An air filtration media pack (301, 302, 402, 502, 602) comprising: (a) a plurality of layers of single facer media (5, 117, 137, 147) wherein the layer of single facer media (5, 117, 137, 147) comprising a fluted sheet (3, 112, 122, 142), a facing sheet (4, 113, 123, 143), and a plurality of flutes (11, 15) extending between the fluted sheet (3, 112, 122, 142) and the facing sheet (4, 113, 123, 143) and having a flute length extending from a first face (9, 304) of the filtration media pack (301, 302, 402, 502, 602) to a second face (8, 306) of the filtration media pack (301, 302, 402, 502, 602); (b) a first portion of the plurality of flutes (11, 15) being closed to unfiltered air flowing into the first portion of the plurality of flutes (11, 15), and a second portion of the plurality of flutes (11, 15) being closed to unfiltered air flowing out of the second portion of the plurality of flutes (11, 15) so that air passing into one of the first face (9, 304) or the second face (8, 306) of the media pack (301, 302, 402, 502, 602) and out the other of the first face (9, 304) or the second face (8, 306) of the media pack (301, 302, 402, 502, 602) passes through media to provide filtration of the air; and (c) wherein the fluted sheet (3, 112, 122, 142) comprises: (i) repeating internal peaks facing (76, 114, 124, 144) toward the facing sheet (4, 113, 123, 143) and repeating external peaks (7a, 115, 125, 145) facing away from the facing sheet (4, 113, 123, 143); (ii) a repeating pattern of flutes (11, 15) comprising a flute having ridges (118, 128, 129, 148, 149) extending along at least a portion of the flute length between an internal peak and an adjacent external peak; and (iii) a tapered configuration tapering from a first configuration to a second configuration; and when tapering from the first configuration to the second configuration the ridges (118, 128, 129, 148, 149) approach the internal peaks (76, 114, 124, 144) and moves away from the external peaks (7a, 115, 125, 145). (a) a plurality of layers of single facer media (5, 117, 137, 147) wherein the layer of single facer media (5, 117, 137, 147) comprising a fluted sheet (3, 112, 122, 142), a facing sheet (4, 113, 123, 143), and a plurality of flutes (11, 15) extending between the fluted sheet (3, 112, 122, 142) and the facing sheet (4, 113, 123, 143) and having a flute length extending from a first face (9, 304) of the filtration media pack (301, 302, 402, 502, 602) to a second face (8, 306) of the filtration media pack (301, 302, 402, 502, 602); (b) a first portion of the plurality of flutes (11, 15) being closed to unfiltered air flowing into the first portion of the plurality of flutes (11, 15), and a second portion of the plurality of flutes (11, 15) being closed to unfiltered air flowing out of the second portion of the plurality of flutes (11, 15) so that air passing into one of the first face (9, 304) or the second face (8, 306) of the media pack (301, 302, 402, 502, 602) and out the other of the first face (9, 304) or the second face (8, 306) of the media pack (301, 302, 402, 502, 602) passes through media to provide filtration of the air; and (c) wherein the fluted sheet (3, 112, 122, 142) comprises: (i) repeating internal peaks facing (76, 114, 124, 144) toward the facing sheet (4, 113, 123, 143) and repeating external peaks (7a, 115, 125, 145) facing away from the facing sheet (4, 113, 123, 143); (ii) a repeating pattern of flutes (11, 15) comprising a flute having ridges (118, 128, 129, 148, 149) extending along at least a portion of the flute length between an internal peak and an adjacent external peak; and (iii) a tapered configuration tapering from a first configuration to a second configuration; and when tapering from the first configuration to the second configuration the ridges (118, 128, 129, 148, 149) approach the internal peaks (76, 114, 124, 144) and moves away from the external peaks (7a, 115, 125, 145). (i) repeating internal peaks facing (76, 114, 124, 144) toward the facing sheet (4, 113, 123, 143) and repeating external peaks (7a, 115, 125, 145) facing away from the facing sheet (4, 113, 123, 143); (ii) a repeating pattern of flutes (11, 15) comprising a flute having ridges (118, 128, 129, 148, 149) extending along at least a portion of the flute length between an internal peak and an adjacent external peak; and (iii) a tapered configuration tapering from a first configuration to a second configuration; and when tapering from the first configuration to the second configuration the ridges (118, 128, 129, 148, 149) approach the internal peaks (76, 114, 124, 144) and moves away from the external peaks (7a, 115, 125, 145).