page_title: Tarsophlebiidae
section_title: Classification
hierarchical_section_title: Tarsophlebiidae / Classification
caption_attribution_description: English: Tarsophlebia minor (Odonata: Tarsophlebioptera: Tarsophlebiidae), Upper Jurassic, Solnhofen Plattenkalk, female specimen at Bürgermeister Müller Museum in Solnhofen (Germany) (ex coll. Wittmann), badly preserved venation, but visible hypertrophied ovipositor
context_page_description: The Tarsophlebiidae is an extinct family of medium-sized fossil odonates from the Upper Jurassic and Lower Cretaceous period of Eurasia. They are either the most basal member of the damsel-dragonfly grade within the stem group of Anisoptera, or the sister group of all Recent odonates. They are characterized by the basally open discoidal cell in both pairs of wings, very long legs, paddle-shaped male cerci, and a hypertrophied ovipositor in females.
context_section_description: The family was established by Handlirsch (1906) on the basis of the type genus Tarsophlebia Hagen, 1866.
  Tarsophlebiopsis mayi was recognized as valid by most authors until the most recent revision of the family by Fleck et al. (2004), who demonstrated that the holotype of this taxon most probably is only an aberrant specimen of Tarsophlebia eximia.
  The family contains 2 described genera with totally 9 valid species:
  Genus †Tarsophlebia Hagen, 1866 (type genus; synonym: Tarsophlebiopsis Tillyard, 1923)
  Diagnosis: cubito-anal areas of forewings and hindwings with four rows of cells or less; 16 or less postnodal crossveins; less than ten secondary antenodal crossveins in hindwing; and IR1 relatively short.
  Tarsophlebia eximia (Hagen, 1862) - (type species; type locality: Eichstätt, Bavaria, Germany; type horizon: Upper Jurassic, Malm zeta 2b, Lower Tithonian / 150 mya, Hybonotum Zone, Solnhofener Plattenkalk Formation; holotype: no. AS VI 44a,b at BSPG Munich Museum; size: forewing 37.1, hindwing 34.7 mm; synonyms: Tarsophlebia major Handlirsch, 1906, Tarsophlebia longissima Handlirsch, 1906, Tarsophlebiopsis mayi Tillyard, 1923)
  Tarsophlebia minor Fleck et al., 2004 - (type locality: Eichstätt, Bavaria, Germany; type horizon: Upper Jurassic, Malm zeta 2b, Lower Tithonian / 150 mya, Hybonotum Zone, Solnhofener Plattenkalk Formation; holotype: no. 55 at MCZ; size: hindwing 26 mm)
  Genus Turanophlebia Pritykina, 1968
  Diagnosis: wings with more dense venation; presence of more than 25 postnodal crossveins (11-16 in Tarsophlebia); six or more rows of cells between CuA and posterior hindwing margin (less than five rows in Tarsophlebia); more than 10 secondary antenodal crossveins in hindwing (less than 10 in Tarsophlebia); IR1 longer than in Tarsophlebia; presence of long secondary longitudinal not zigzagged veins in area between IR2 and RP2.
  Turanophlebia anglicana Fleck et al., 2004 - (type locality: Clockhouse (Butterley) Brickworks, England; type horizon: Lower Cretaceous, Upper Hauterivian / 130 mya, Lower Weald Clay; holotype: no. BMB 018531 at Booth Museum in Brighton / UK; size: hindwing 39.7 mm)
  Turanophlebia martynovi Pritykina, 1968 - (type species; type locality: Karatau-Mikhailovka, Chimkent region, Southern Kazakhstan; type horizon: Upper Jurassic, Callovian- Kimmeridgian or Oxfordian-Kimmeridgian / 163 mya, Karabastau Formation; holotype: no. PIN 2554/210 at Palaeontological Institute Moscow; size: hindwing 41.0 mm)
  Turanophlebia mongolica Fleck et al., 2004 - (type locality: Bon-Tsagaan, Bayanhongor Aimak, Central Mongolia; type horizon: Lower Cretaceous, Hauterivian-Barremian / 129 mya, Dzun-Bain Formation; holotype: no. PIN 3559/69 at Palaeontological Institute Moscow; size: hindwing 41.0 mm)
  Turanophlebia neckini (Martynov, 1927) - (type locality: Karatau-Karabastau, Chimkent region, Southern Kazakhstan; type horizon: Upper Jurassic, Callovian- Kimmeridgian or Oxfordian-Kimmeridgian / 163 mya, Karabastau Formation; holotype: no. PIN 2452/3 at Palaeontological Institute Moscow; size: hindwing 39.0 mm; originally described in the genus Tarsophlebia)
  Turanophlebia? sibirica Pritykina,1977 - (type locality: Baissa, Vitim River, Buryat Republik, Siberia / Transbaikalia, Russia; type horizon: Lower Cretaceous, Neocomian-Berriasian / 145-140 mya or Barremian-Aptian / 125 mya, Zaza Formation; holotype: no. PIN 1989/1258 at Palaeontological Institute Moscow; estimated size: hindwing 46.9, based on a comparison of the distance between wing base and end of MP, which is 24.7 mm in T. sibirica and 21.6 mm in T. martynovi that has a total hindwing length of 41.0 mm)
  Turanophlebia sinica Huang & Nel, 2009 - (type locality: Western Liaoning, PR China; type horizon: Lower Cretaceous, Barremian-Aptian / 125-121 mya, Yixian Formation; holotype: no. NIGP 148201 at Nanjing Institute of Geology and Palaeontology; size: forewing 49.5 and hindwing 45.5 mm)
  Turanophlebia vitimensis Fleck et al., 2004 - (type locality: right side of Vitim River downstream Romanovka village, Eravna district, Buryat Republic, Siberia / Transaikalia, Russ
caption: Tarsophlebia minor, Upper Jurassic, Solnhofen Plattenkalk, female with ovipositor, specimen at BMMS
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