['it', 'apply', 'EmptyParameter', 'species', 'to']
['<PRP>', 'put-9.1-1', 'EmptyParameter', "Synset('biological_group.n.01')", 'to']
It can at least be applied to their own species.
<PRP> can at Synset('least.a.01') be applied to <PRP> own Synset('biological_group.n.01').