeletal elements (eg., bone, cartilage, and teeth) of the body.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000026</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004288"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical cluster that consists of all the skeletal elements (eg., bone, cartilage, and teeth) of the body.[VSAO]</owl:annotatedTarget>
+        <oboInOwl:date_retrieved rdf:datatype="http://www.w3.org/2001/XMLSchema#string">2012-08-14</oboInOwl:date_retrieved>
+        <oboInOwl:external_class rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000026</oboInOwl:external_class>
+        <oboInOwl:ontology rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO</oboInOwl:ontology>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO_REF:0000034, http://dx.plos.org/10.1371/journal.pone.0051070</oboInOwl:source>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0004770 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0004770">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000467"/>
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000480"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001434"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of all the joints of the body.</obo:IAO_0000115>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/361827000"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:35150</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:23878</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0003007</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000181</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">joint system</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all joints of body</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all joints</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of joints of body</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0004770</oboInOwl:id>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">articular system</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004770"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Anatomical system that consists of all the joints of the body.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">VSAO:0000181</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004770"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">joint system</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:23878</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004770"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of all joints of body</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:23878</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0004770"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">set of joints of body</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:23878</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0005769 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0005769">
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">An acellular membrane that is part of the epithelium, lies adjacent to the epithelial cells, and is the fusion of the the basal lamina and the reticular lamina.</obo:IAO_0000115>
+        <obo:IAO_0000232 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">this class represents a continuous sheet of basement membrane which can underlie multiple epithelial cells over large regions. In contrast, the GO class &apos;basal membrane&apos; represents a portion of substance on the scale of a single cell.</obo:IAO_0000232>
+        <obo:RO_0002175 rdf:resource="http://purl.obolibrary.org/obo/NCBITaxon_80999"/>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Basement_membrane"/>
+        <oboInOwl:hasDbXref rdf:resource="http://linkedlifedata.com/resource/umls/id/C0004799"/>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Basement_Membrane"/>
+        <oboInOwl:hasDbXref rdf:resource="http://www.snomedbrowser.com/Codes/Details/68989006"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AAO:0010596</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:63872</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GAID:915</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0004799</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">basement membrane of connective tissue</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">membrana basalis</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">basement membrane</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0005769</oboInOwl:id>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">basement membrane of epithelium</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0005769"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">An acellular membrane that is part of the epithelium, lies adjacent to the epithelial cells, and is the fusion of the the basal lamina and the reticular lamina.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Basement_membrane"/>
+        <oboInOwl:hasDbXref rdf:resource="http://orcid.org/0000-0002-6601-2165"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0005769"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/RO_0002175"/>
+        <owl:annotatedTarget rdf:resource="http://purl.obolibrary.org/obo/NCBITaxon_80999"/>
+        <oboInOwl:source rdf:resource="http://palaeos.com/metazoa/porifera/homoscleromorpha.html"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0005769"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasDbXref"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UMLS:C0004799</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Basement_Membrane</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0005769"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">basement membrane of connective tissue</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:63872</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0005769"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">membrana basalis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Basement_membrane"/>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#LATIN"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0005769"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">basement membrane</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:63872</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0007376 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0007376">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000483"/>
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0003102"/>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The epidermis is the entire outer epithelial layer of an animal, it may be a single layer that produces an extracellular material (e.g. the cuticle of arthropods) or a complex stratified squamous epithelium, as in the case of many vertebrate species[GO].</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">this grouping class exists primarily to align with GO - see GO:0008544.</obo:IAO_0000116>
+        <oboInOwl:hasDbXref rdf:resource="http://en.wikipedia.org/wiki/Epidermis_(zoology)"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BSA:0000073</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000313</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FBbt:00004993</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">HAO:0000298</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TADS:0000109</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0005733</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epidermis</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">outer epidermal layer</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">outer epithelial layer</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">hypoderm</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">hypodermis</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0007376</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#grouping_class"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">outer epithelium</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0007376"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The epidermis is the entire outer epithelial layer of an animal, it may be a single layer that produces an extracellular material (e.g. the cuticle of arthropods) or a complex stratified squamous epithelium, as in the case of many vertebrate species[GO].</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO:0008544</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0007376"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">epidermis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO:0008544</oboInOwl:hasDbXref>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#HUMAN_PREFERRED"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0007376"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">hypoderm</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0000313</oboInOwl:hasDbXref>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#INCONSISTENT"/>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0007376"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">hypodermis</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">GO:0008544</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">WBbt:0005733</oboInOwl:hasDbXref>
+        <oboInOwl:hasSynonymType rdf:resource="http://purl.obolibrary.org/obo/uberon/core#INCONSISTENT"/>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0010133 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0010133">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000062"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000949"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0001016"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">any of the organized aggregations of cells that function as secretory or excretory organs and that release hormones in response to neural stimuli</obo:IAO_0000115>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MA:0000720</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0010133</oboInOwl:id>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">neuroendocrine gland</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0010133"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">any of the organized aggregations of cells that function as secretory or excretory organs and that release hormones in response to neural stimuli</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">MP:0000631,ISBN:0-683-40008-8</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0013701 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0013701">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">A principle subdivision of an organism that includes all structures along the primary axis, typically the anterior-posterior axis, from head to tail, including structures of the body proper where present (for example, ribs), but excluding appendages.</obo:IAO_0000115>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0013701</oboInOwl:id>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">main body axis</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013701"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">A principle subdivision of an organism that includes all structures along the primary axis, typically the anterior-posterior axis, from head to tail, including structures of the body proper where present (for example, ribs), but excluding appendages.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:resource="https://orcid.org/0000-0002-6601-2165"/>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0013702 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0013702">
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000475"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013701"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The region of the organism associated with the visceral organs.</obo:IAO_0000115>
+        <obo:UBPROP_0000001 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Cardinal body part, which consists of a maximal set of diverse subclasses of organ and organ part spatially associated with the vertebral column and ribcage. Examples: There is only one body proper[FMA:231424].</obo:UBPROP_0000001>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000103</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001489</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">EMAPA:36031</oboInOwl:hasDbXref>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:231424</oboInOwl:hasDbXref>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:hasRelatedSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">whole body</oboInOwl:hasRelatedSynonym>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0013702</oboInOwl:id>
+        <oboInOwl:inSubset rdf:resource="http://purl.obolibrary.org/obo/uberon/core#non_informative"/>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body proper</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013702"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">The region of the organism associated with the visceral organs.</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000103</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013702"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/UBPROP_0000001"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Cardinal body part, which consists of a maximal set of diverse subclasses of organ and organ part spatially associated with the vertebral column and ribcage. Examples: There is only one body proper[FMA:231424].</owl:annotatedTarget>
