page_title: Utricularia
section_title: Distribution and habitat
hierarchical_section_title: Utricularia / Distribution and habitat
caption_attribution_description: Utricularia aurea
context_page_description: Utricularia, commonly and collectively called the bladderworts, is a genus of carnivorous plants consisting of approximately 233 species. They occur in fresh water and wet soil as terrestrial or aquatic species across every continent except Antarctica. Utricularia are cultivated for their flowers, which are often compared with those of snapdragons and orchids, especially amongst carnivorous plant enthusiasts.
  All Utricularia are carnivorous and capture small organisms by means of bladder-like traps. Terrestrial species tend to have tiny traps that feed on minute prey such as protozoa and rotifers swimming in water-saturated soil. The traps can range in size from 0.2 mm to 1.2 cm. Aquatic species, such as U. vulgaris, possess bladders that are usually larger and can feed on more substantial prey such as water fleas, nematodes and even fish fry, mosquito larvae and young tadpoles. Despite their small size, the traps are extremely sophisticated. In the active traps of the aquatic species, prey brush against trigger hairs connected to the trapdoor.
context_section_description: Utricularia can survive almost anywhere where there is fresh water for at least part of the year; only Antarctica and some oceanic islands have no native species. The greatest species diversity for the genus is seen in South America, with Australia coming a close second. In common with most carnivorous plants, they grow in moist soils which are poor in dissolved minerals, where their carnivorous nature gives them a competitive advantage; terrestrial varieties of Utricularia can frequently be found alongside representatives of the carnivorous genera–Sarracenia, Drosera and others–in very wet areas where continuously moving water removes most soluble minerals from the soil.
  About 80% of the species are terrestrial, and most inhabit waterlogged or wet soils, where their tiny bladders can be permanently exposed to water in the substrate. Frequently they will be found in marshy areas where the water table is very close to the surface. Most of the terrestrial species are tropical, although they occur worldwide.
  Approximately 20% of the species are aquatic. Most of these drift freely over the surface of ponds and other still, muddy-bottomed waters and only protrude above the surface when flowering, although a few species are lithophytic and adapted to rapidly moving streams or even waterfalls. The plants are usually found in acidic waters, but they are quite capable of growing in alkaline waters and would very likely do so were it not for the higher level of competition from other plants in such areas. Utricularia vulgaris is an aquatic species and grows into branching rafts with individual stolons up to one metre or longer in ponds and ditches throughout Eurasia.
  Some South American tropical species are epiphytes, and can be found growing in wet moss and spongy bark on trees in rainforests, or even in the watery leaf-rosettes of other epiphytes such as various Tillandsia (a type of bromeliad) species. Rosette-forming epiphytes such as U. nelumbifolia put out runners, searching for other nearby bromeliads to colonise.
  The plants are as highly adapted in their methods of surviving seasonally inclement conditions as they are in their structure and feeding habits. Temperate perennials can require a winter period in which they die back each year, and they will weaken in cultivation if they are not given it; tropical and warm-temperate species, on the other hand, require no dormancy. Floating bladderworts in cold temperate zones such as the UK and Siberia can produce winter buds called turions at the extremities of their stems: as the autumnal light fails and growth slows down, the main plant may rot away or be killed by freezing conditions, but the turions will separate and sink to the bottom of the pond to rest beneath the coming ice until the spring, when they will return to the surface and resume growth. Many Australian species will grow only during the wet season, reducing themselves to tubers only 10 mm long to wait out the dry season. Other species are annual, returning from seed each year.
caption: Utricularia aurea growing in a rice paddy in Thailand.
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