remains the same: proton pumping rate, electron transport rate, rate of oxygen uptakedecreases (or goes to zero): rate of ATP synthesis, size of the proton gradientGramicidin causes membranes to become very leaky to protons, so that a proton gradient cannot be maintained and ATP synthesis stops. However, the leakiness of the membrane has no effect on the ability of electron transport to pump protons. Thus, the rates of proton pumping, electron transport, and oxygen uptake remain unchanged.