+        <oboInOwl:source rdf:datatype="http://www.w3.org/2001/XMLSchema#string">FMA:231424</oboInOwl:source>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013702"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">body</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">AEO:0000103</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0013702"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasRelatedSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">whole body</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">BTO:0001489</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    
+
+
+    <!-- http://purl.obolibrary.org/obo/UBERON_0018303 -->
+
+    <owl:Class rdf:about="http://purl.obolibrary.org/obo/UBERON_0018303">
+        <owl:equivalentClass>
+            <owl:Class>
+                <owl:intersectionOf rdf:parseType="Collection">
+                    <rdf:Description rdf:about="http://purl.obolibrary.org/obo/UBERON_0000479"/>
+                    <owl:Restriction>
+                        <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                        <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+                    </owl:Restriction>
+                </owl:intersectionOf>
+            </owl:Class>
+        </owl:equivalentClass>
+        <rdfs:subClassOf rdf:resource="http://purl.obolibrary.org/obo/UBERON_0000479"/>
+        <rdfs:subClassOf>
+            <owl:Restriction>
+                <owl:onProperty rdf:resource="http://purl.obolibrary.org/obo/BFO_0000050"/>
+                <owl:someValuesFrom rdf:resource="http://purl.obolibrary.org/obo/UBERON_0002369"/>
+            </owl:Restriction>
+        </rdfs:subClassOf>
+        <obo:IAO_0000115 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Tissue that is part of some adrenal gland</obo:IAO_0000115>
+        <obo:IAO_0000116 rdf:datatype="http://www.w3.org/2001/XMLSchema#string">TODO - describe typology of adrenal gland tissue used in this ontology</obo:IAO_0000116>
+        <oboInOwl:hasDbXref rdf:resource="http://ncicb.nci.nih.gov/xml/owl/EVS/Adrenal_Gland_Tissue"/>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERONTEMP:a7bf197b-0db2-467e-824e-be45b39e2672</oboInOwl:hasDbXref>
+        <oboInOwl:hasExactSynonym rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal gland tissue</oboInOwl:hasExactSynonym>
+        <oboInOwl:hasOBONamespace rdf:datatype="http://www.w3.org/2001/XMLSchema#string">uberon</oboInOwl:hasOBONamespace>
+        <oboInOwl:id rdf:datatype="http://www.w3.org/2001/XMLSchema#string">UBERON:0018303</oboInOwl:id>
+        <rdfs:label rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal tissue</rdfs:label>
+    </owl:Class>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0018303"/>
+        <owl:annotatedProperty rdf:resource="http://purl.obolibrary.org/obo/IAO_0000115"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">Tissue that is part of some adrenal gland</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">PHENOSCAPE:Alex</oboInOwl:hasDbXref>
+    </owl:Axiom>
+    <owl:Axiom>
+        <owl:annotatedSource rdf:resource="http://purl.obolibrary.org/obo/UBERON_0018303"/>
+        <owl:annotatedProperty rdf:resource="http://www.geneontology.org/formats/oboInOwl#hasExactSynonym"/>
+        <owl:annotatedTarget rdf:datatype="http://www.w3.org/2001/XMLSchema#string">adrenal gland tissue</owl:annotatedTarget>
+        <oboInOwl:hasDbXref rdf:datatype="http://www.w3.org/2001/XMLSchema#string">ncithesaurus:Adrenal_Gland_Tissue</oboInOwl:hasDbXref>
+    </owl:Axiom>
+</rdf:RDF>
+
+
+
+<!-- Generated by the OWL API (version 4.5.6) https://github.com/owlcs/owlapi -->
+
diff --git a/robot-command/src/main/java/org/obolibrary/robot/CollapseCommand.java b/robot-command/src/main/java/org/obolibrary/robot/CollapseCommand.java
new file mode 100644
index 00000000..e53befdd
--- /dev/null
+++ b/robot-command/src/main/java/org/obolibrary/robot/CollapseCommand.java
@@ -0,0 +1,124 @@
+package org.obolibrary.robot;
+
+import java.util.Set;
+import org.apache.commons.cli.CommandLine;
+import org.apache.commons.cli.Options;
+import org.semanticweb.owlapi.model.IRI;
+import org.semanticweb.owlapi.model.OWLOntology;
+
+/** */
+public class CollapseCommand implements Command {
+
+  /** Namespace for error messages. */
+  private static final String NS = "minimize#";
+
+  /** Error message when --threshold is not an integer. */
+  private static final String thresholdError =
+      NS + "THRESHOLD ERROR threshold ('%s') must be a valid integer.";
+
+  /** Error message when --threshold is not an integer. */
+  private static final String thresholdValueError =
+      NS + "THRESHOLD VALUE ERROR threshold ('%d') must be 2 or greater.";
+
+  /** Store the command-line options for the command. */
+  private Options options;
+
+  /** Initialize the command. */
+  public CollapseCommand() {
+    Options o = CommandLineHelper.getCommonOptions();
+    o.addOption("i", "input", true, "collapse ontology from a file");
+    o.addOption("I", "input-iri", true, "collapse ontology from an IRI");
+    o.addOption("t", "threshold", true, "threshold to collapse below");
+    o.addOption("r", "precious", true, "CURIE or IRI of a class to keep");
+    o.addOption("R", "precious-terms", true, "set of CURIEs or IRIs of classes to keep");
+    o.addOption("o", "output", true, "save minimized ontology to a file");
+    options = o;
+  }
+
+  /**
+   * Name of the command.
+   *
+   * @return name
+   */
+  public String getName() {
+    return "collapse";
+  }
+
+  /**
+   * Brief description of the command.
+   *
+   * @return description
+   */
+  public String getDescription() {
+    return "minimize an ontology based on a threshold";
+  }
+
+  /**
+   * Command-line usage for the command.
+   *
+   * @return usage
+   */
+  public String getUsage() {
+    return "robot collapse --input <file> " + "--threshold <int> " + "--output <file>";
+  }
+
+  /**
+   * Command-line options for the command.
+   *
+   * @return options
+   */
+  public Options getOptions() {
+    return options;
+  }
+
+  /**
+   * Handle the command-line and file operations for the ReasonOperation.
