A touch substrate, comprising a periphery area and a touch area, wherein the touch area comprises a first edge and a second edge opposite to each other, wherein the touch substrate comprises bonding terminals (1041, 1042) arranged in the periphery area, touch electrodes (1021, 1022) arranged in the touch area and signal transmission lines (1051, 1052) connecting the bonding terminals (1041, 1042) and the touch electrodes (1021, 1022), wherein the signal transmission lines (1041, 1042) are arranged between the first edge and the second edge, wherein: the bonding terminals (1041, 1042) comprise a first bonding terminal and a second bonding terminal arranged between the first edge and the second edge; the touch electrodes (1021, 1022) comprise a first touch electrode and a second touch electrode; the signal transmission lines (1051, 1052) comprise a first signal transmission line connecting the first bonding terminal and the first touch electrode and a second signal transmission line connecting the second bonding terminal and the second touch electrode; the first touch electrode extends toward the touch area from the bonding terminals (1041, 1042), and the second touch electrode and the first touch electrode are crossed to each other, and the first signal transmission line and the first touch electrode are arranged in a same layer; the second signal transmission line comprises a first part extending along the first touch electrode and arranged in the touch area; characterized in that: the first part is arranged in a different layer from the second touch electrode; the second signal transmission line further comprises a second part arranged in the touch area and connected to the first part through a via hole, wherein the second part extends along the second touch electrode and is in a same layer with the second touch electrode and electrically connected to the second touch electrode; and the first touch electrode and the second touch electrode are all of a stripe shape and arranged at different layers. the bonding terminals (1041, 1042) comprise a first bonding terminal and a second bonding terminal arranged between the first edge and the second edge; the touch electrodes (1021, 1022) comprise a first touch electrode and a second touch electrode; the signal transmission lines (1051, 1052) comprise a first signal transmission line connecting the first bonding terminal and the first touch electrode and a second signal transmission line connecting the second bonding terminal and the second touch electrode; the first touch electrode extends toward the touch area from the bonding terminals (1041, 1042), and the second touch electrode and the first touch electrode are crossed to each other, and the first signal transmission line and the first touch electrode are arranged in a same layer; the second signal transmission line comprises a first part extending along the first touch electrode and arranged in the touch area; characterized in that: the first part is arranged in a different layer from the second touch electrode; the second signal transmission line further comprises a second part arranged in the touch area and connected to the first part through a via hole, wherein the second part extends along the second touch electrode and is in a same layer with the second touch electrode and electrically connected to the second touch electrode; and the first touch electrode and the second touch electrode are all of a stripe shape and arranged at different layers. the first part is arranged in a different layer from the second touch electrode; the second signal transmission line further comprises a second part arranged in the touch area and connected to the first part through a via hole, wherein the second part extends along the second touch electrode and is in a same layer with the second touch electrode and electrically connected to the second touch electrode; and the first touch electrode and the second touch electrode are all of a stripe shape and arranged at different layers.