A ferrule (10K, 10L) that is mutually connectable to a second ferrule (10K, 10L) having the same structure, the ferrule comprising: a plurality of holding portions aligned in a first direction and further arranged in one or a plurality of stages in a second direction orthogonal to the first direction, and configured to provide a first optical axis by holding optical waveguide members, the first optical axis extending in a third direction orthogonal to the first direction and the second direction; a light incidence/emission plane configured to pass light entering or emitted from the optical waveguide members respectively held at the plurality of holding portions, and further configured to face the second ferrule; a plurality of lenses disposed on optical axes between the respective plural holding portions and the light incidence/emission plane; and two or more guide portions configured to determine a relative position with the second ferrule in the first direction and the second direction, wherein a positional relation between the two or more guide portions is 180-degree rotationally symmetric around a reference axial line extending in the third direction, characterized in that a second optical axis between the light incidence/emission plane and the second ferrule is inclined relative to the third direction, wherein the plurality of lenses is disposed disproportionately in a direction opposite to an inclination direction of the second optical axis from a position line-symmetric relative to a straight line crossing with the reference axial line and parallel to the second direction, the number of the stages of the holding portions in the second direction is smaller than the number of holding portions aligned in the first direction, and wherein the second optical axis is inclined in the first direction. a plurality of holding portions aligned in a first direction and further arranged in one or a plurality of stages in a second direction orthogonal to the first direction, and configured to provide a first optical axis by holding optical waveguide members, the first optical axis extending in a third direction orthogonal to the first direction and the second direction; a light incidence/emission plane configured to pass light entering or emitted from the optical waveguide members respectively held at the plurality of holding portions, and further configured to face the second ferrule; a plurality of lenses disposed on optical axes between the respective plural holding portions and the light incidence/emission plane; and two or more guide portions configured to determine a relative position with the second ferrule in the first direction and the second direction, wherein a positional relation between the two or more guide portions is 180-degree rotationally symmetric around a reference axial line extending in the third direction, characterized in that a second optical axis between the light incidence/emission plane and the second ferrule is inclined relative to the third direction, wherein the plurality of lenses is disposed disproportionately in a direction opposite to an inclination direction of the second optical axis from a position line-symmetric relative to a straight line crossing with the reference axial line and parallel to the second direction, the number of the stages of the holding portions in the second direction is smaller than the number of holding portions aligned in the first direction, and wherein the second optical axis is inclined in the first direction.