page_title: Moa
section_title: Surviving remains
hierarchical_section_title: Moa / Relationship with humans / Surviving remains
caption_attribution_description: Moa excavation in Kapua Swamp.
context_page_description: Moa were nine species of now-extinct flightless birds endemic to New Zealand. The two largest species, Dinornis robustus and Dinornis novaezelandiae, reached about 3.6 m in height with neck outstretched, and weighed about 230 kg while the smallest, the bush moa, was around the size of a turkey. Estimates of the Moa population when Polynesians settled New Zealand circa 1280, vary between 58,000 to c. 2.5 million.
  Moa belong to the order Dinornithiformes, traditionally placed in the ratite group. However, their closest relatives have been found by genetic studies to be the flighted South American tinamous, once considered to be a sister group to ratites. The nine species of moa were the only wingless birds lacking even the vestigial wings that all other ratites have. They were the largest terrestrial animals and dominant herbivores in New Zealand's forest, shrubland, and subalpine ecosystems until the arrival of the Māori, and were hunted only by the Haast's eagle. Moa extinction occurred around 200 years after human settlement primarily due to overhunting by the Māori.
context_section_description: Joel Polack, a trader who lived on the East Coast of the North Island from 1834 to 1837, recorded in 1838 that he had been shown "several large fossil ossifications" found near Mt Hikurangi. He was certain that these were the bones of a species of emu or ostrich, noting that "the Natives add that in times long past they received the traditions that very large birds had existed, but the scarcity of animal food, as well as the easy method of entrapping them, has caused their extermination". Polack further noted that he had received reports from Māori that a "species of Struthio" still existed in remote parts of the South Island.
  Dieffenbach also refers to a fossil from the area near Mt Hikurangi, and surmises that it belongs to "a bird, now extinct, called Moa (or Movie) by the natives". 'Movie' is the first transcribed name for the bird. In 1839, John W. Harris, a Poverty Bay flax trader who was a natural-history enthusiast, was given a piece of unusual bone by a Māori who had found it in a river bank. He showed the 15 cm (6 in) fragment of bone to his uncle, John Rule, a Sydney surgeon, who sent it to Richard Owen, who at that time was working at the Hunterian Museum at the Royal College of Surgeons in London.
  Owen puzzled over the fragment for almost four years. He established it was part of the femur of a big animal, but it was uncharacteristically light and honeycombed. Owen announced to a skeptical scientific community and the world that it was from a giant extinct bird like an ostrich, and named it Dinornis. His deduction was ridiculed in some quarters, but was proved correct with the subsequent discoveries of considerable quantities of moa bones throughout the country, sufficient to reconstruct skeletons of the birds.
  In July 2004, the Natural History Museum in London placed on display the moa bone fragment Owen had first examined, to celebrate 200 years since his birth, and in memory of Owen as founder of the museum.
  Since the discovery of the first moa bones in the late 1830s, thousands more have been found. They occur in a range of late Quaternary and Holocene sedimentary deposits, but are most common in three main types of site: caves, dunes, and swamps.
  Bones are commonly found in caves or tomo (the Maori word for doline or sinkhole, often used to refer to pitfalls or vertical cave shafts). The two main ways that the moa bones were deposited in such sites were birds that entered the cave to nest or escape bad weather, and subsequently died in the cave and birds that fell into a vertical shaft and were unable to escape. Moa bones (and the bones of other extinct birds) have been found in caves throughout New Zealand, especially in the limestone/marble areas of northwest Nelson, Karamea, Waitomo, and Te Anau.
  Moa bones and eggshell fragments sometimes occur in active coastal sand dunes, where they may erode from paleosols and concentrate in 'blowouts' between dune ridges. Many such moa bones antedate human settlement, although some originate from Maori midden sites, which frequently occur in dunes near harbours and river mouths (for example the large moa hunter sites at Shag River, Otago, and Wairau Bar, Marlborough).
  Densely intermingled moa bones have been encountered in swamps throughout New Zealand. The most well-known example is at Pyramid Valley in north Canterbury, where bones from at least 183 individual moa have been excavated, mostly by Roger Duff of Canterbury Museum. Many explanations have been proposed to account for how these deposits formed, ranging from poisonous spring waters to floods and wildfires. However, the currently accepted explanation is that the bones accumulated slowly over thousands of years, from birds that entered the swamps to feed and became trapped in the soft sediment.
caption: An excavation in Kapua Swamp, 1894
exif: {"Image Orientation": "Horizontal (normal)", "Image XResolution": "72", "Image YResolution": "72", "Image ResolutionUnit": "Pixels/Inch", "Image Software": "Adobe Photoshop CS3 Windows", "Image DateTime": "2012:01:13 11:54:36", "Image ExifOffset": "164", "Thumbnail Compression": "JPEG (old-style)", "Thumbnail XResolution": "72", "Thumbnail YResolution": "72", "Thumbnail ResolutionUnit": "Pixels/Inch", "Thumbnail JPEGInterchangeFormat": "302", "Thumbnail JPEGInterchangeFormatLength": "8450", "EXIF ColorSpace": "sRGB", "EXIF ExifImageWidth": "883", "EXIF ExifImageLength": "549"}