A filter media pack (20, 302, 402, 502, 602) comprising: (a) a media construction comprising alternating first media sheet (24, 40) and a second media sheet (26, 40) secured together and forming a first flow face and a second flow face, wherein each of the first media sheet (24, 40) and the second media sheet (26, 40) includes a first side and a second side; (i) the first media sheet (24, 40) including a plurality of first bosses extending from the first media sheet (24, 40) first side toward the second media sheet (26, 40) to provide a first separation between the first media sheet (24, 40) first side and the second media sheet (26, 40) second side, and a plurality of second bosses extending from the first media sheet (24, 40) second side in a direction opposite the plurality of first bosses to provide a second separation between the first media sheet (24, 40) second side and the second media sheet (26, 40) first side; wherein the first separation is greater than the second separation; (ii) the first media sheet (24, 40) and the second media sheet (26, 40) secured together at a first location to form the first separation open at the second flow face, and the first media sheet (24, 40) and the second media sheet (26, 40) secured together at a second location to form the second separation open at the first flow face; (b) the media construction being closed to passage of unfiltered fluid therethrough from the first flow face to the second flow face or from the second flow face to the first flow face without filtering passage through the first media sheet (24, 40) or the second media sheet (26, 40). (a) a media construction comprising alternating first media sheet (24, 40) and a second media sheet (26, 40) secured together and forming a first flow face and a second flow face, wherein each of the first media sheet (24, 40) and the second media sheet (26, 40) includes a first side and a second side; (i) the first media sheet (24, 40) including a plurality of first bosses extending from the first media sheet (24, 40) first side toward the second media sheet (26, 40) to provide a first separation between the first media sheet (24, 40) first side and the second media sheet (26, 40) second side, and a plurality of second bosses extending from the first media sheet (24, 40) second side in a direction opposite the plurality of first bosses to provide a second separation between the first media sheet (24, 40) second side and the second media sheet (26, 40) first side; wherein the first separation is greater than the second separation; (ii) the first media sheet (24, 40) and the second media sheet (26, 40) secured together at a first location to form the first separation open at the second flow face, and the first media sheet (24, 40) and the second media sheet (26, 40) secured together at a second location to form the second separation open at the first flow face; (i) the first media sheet (24, 40) including a plurality of first bosses extending from the first media sheet (24, 40) first side toward the second media sheet (26, 40) to provide a first separation between the first media sheet (24, 40) first side and the second media sheet (26, 40) second side, and a plurality of second bosses extending from the first media sheet (24, 40) second side in a direction opposite the plurality of first bosses to provide a second separation between the first media sheet (24, 40) second side and the second media sheet (26, 40) first side; wherein the first separation is greater than the second separation; (ii) the first media sheet (24, 40) and the second media sheet (26, 40) secured together at a first location to form the first separation open at the second flow face, and the first media sheet (24, 40) and the second media sheet (26, 40) secured together at a second location to form the second separation open at the first flow face; (b) the media construction being closed to passage of unfiltered fluid therethrough from the first flow face to the second flow face or from the second flow face to the first flow face without filtering passage through the first media sheet (24, 40) or the second media sheet (26, 40).