Clan Donald, also known as Clan MacDonald, is one of the largest Scottish clans. They had strongholds throughout the Western Isles including Islay, Mull, Jura, Arran and Skye, as well as castles on the mainland. The clan can trace its roots to the Norse Lords of the Isles and the ruler Somerled, whose descendent Dugall mac Somhairle (died 1175) was a King of the Isles. The MacDonald dynasty also has bloodlines mixed with the High Kings of Ireland. Recent DNA studies of living clan members proves the lineage has a mix of Norse and Irish descendants.