+   *
+   * @param args strings to use as arguments
+   */
+  public void main(String[] args) {
+    try {
+      execute(null, args);
+    } catch (Exception e) {
+      CommandLineHelper.handleException(e);
+    }
+  }
+
+  public CommandState execute(CommandState state, String[] args) throws Exception {
+    CommandLine line = CommandLineHelper.getCommandLine(getUsage(), getOptions(), args);
+    if (line == null) {
+      return null;
+    }
+
+    if (state == null) {
+      state = new CommandState();
+    }
+
+    IOHelper ioHelper = CommandLineHelper.getIOHelper(line);
+    state = CommandLineHelper.updateInputOntology(ioHelper, state, line);
+    OWLOntology ontology = state.getOntology();
+
+    String thresholdString = CommandLineHelper.getDefaultValue(line, "threshold", "2");
+    int threshold;
+    try {
+      threshold = Integer.parseInt(thresholdString);
+    } catch (NumberFormatException e) {
+      throw new IllegalArgumentException(String.format(thresholdError, thresholdString));
+    }
+
+    // If threshold is not 2 or greater
+    // Nothing will be done
+    if (threshold < 2) {
+      throw new Exception(String.format(thresholdValueError, threshold));
+    }
+
+    Set<IRI> precious = CommandLineHelper.getTerms(ioHelper, line, "precious", "precious-terms");
+
+    OntologyHelper.collapseOntology(ontology, threshold, precious, true);
+
+    // Save the changed ontology and return the state
+    CommandLineHelper.maybeSaveOutput(line, ontology);
+    state.setOntology(ontology);
+    return state;
+  }
+}
diff --git a/robot-command/src/main/java/org/obolibrary/robot/CommandLineInterface.java b/robot-command/src/main/java/org/obolibrary/robot/CommandLineInterface.java
index b9712dec..48cc8d06 100644
--- a/robot-command/src/main/java/org/obolibrary/robot/CommandLineInterface.java
+++ b/robot-command/src/main/java/org/obolibrary/robot/CommandLineInterface.java
@@ -20,6 +20,7 @@ public class CommandLineInterface {
   private static CommandManager initManager() {
     CommandManager m = new CommandManager();
     m.addCommand("annotate", new AnnotateCommand());
+    m.addCommand("collapse", new CollapseCommand());
     m.addCommand("convert", new ConvertCommand());
     m.addCommand("diff", new DiffCommand());
     m.addCommand("explain", new ExplainCommand());
diff --git a/robot-core/src/main/java/org/obolibrary/robot/OntologyHelper.java b/robot-core/src/main/java/org/obolibrary/robot/OntologyHelper.java
index 14e0b88d..e9eaf787 100644
--- a/robot-core/src/main/java/org/obolibrary/robot/OntologyHelper.java
+++ b/robot-core/src/main/java/org/obolibrary/robot/OntologyHelper.java
@@ -141,7 +141,8 @@ public class OntologyHelper {
   /**
    * Given an ontology and a set of IRIs that must be retained, remove intermediate superclasses
    * (classes that only have one child) and update the subclass relationships to preserve structure.
-   * The ontology passed in is updated.
+   * The ontology passed in is updated. Do not perform collapse multiple times (some intermediate
+   * classes will remain after only one pass).
    *
    * @param ontology ontology to remove intermediates in
    * @param precious set of OWLEntities that should not be removed
@@ -149,60 +150,112 @@ public class OntologyHelper {
    */
   public static void collapseOntology(OWLOntology ontology, Set<IRI> precious)
       throws OWLOntologyCreationException {
+    collapseOntology(ontology, 2, precious, false);
+  }
 
-    OWLOntologyManager manager = OWLManager.createOWLOntologyManager();
-    OWLClass thing = manager.getOWLDataFactory().getOWLThing();
+  /**
+   * Given an ontology, a threshold, and a set of precious IRIs (or empty set), minimize the input
+   * ontology's class hierarchy based on the threshold. The threshold is the minimum number of child
+   * classes that an intermediate class should have. Any intermediate class that has less than the
+   * threshold number of children will be removed and its children will become children of the next
+   * level up. Bottom-level and top-level classes are not removed. Any class with an IRI in the
+   * precious set is not removed. If not repeat, collapse will only be performed once, meaning that
+   * some intermediate classes may remain.
