An on-machine development type negative planographic printing plate precursor, comprising: a support; and an image-recording layer on the support, wherein the image-recording layer contains an electron-accepting polymerization initiator, an infrared absorber capable of donating electrons to the initiator, and a color-developing substance precursor, characterized in that, in a case where the image-recording layer is exposed to an infrared laser with a wavelength of 830 nm at an energy density of 110 mJ/cm 2, a brightness change ΔL of the image-recording layer before and after the exposure is 3.0 or more, a molar light absorption coefficient ε of a color-developing substance generated from the color-developing substance precursor is 35,000 or more, a ring-opening rate of the color-developing substance precursor calculated by the following equation is 40 mol% to 99 mol%, ring-opening rate = molar light absorption coefficient of the color-developing substance precursor to which 1 molar equivalent of acid is added/molar light absorption coefficient ε of a color-developing substance generated from the color-developing substance precursor × 100, a color-developing substance generated from the color-developing substance precursor has a maximum absorption wavelength of 500 nm to 650 nm in a wavelength range of 380 nm to 750 nm, the color-developing substance precursor has 2 or more electron donating groups each directly bonded to an aromatic ring, and the electron donating groups are each independently a di(unsubstituted alkyl)amino group, a monoalkyl monoarylamino group, or a diarylamino group, and printing plate precursors as prepared in Examples 23, 24, 26 and 31-34 of EP 3 991 983 A1 and in Examples 14-17 of EP 3 991 988 A1 are excluded, wherein the color-developing substance precursor is a compound represented by Formula (Z-1) or Formula (Z-4), in Formula (Z-1), EDG each independently represents a di(unsubstituted alkyl)amino group, a monoalkyl monoarylamino group, or a diarylamino group; X represents O or NR; R represents a hydrogen atom, an alkyl group, an aryl group, or a heteroaryl group; and Y 1 and Y 2 each independently represent CH or N, in Formula (Z-4), Ra 1 represents a hydrogen atom, an alkyl group, or an alkoxy group; Rb 1 and Rb 2 each independently represent an unsubstituted alkyl group, or Rb 1 and Rb 2 represent an alkyl group and an aryl group, respectively, or Rb 2 and Rb 1 represent an alkyl group and an aryl group, respectively; Rb 3 and Rb 4 each independently represent an aryl group; Rb 1 and Rb 2 or Rb 3 and Rb 4 may form a ring; X represents O or NR; R represents a hydrogen atom, an alkyl group, an aryl group, or a heteroaryl group; and Y 1 and Y 2 each independently represent CH or N. a support; and an image-recording layer on the support, wherein the image-recording layer contains an electron-accepting polymerization initiator, an infrared absorber capable of donating electrons to the initiator, and a color-developing substance precursor, characterized in that, in a case where the image-recording layer is exposed to an infrared laser with a wavelength of 830 nm at an energy density of 110 mJ/cm 2, a brightness change ΔL of the image-recording layer before and after the exposure is 3.0 or more, a molar light absorption coefficient ε of a color-developing substance generated from the color-developing substance precursor is 35,000 or more, a ring-opening rate of the color-developing substance precursor calculated by the following equation is 40 mol% to 99 mol%, ring-opening rate = molar light absorption coefficient of the color-developing substance precursor to which 1 molar equivalent of acid is added/molar light absorption coefficient ε of a color-developing substance generated from the color-developing substance precursor × 100, a color-developing substance generated from the color-developing substance precursor has a maximum absorption wavelength of 500 nm to 650 nm in a wavelength range of 380 nm to 750 nm, the color-developing substance precursor has 2 or more electron donating groups each directly bonded to an aromatic ring, and the electron donating groups are each independently a di(unsubstituted alkyl)amino group, a monoalkyl monoarylamino group, or a diarylamino group, and printing plate precursors as prepared in Examples 23, 24, 26 and 31-34 of EP 3 991 983 A1 and in Examples 14-17 of EP 3 991 988 A1 are excluded, wherein the color-developing substance precursor is a compound represented by Formula (Z-1) or Formula (Z-4), in Formula (Z-1), EDG each independently represents a di(unsubstituted alkyl)amino group, a monoalkyl monoarylamino group, or a diarylamino group; X represents O or NR; R represents a hydrogen atom, an alkyl group, an aryl group, or a heteroaryl group; and Y 1 and Y 2 each independently represent CH or N, in Formula (Z-4), Ra 1 represents a hydrogen atom, an alkyl group, or an alkoxy group; Rb 1 and Rb 2 each independently represent an unsubstituted alkyl group, or Rb 1 and Rb 2 represent an alkyl group and an aryl group, respectively, or Rb 2 and Rb 1 represent an alkyl group and an aryl group, respectively; Rb 3 and Rb 4 each independently represent an aryl group; Rb 1 and Rb 2 or Rb 3 and Rb 4 may form a ring; X represents O or NR; R represents a hydrogen atom, an alkyl group, an aryl group, or a heteroaryl group; and Y 1 and Y 2 each independently represent CH or N.