A dispenser for dispensing fluid comprising: a housing, a reservoir, the housing supporting the reservoir in a dispensing position, the reservoir being insertable into said housing to assume said dispensing position and removable therefrom for replacement, the reservoir comprising: (i) a chamber for retaining fluid having an outlet, and (ii) a valving mechanism disposed across the outlet for dispensing fluid from the chamber, the valving mechanism comprising a reciprocally movable element for reciprocal movement along an axis relative to the housing when the reservoir is in the dispensing position to dispense fluid, the housing including an actuator mechanism for moving the reciprocally movable element along the axis, the actuator mechanism movable relative the housing between a first position and a second position, the actuator mechanism including a catch member, the reciprocally movable element including an engagement mechanism to engage the catch member and couple the reciprocally movable element to the actuator mechanism, characterized by: the engagement mechanism comprising a resilient member extending radially outwardly relative the axis from the reciprocally movable element in an unbiased inherent position, the resilient member being resiliently deflectable radially from the unbiased inherent position to a deflected position of a different radial extent relative the axis than the unbiased inherent position and biased to return to the unbiased inherent position when deflected away from the unbiased inherent position toward the deflected position, wherein when the reservoir is inserted into the dispensing position, the engagement mechanism and catch member assume an uncoupled orientation from which on a first cycling of the actuator mechanism between the first position and the second position, the engagement mechanism and catch member are moved relative to each other such that the catch member is forced axially into the resilient member, wherein from the uncoupled orientation on relative forcing the catch member axially into the resilient member engagement therebetween deflects the resilient member radially sufficiently to permit the catch member to move axially from the uncoupled orientation to a coupled orientation in which the engagement mechanism engages the catch member for reciprocal movement of the reciprocally movable element to dispense fluid with movement of the actuator mechanism between the first position and the second position, and wherein in the coupled orientation, the resilient member returns to the unbiased inherent position. a housing, a reservoir, the housing supporting the reservoir in a dispensing position, the reservoir being insertable into said housing to assume said dispensing position and removable therefrom for replacement, (i) a chamber for retaining fluid having an outlet, and (ii) a valving mechanism disposed across the outlet for dispensing fluid from the chamber, the valving mechanism comprising a reciprocally movable element for reciprocal movement along an axis relative to the housing when the reservoir is in the dispensing position to dispense fluid, the housing including an actuator mechanism for moving the reciprocally movable element along the axis, the actuator mechanism movable relative the housing between a first position and a second position, the actuator mechanism including a catch member, the reciprocally movable element including an engagement mechanism to engage the catch member and couple the reciprocally movable element to the actuator mechanism, the engagement mechanism comprising a resilient member extending radially outwardly relative the axis from the reciprocally movable element in an unbiased inherent position, the resilient member being resiliently deflectable radially from the unbiased inherent position to a deflected position of a different radial extent relative the axis than the unbiased inherent position and biased to return to the unbiased inherent position when deflected away from the unbiased inherent position toward the deflected position, wherein when the reservoir is inserted into the dispensing position, the engagement mechanism and catch member assume an uncoupled orientation from which on a first cycling of the actuator mechanism between the first position and the second position, the engagement mechanism and catch member are moved relative to each other such that the catch member is forced axially into the resilient member, wherein from the uncoupled orientation on relative forcing the catch member axially into the resilient member engagement therebetween deflects the resilient member radially sufficiently to permit the catch member to move axially from the uncoupled orientation to a coupled orientation in which the engagement mechanism engages the catch member for reciprocal movement of the reciprocally movable element to dispense fluid with movement of the actuator mechanism between the first position and the second position, and wherein in the coupled orientation, the resilient member returns to the unbiased inherent position.