A filter element for an air filter, the filter element comprising: a filter media pack having an inlet end, an outlet end and an outer periphery extending between the inlet end and the outlet end, the inlet end and the outlet end being separated along a central axis, the outer periphery including a first pair of opposed sides between the inlet end and the outlet end, and a second pair of opposed sides extending transversely between the first pair of opposed sides between the inlet end and the outlet end; and a frame supporting the outer periphery of the filter media pack, the frame including opposing first frame side segments along the first pair of opposed sides of the media pack, and a second frame segment along at least one of the second opposed sides of the media pack, the first frame side segments each including a pair of guide projections along the respective first sides of the media pack from one of the second sides to the other of the second sides, the guide projections being spaced apart from one another from a greater distance at the one second side, to a smaller distance at the other second side and arranged to receive axial forces when received within a housing having complementary engagement surfaces; and a housing seal supported by the frame along at least one guide projection on each side of the media pack, each housing seal being arranged to seal against one of the complementary engagement surfaces, characterized in that one guide projection includes a plurality of spaced-apart, lateral projection components, with one of the projection components being closer to the one of the second sides, and another of the projection components being closer to the other of the second sides. a filter media pack having an inlet end, an outlet end and an outer periphery extending between the inlet end and the outlet end, the inlet end and the outlet end being separated along a central axis, the outer periphery including a first pair of opposed sides between the inlet end and the outlet end, and a second pair of opposed sides extending transversely between the first pair of opposed sides between the inlet end and the outlet end; and a frame supporting the outer periphery of the filter media pack, the frame including opposing first frame side segments along the first pair of opposed sides of the media pack, and a second frame segment along at least one of the second opposed sides of the media pack, the first frame side segments each including a pair of guide projections along the respective first sides of the media pack from one of the second sides to the other of the second sides, the guide projections being spaced apart from one another from a greater distance at the one second side, to a smaller distance at the other second side and arranged to receive axial forces when received within a housing having complementary engagement surfaces; and a housing seal supported by the frame along at least one guide projection on each side of the media pack, each housing seal being arranged to seal against one of the complementary engagement surfaces, characterized in that one guide projection includes a plurality of spaced-apart, lateral projection components, with one of the projection components being closer to the one of the second sides, and another of the projection components being closer to the other of the second sides.