An image sensor comprising: a first image-capturing unit that includes a plurality of microlenses arranged in a two dimensional pattern and a plurality of light reception units that are provided in correspondence to each of the microlenses, respectively, and that receive, absorb and photoelectrically convert light having a predetermined wavelength and transmit lights having wavelengths different from the predetermined wavelength; and a second image-capturing unit that receives the lights that are transmitted through the first image-capturing unit, wherein the first image-capturing unit includes first, second and third photoelectric conversion units absorbing and photoelectrically converting lights having first, second and third wavelengths, respectively, differing from each other, and being arranged in a two dimensional pattern, the second image-capturing unit includes fourth, fifth and sixth photoelectric conversion units absorbing and photoelectrically converting lights having fourth, fifth and sixth wavelengths, respectively, that are complementary to the lights having the first, second, and third wavelengths, respectively, in the first image-capturing unit, being arranged in a two dimensional pattern, and positions of the first, second and third photoelectric conversion units in the first image-capturing unit and positions of the fourth, fifth and sixth photoelectric conversion units in the second image-capturing unit are determined such that the fourth, fifth and sixth photoelectric conversion units receive light fluxes that pass through the first, second and third photoelectric conversion units, respectively. a first image-capturing unit that includes a plurality of microlenses arranged in a two dimensional pattern and a plurality of light reception units that are provided in correspondence to each of the microlenses, respectively, and that receive, absorb and photoelectrically convert light having a predetermined wavelength and transmit lights having wavelengths different from the predetermined wavelength; and a plurality of microlenses arranged in a two dimensional pattern and a plurality of light reception units that are provided in correspondence to each of the microlenses, respectively, and that receive, absorb and photoelectrically convert light having a predetermined wavelength and transmit lights having wavelengths different from the predetermined wavelength; and a second image-capturing unit that receives the lights that are transmitted through the first image-capturing unit, wherein the first image-capturing unit includes first, second and third photoelectric conversion units absorbing and photoelectrically converting lights having first, second and third wavelengths, respectively, differing from each other, and being arranged in a two dimensional pattern, the second image-capturing unit includes fourth, fifth and sixth photoelectric conversion units absorbing and photoelectrically converting lights having fourth, fifth and sixth wavelengths, respectively, that are complementary to the lights having the first, second, and third wavelengths, respectively, in the first image-capturing unit, being arranged in a two dimensional pattern, and positions of the first, second and third photoelectric conversion units in the first image-capturing unit and positions of the fourth, fifth and sixth photoelectric conversion units in the second image-capturing unit are determined such that the fourth, fifth and sixth photoelectric conversion units receive light fluxes that pass through the first, second and third photoelectric conversion units, respectively.