+   *
+   * @param ontology OWLOntology to minimize
+   * @param threshold minimum number of child classes
+   * @param precious set of IRIs to keep
+   * @param repeat if true, repeat collapsing until no intermediate classes remain
+   */
+  public static void collapseOntology(
+      OWLOntology ontology, int threshold, Set<IRI> precious, boolean repeat)
+      throws OWLOntologyCreationException {
+    OWLOntology copy =
+        OWLManager.createOWLOntologyManager().copyOntology(ontology, OntologyCopy.DEEP);
+    logger.debug("Classes before collapsing: " + ontology.getClassesInSignature().size());
 
-    Set<OWLObject> removeObjects = new HashSet<>();
+    Set<OWLObject> removeClasses = getClassesToRemove(ontology, threshold, precious);
 
-    for (OWLEntity e : OntologyHelper.getEntities(ontology)) {
-      // Skip anything that is not a class
-      if (!e.isOWLClass()) {
-        continue;
-      }
-      // Skip any precious classes
-      OWLClass cls = e.asOWLClass();
-      if (precious.contains(cls.getIRI())) {
-        continue;
+    boolean collapsedOnce = false;
+
+    // Remove axioms based on classes
+    // Get all axioms that involve these classes
+    // Continue to get remove classes until there's no more to remove
+    while (!removeClasses.isEmpty()) {
+      if (collapsedOnce && !repeat) {
+        break;
       }
-      // Skip anything that is a SC of owl:Thing
-      Collection<OWLClassExpression> superClasses = EntitySearcher.getSuperClasses(cls, ontology);
-      if (superClasses.contains(thing)) {
+      Set<OWLAxiom> axiomsToRemove =
+          RelatedObjectsHelper.getPartialAxioms(ontology, removeClasses, null);
+      OWLOntologyManager manager = ontology.getOWLOntologyManager();
+      manager.removeAxioms(ontology, axiomsToRemove);
+      // Span gaps to maintain hierarchy
+      manager.addAxioms(
+          ontology, RelatedObjectsHelper.spanGaps(copy, OntologyHelper.getObjects(ontology)));
+      // Repeat until there's no more to remove
+      removeClasses = getClassesToRemove(ontology, threshold, precious);
+      collapsedOnce = true;
+    }
+    logger.debug("Classes after collapsing: " + ontology.getClassesInSignature().size());
+  }
+
+  /**
+   * Given an ontology, a threshold, and a set of precious IRIs (or empty set), return the classes
+   * to remove to minimize the class hierarchy. Top-level and bottom-level classes are not removed.
+   * Any class with a precious IRI is not removed. Any class with a number of named subclasses that
+   * is less than the threshold will be removed.
+   *
+   * @param ontology OWLOntology to minimize
+   * @param threshold minimum number of child classes
+   * @param precious set of IRIs to keep
+   */
+  private static Set<OWLObject> getClassesToRemove(
+      OWLOntology ontology, int threshold, Set<IRI> precious) {
+    Set<OWLClass> classes = ontology.getClassesInSignature();
+    Set<OWLObject> remove = new HashSet<>();
+
+    for (OWLClass cls : classes) {
+      if (cls.isOWLThing() || precious.contains(cls.getIRI())) {
+        // Ignore if the IRI is in precious or is OWL Thing
         continue;
       }
-      // Skip anything that does not have a named SC (default SC of owl:Thing)
-      boolean hasNamedSuperClass = false;
-      for (OWLClassExpression superExpr : superClasses) {
-        if (!superExpr.isAnonymous()) {
-          hasNamedSuperClass = true;
+
+      // Check for superclasses
+      Set<OWLSubClassOfAxiom> superAxioms = ontology.getSubClassAxiomsForSubClass(cls);
+      boolean hasNamedSuperclass = false;
+      for (OWLSubClassOfAxiom superAx : superAxioms) {
+        OWLClassExpression expr = superAx.getSuperClass();
+        if (!expr.isAnonymous() && !expr.asOWLClass().isOWLThing()) {
+          hasNamedSuperclass = true;
           break;
         }
       }
-      if (!hasNamedSuperClass) {
+
+      if (!hasNamedSuperclass) {
+        // Also ignore if there are no named superclasses
