A surgical instrument, comprising: a surgical end effector comprising: a first jaw; and a second jaw movably supported relative to said first jaw for selective movement relative thereto between an open position and closed positions and wherein said surgical instrument further comprises a closure member axially movable in response to closing and opening motions applied thereto, said closure member comprising at least one opening cam protruding therefrom to movably engage a corresponding slotted cam surface on said second jaw such that upon application of said opening motion to said closure member, said at least one opening cam movably engages said corresponding slotted cam surface to move said second jaw to said open position and upon application of said closure motion to said closure member, said closure member engages said second jaw to move said second jaw to one of said closed positions, wherein said at least one opening cam comprises: a first opening cam extending laterally inwardly from said closure member and engaging a first one of said corresponding slotted cam surfaces; and a second opening cam extending laterally inwardly from said closure member and engaging a second one of said corresponding slotted cam surfaces. a surgical end effector comprising: a first jaw; and a second jaw movably supported relative to said first jaw for selective movement relative thereto between an open position and closed positions and wherein said surgical instrument further comprises a closure member axially movable in response to closing and opening motions applied thereto, said closure member comprising at least one opening cam protruding therefrom to movably engage a corresponding slotted cam surface on said second jaw such that upon application of said opening motion to said closure member, said at least one opening cam movably engages said corresponding slotted cam surface to move said second jaw to said open position and upon application of said closure motion to said closure member, said closure member engages said second jaw to move said second jaw to one of said closed positions, a first jaw; and a second jaw movably supported relative to said first jaw for selective movement relative thereto between an open position and closed positions and wherein said surgical instrument further comprises a closure member axially movable in response to closing and opening motions applied thereto, said closure member comprising at least one opening cam protruding therefrom to movably engage a corresponding slotted cam surface on said second jaw such that upon application of said opening motion to said closure member, said at least one opening cam movably engages said corresponding slotted cam surface to move said second jaw to said open position and upon application of said closure motion to said closure member, said closure member engages said second jaw to move said second jaw to one of said closed positions, wherein said at least one opening cam comprises: a first opening cam extending laterally inwardly from said closure member and engaging a first one of said corresponding slotted cam surfaces; and a second opening cam extending laterally inwardly from said closure member and engaging a second one of said corresponding slotted cam surfaces. a first opening cam extending laterally inwardly from said closure member and engaging a first one of said corresponding slotted cam surfaces; and a second opening cam extending laterally inwardly from said closure member and engaging a second one of said corresponding slotted cam surfaces.