Highly selective journals have high publishing costs due to high rejection rates and bespoke author services. To make these articles Open Access using individual article processing charges (APCs), the resulting APCs are usually very high ($5K USD or more). Rather than charge the costly “true APC” that would cover the cost of publishing in those journals (~$6100 USD for PLOS Medicine and ~$5400 for PLOS Biology), we have historically deflated the APC to make it more affordable. With the PLOS Community Action Publishing effort, we will eliminate APCs altogether.