page_title: CLE peptide
section_title: Signaling in the shoot apical meristem
hierarchical_section_title: CLE peptide / Signaling in the shoot apical meristem
context_page_description: CLE peptides are a group of peptides found in plants that are involved with cell signaling. Production is controlled by the CLE genes. Upon binding to a CLE peptide receptor in another cell, a chain reaction of events occurs, which can lead to various physiological and developmental processes. This signaling pathway is conserved in diverse land plants.
context_section_description: Meristematic cells give rise to various organs of the plant and keep the plant growing. There are two types of meristematic tissues 1) Apical Meristem 2) Lateral Meristem. The Apical Meristem is of two types; the shoot apical meristem (SAM) gives rise to organs like the leaves and flowers, while the root apical meristem (RAM) provides the meristematic cells for the future root growth. SAM and RAM cells divide rapidly and are considered indeterminate, in that they do not possess any defined end status. In that sense, the meristematic cells are frequently compared to the stem cells in animals, which have an analogous behavior and function. Within plants SAM cells play a major role in the overall growth and development, this is due to the fact that all cells making up the major parts of the plant come from the shoot apical meristem (SAM). There are three different important area found within the SAM and they include the central zone, the peripheral zone), and the rib meristem. Each of these areas play an important in the production of new stem cells within the SAM. All SAMs are usually dome shaped and have structures that are layered and are described as the tunica and corpus. CLV3 plays an important role in regulating the production of stem cells within the Central Zone region of the (SAM), this is also true for the cell promoting WUSCHEL (WUS) gene. The combination of these two genes regulates stem cell production by WUS negatively or positively regulating the production of stem cells by controlling the CLV3 gene.;
caption: Tunica-Corpus model of the apical meristem (growing tip). The epidermal (L1) and subepidermal (L2) layers form the outer layers called the tunica. The inner L3 layer is called the corpus. Cells in the L1 and L2 layers divide in a sideways fashion, which keeps these layers distinct, whereas the L3 layer divides in a more random fashion.
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