+        // Or just no superclasses in general
+        // This means it is directly placed under owl:Thing
         continue;
       }
-      Collection<OWLClassExpression> subClasses = EntitySearcher.getSubClasses(cls, ontology);
-      if (subClasses.size() == 1) {
-        removeObjects.add(cls);
-      }
-    }
-
-    logger.debug("Removing " + removeObjects.size() + " classes and associated axioms...");
 
-    // Remove the unnecessary intermediate classes
-    Set<OWLAxiom> axiomsToRemove =
-        RelatedObjectsHelper.getPartialAxioms(
-            ontology, removeObjects, new HashSet<>(Collections.singletonList(OWLAxiom.class)));
-    // Make copy before removing to preserve structure
-    OWLOntology copy = manager.copyOntology(ontology, OntologyCopy.DEEP);
-    manager.removeAxioms(ontology, axiomsToRemove);
+      Set<OWLSubClassOfAxiom> subAxioms = ontology.getSubClassAxiomsForSuperClass(cls);
+      int scCount = 0;
+      for (OWLSubClassOfAxiom subAx : subAxioms) {
+        OWLClassExpression expr = subAx.getSubClass();
+        if (!expr.isAnonymous()) {
+          // Only count the named subclasses
+          scCount++;
+        }
+      }
 
-    // Then re-associate classes with new superclasses
-    Set<OWLObject> relatedObjects = RelatedObjectsHelper.selectComplement(ontology, removeObjects);
-    manager.addAxioms(ontology, RelatedObjectsHelper.spanGaps(copy, relatedObjects));
+      if (scCount != 0 && scCount < threshold) {
+        // If the class has subclasses, but LESS subclasses than the threshold,
+        // add it to the set of classes to be removed
+        remove.add(cls);
+      }
+    }
 
-    // Discard copy
-    manager.removeOntology(copy);
+    return remove;
   }
 
   /**


Test patch:
diff --git a/robot-core/src/test/java/org/obolibrary/robot/OntologyHelperTest.java b/robot-core/src/test/java/org/obolibrary/robot/OntologyHelperTest.java
index f3572746..c3385610 100644
--- a/robot-core/src/test/java/org/obolibrary/robot/OntologyHelperTest.java
+++ b/robot-core/src/test/java/org/obolibrary/robot/OntologyHelperTest.java
@@ -7,11 +7,9 @@ import java.util.HashMap;
 import java.util.HashSet;
 import java.util.Map;
 import java.util.Set;
+import junit.framework.TestCase;
 import org.junit.Test;
-import org.semanticweb.owlapi.model.IRI;
-import org.semanticweb.owlapi.model.OWLAnnotation;
-import org.semanticweb.owlapi.model.OWLDataFactory;
-import org.semanticweb.owlapi.model.OWLOntology;
+import org.semanticweb.owlapi.model.*;
 
 /** Tests for OntologyHelper. */
 public class OntologyHelperTest extends CoreTest {
@@ -110,4 +108,35 @@ public class OntologyHelperTest extends CoreTest {
     OntologyHelper.removeOntologyAnnotations(simple);
     assertEquals(0, simple.getAnnotations().size());
   }
+
+  /**
+   * Test collapse ontology method.
+   *
+   * @throws IOException on issue loading ontology or running minimize
+   * @throws OWLOntologyCreationException on loading ontology
+   */
+  @Test
+  public void testCollapse() throws IOException, OWLOntologyCreationException {
+    OWLOntology ontology = loadOntology("/uberon.owl");
+    OntologyHelper.collapseOntology(ontology, 3, new HashSet<>(), true);
+    int after = ontology.getClassesInSignature().size();
+    TestCase.assertEquals(7, after);
+  }
+
+  /**
+   * Test collapse ontology method with a precious class.
+   *
+   * @throws IOException on issue loading ontology or running minimize
+   * @throws OWLOntologyCreationException on loading ontology
+   */
+  @Test
+  public void testCollapseWithPrecious() throws IOException, OWLOntologyCreationException {
+    OWLOntology ontology = loadOntology("/uberon.owl");
+    Set<IRI> precious = new HashSet<>();
+    // 'skeletal joint' will be kept
+    precious.add(IRI.create("http://purl.obolibrary.org/obo/UBERON_0000982"));
+    OntologyHelper.collapseOntology(ontology, 3, precious, true);
+    int after = ontology.getClassesInSignature().size();
+    TestCase.assertEquals(8, after);
+  }
 }