About Clements ' distinction between primary succession and secondary succession , Cowles wrote ( 1911 ) : Cowles ( 1911 ) wrote about clements ' ; the distinction between primary and secondary succession :
Mengenai perbezaan Clements antara penggantian primer dan penggantian sekunder, Cowles menulis (1911): Cowles (1911) menulis tentang clements'; perbezaan antara penggantian primer dan sekunder: