An image sensor , comprising a plurality of pixels: each pixel of the plurality of pixels including: - a photoelectric conversion unit that photoelectrically converts light and generates an electric charge; an accumulation unit (FD) that accumulates the electric charge generated by the photoelectric conversion unit; and a transfer unit (Tx) that transfers the electric charge generated by the photoelectric conversion unit to the accumulation unit; and - a supply unit that supplies a signal based on a first voltage, Vneg, being lower than a substrate voltage, GND and being supplied from a first power supply unit and a second voltage, V1, being higher than the substrate voltage, GND, and being supplied from a second power supply unit provided in the pixel or a signal based on the first voltage and a predetermined voltage being the substrate voltage, GND, to the transfer unit, wherein the supply unit supplies the signal based on the first voltage, and the second voltage to the transfer unit such that the electric charge is transferred to the accumulation unit, and supplies the signal based on the first voltage and the predetermined voltage to the transfer unit such that the electric charge is not transferred to the accumulation unit, wherein the second voltage source is an individual power supply unit provided for each of the pixels. each pixel of the plurality of pixels including: - a photoelectric conversion unit that photoelectrically converts light and generates an electric charge; an accumulation unit (FD) that accumulates the electric charge generated by the photoelectric conversion unit; and a transfer unit (Tx) that transfers the electric charge generated by the photoelectric conversion unit to the accumulation unit; and - a supply unit that supplies a signal based on a first voltage, Vneg, being lower than a substrate voltage, GND and being supplied from a first power supply unit and a second voltage, V1, being higher than the substrate voltage, GND, and being supplied from a second power supply unit provided in the pixel or a signal based on the first voltage and a predetermined voltage being the substrate voltage, GND, to the transfer unit, - a photoelectric conversion unit that photoelectrically converts light and generates an electric charge; an accumulation unit (FD) that accumulates the electric charge generated by the photoelectric conversion unit; and a transfer unit (Tx) that transfers the electric charge generated by the photoelectric conversion unit to the accumulation unit; and - a supply unit that supplies a signal based on a first voltage, Vneg, being lower than a substrate voltage, GND and being supplied from a first power supply unit and a second voltage, V1, being higher than the substrate voltage, GND, and being supplied from a second power supply unit provided in the pixel or a signal based on the first voltage and a predetermined voltage being the substrate voltage, GND, to the transfer unit, wherein the supply unit supplies the signal based on the first voltage, and the second voltage to the transfer unit such that the electric charge is transferred to the accumulation unit, and supplies the signal based on the first voltage and the predetermined voltage to the transfer unit such that the electric charge is not transferred to the accumulation unit, wherein the second voltage source is an individual power supply unit provided for each of the pixels. <EOS>