A recombinant nucleic acid molecule encoding a 4-hydroxyphenylpyruvate dioxygenase (HPPD) protein consisting of an amino acid sequence comprising a proline at the amino acid position corresponding to amino acid position 335 of SEQ ID NO:1 and a phenylalanine or a tyrosine at the position corresponding to amino acid position 336 of SEQ ID NO:1, and wherein said HPPD protein is tolerant to an HPPD inhibitor herbicide, wherein said encoded HPPD protein consists of an amino acid sequence further comprising: (a) an alanine at the amino acid position corresponding to amino acid position 188 of SEQ ID NO:1, a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (b) an isoleucine at the amino acid position corresponding to amino acid position 200 of SEQ ID NO:1, a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (c) a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (d) a glycine at the amino acid position corresponding to amino acid position 172 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (e) an isoleucine at the amino acid position corresponding to amino acid position 200 of SEQ ID NO:1 and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; or (f) an alanine at the amino acid position corresponding to amino acid position 188 of SEQ ID NO:1, an isoleucine at the amino acid position corresponding to amino acid position 200 of SEQ ID NO:1, a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1. (a) an alanine at the amino acid position corresponding to amino acid position 188 of SEQ ID NO:1, a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (b) an isoleucine at the amino acid position corresponding to amino acid position 200 of SEQ ID NO:1, a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (c) a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (d) a glycine at the amino acid position corresponding to amino acid position 172 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; (e) an isoleucine at the amino acid position corresponding to amino acid position 200 of SEQ ID NO:1 and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1; or (f) an alanine at the amino acid position corresponding to amino acid position 188 of SEQ ID NO:1, an isoleucine at the amino acid position corresponding to amino acid position 200 of SEQ ID NO:1, a histidine at the amino acid position corresponding to amino acid position 226 of SEQ ID NO:1, and a glycine at the position corresponding to amino acid position 340 of SEQ ID NO:1.