Storage system having a storage arrangement for storing loading units, wherein - the storage arrangement has at least one first storage region and one second storage region arranged vertically above the first storage region, characterised in that - the first storage region has a first storage structure, which is formed by first horizontal storage and retrieval routes and the second storage region has a second storage structure, which is formed by second horizontal storage and retrieval routes, so that such a storage structure is a parallel rack structure of respective rows of racks parallel to each other alternating with storage and retrieval routes parallel to each other, - wherein each storage and retrieval route is a passageway and/or an aisle in a storage region which is respectively delimited on both sides by rack units in the respective rack structure in its transverse direction and wherein each storage and retrieval route is an at least approximately rectilinear route extending in a longitudinal direction via which an operating system/resource moves; - wherein the operating system/resource is equipped with first operating units embodied for the first storage region and second operating units for the second storage region, which in each case is embodied in the form of a combined floor-level conveyor-hoist for shifting and/or retrieving/storing the loading units in the first and the second storage area (4 and 5) and also can be referred to as a shuttle; and - a first orientation of the first storage structure is rotated in relation to a second orientation of the second storage structure; - wherein the storage system comprises a processing area for the loading units, which is arranged vertically below the storage arrangement; - and wherein the storage system comprises at least one lifting device for the loading units, which can also be referred to as a lift and by means of which the first storage region and the second storage region are connected to the processing area; and - wherein the shuttle transports the loading units in its storage and retrieval routes to the nearest lift. - the storage arrangement has at least one first storage region and one second storage region arranged vertically above the first storage region, characterised in that - the first storage region has a first storage structure, which is formed by first horizontal storage and retrieval routes and the second storage region has a second storage structure, which is formed by second horizontal storage and retrieval routes, so that such a storage structure is a parallel rack structure of respective rows of racks parallel to each other alternating with storage and retrieval routes parallel to each other, - wherein each storage and retrieval route is a passageway and/or an aisle in a storage region which is respectively delimited on both sides by rack units in the respective rack structure in its transverse direction and wherein each storage and retrieval route is an at least approximately rectilinear route extending in a longitudinal direction via which an operating system/resource moves; - wherein the operating system/resource is equipped with first operating units embodied for the first storage region and second operating units for the second storage region, which in each case is embodied in the form of a combined floor-level conveyor-hoist for shifting and/or retrieving/storing the loading units in the first and the second storage area (4 and 5) and also can be referred to as a shuttle; and - a first orientation of the first storage structure is rotated in relation to a second orientation of the second storage structure; - wherein the storage system comprises a processing area for the loading units, which is arranged vertically below the storage arrangement; - and wherein the storage system comprises at least one lifting device for the loading units, which can also be referred to as a lift and by means of which the first storage region and the second storage region are connected to the processing area; and - wherein the shuttle transports the loading units in its storage and retrieval routes to the nearest lift.