A surgical instrument assembly, comprising: a shaft; an articulation joint; an end effector rotatably connected to said shaft about said articulation joint, wherein said end effector comprises: a first jaw; a second jaw movable relative to said first jaw; a staple cartridge comprising a plurality of staples removably stored therein; an anvil configured to deform said staples; a drive input; a closure system configured to move said second jaw between an open position and a closed position; a staple firing system configured to eject said staples from said staple cartridge; and a transmission positioned in said end effector wherein said transmission is configurable in three operating states: a closure operating state in which said drive input is operably coupled with said closure system and operably decoupled from said staple firing system; a firing operating state in which said drive input is operably coupled with said staple firing system and operably decoupled from said closure system; and a third operating state in which said drive input is operably coupled with said closure system and said staple firing system such that said closure system and said staple firing system are configured to be operated simultaneously. a shaft; an articulation joint; an end effector rotatably connected to said shaft about said articulation joint, wherein said end effector comprises: a first jaw; a second jaw movable relative to said first jaw; a staple cartridge comprising a plurality of staples removably stored therein; an anvil configured to deform said staples; a drive input; a closure system configured to move said second jaw between an open position and a closed position; a staple firing system configured to eject said staples from said staple cartridge; and a transmission positioned in said end effector wherein said transmission is configurable in three operating states: a closure operating state in which said drive input is operably coupled with said closure system and operably decoupled from said staple firing system; a firing operating state in which said drive input is operably coupled with said staple firing system and operably decoupled from said closure system; and a third operating state in which said drive input is operably coupled with said closure system and said staple firing system such that said closure system and said staple firing system are configured to be operated simultaneously. a first jaw; a second jaw movable relative to said first jaw; a staple cartridge comprising a plurality of staples removably stored therein; an anvil configured to deform said staples; a drive input; a closure system configured to move said second jaw between an open position and a closed position; a staple firing system configured to eject said staples from said staple cartridge; and a transmission positioned in said end effector wherein said transmission is configurable in three operating states: a closure operating state in which said drive input is operably coupled with said closure system and operably decoupled from said staple firing system; a firing operating state in which said drive input is operably coupled with said staple firing system and operably decoupled from said closure system; and a third operating state in which said drive input is operably coupled with said closure system and said staple firing system such that said closure system and said staple firing system are configured to be operated simultaneously. a closure operating state in which said drive input is operably coupled with said closure system and operably decoupled from said staple firing system; a firing operating state in which said drive input is operably coupled with said staple firing system and operably decoupled from said closure system; and a third operating state in which said drive input is operably coupled with said closure system and said staple firing system such that said closure system and said staple firing system are configured to be operated simultaneously.