A sole structure for an article of footwear, the sole structure comprising: a sole body portion, the sole body portion including an outsole surface facing away from the article of footwear and an upper surface disposed opposite the outsole surface; and a central sensory node element disposed within an aperture in the sole body portion, the aperture being located at least partially within at least one of a forefoot region and a midfoot region of the sole structure and located between a medial side and a lateral side of the sole structure; the central sensory node element including a bottom surface configured to engage a ground surface and a top surface disposed opposite the bottom surface; the bottom surface of the central sensory node element extending above the outsole surface of the sole body portion when the central sensory node element is in an uncompressed condition; wherein the central sensory node element is configured to move vertically within the aperture in the sole body portion so that the bottom surface of the central sensory node element moves closer towards the outsole surface of the sole body portion when the central sensory node element is in a compressed condition; wherein the top surface of the central sensory node element is attached to a base layer and wherein the base layer is attached to the upper surface of the sole body portion such that the sole body portion and the central sensory node element are not directly joined to each other, wherein the central sensory node element has an approximately truncated cone shape; and wherein the top surface of the central sensory node element is located at a top end where the central sensory node element has a smaller diameter than an opposite bottom end where bottom surface is located. a sole body portion, the sole body portion including an outsole surface facing away from the article of footwear and an upper surface disposed opposite the outsole surface; and a central sensory node element disposed within an aperture in the sole body portion, the aperture being located at least partially within at least one of a forefoot region and a midfoot region of the sole structure and located between a medial side and a lateral side of the sole structure; the central sensory node element including a bottom surface configured to engage a ground surface and a top surface disposed opposite the bottom surface; the bottom surface of the central sensory node element extending above the outsole surface of the sole body portion when the central sensory node element is in an uncompressed condition; wherein the central sensory node element is configured to move vertically within the aperture in the sole body portion so that the bottom surface of the central sensory node element moves closer towards the outsole surface of the sole body portion when the central sensory node element is in a compressed condition; wherein the top surface of the central sensory node element is attached to a base layer and wherein the base layer is attached to the upper surface of the sole body portion such that the sole body portion and the central sensory node element are not directly joined to each other, wherein the central sensory node element has an approximately truncated cone shape; and wherein the top surface of the central sensory node element is located at a top end where the central sensory node element has a smaller diameter than an opposite bottom end where bottom surface is located.