page_title: Apatosaurus
section_title: Neck posture
hierarchical_section_title: Apatosaurus / Palaeobiology / Neck posture
caption_attribution_description: Русский: Апатозавр English: Illustration of Apatosaurus louisae
context_page_description: Apatosaurus is a genus of herbivorous sauropod dinosaur that lived in North America during the Late Jurassic period. Othniel Charles Marsh described and named the first-known species, A. ajax, in 1877, and a second species, A. louisae, was discovered and named by William H. Holland in 1916. Apatosaurus lived about 152 to 151 million years ago, during the late Kimmeridgian to early Tithonian age, and are now known from fossils in the Morrison Formation of modern-day Colorado, Oklahoma, New Mexico, Wyoming, and Utah in the United States. Apatosaurus had an average length of 21–22.8 m, and an average mass of 16.4–22.4 t. A few specimens indicate a maximum length of 11–30% greater than average and a mass of 32.7–72.6 t.
  The cervical vertebrae of Apatosaurus are less elongated and more heavily constructed than those of Diplodocus, a diplodocid like Apatosaurus, and the bones of the leg are much stockier despite being longer, implying that Apatosaurus was a more robust animal. The tail was held above the ground during normal locomotion. Apatosaurus had a single claw on each forelimb and three on each hindlimb.
context_section_description: Diplodocids like Apatosaurus are often portrayed with their necks held high up in the air, allowing them to browse on tall trees. Some studies state diplodocid necks were less flexible than previously believed, because the structure of the neck vertebrae would not have allowed the neck to bend far upwards, and that sauropods like Apatosaurus were adapted to low browsing or ground feeding.
  Other studies by Taylor find that all tetrapods appear to hold their necks at the maximum possible vertical extension when in a normal, alert posture; they argue the same would hold true for sauropods barring any unknown, unique characteristics that set the soft tissue anatomy of their necks apart from that of other animals. Apatosaurus, like Diplodocus, would have held its neck angled upwards with the head pointing downwards in a resting posture. Kent Stevens and Michael Parrish (1999 and 2005) state Apatosaurus had a great feeding range; its neck could bend into a U-shape laterally. The neck's range of movement would have also allowed the head to feed at the level of the feet.
  Matthew Cobley et al. (2013) dispute this, finding that large muscles and cartilage would have limited movement of the neck. They state the feeding ranges for sauropods like Diplodocus were smaller than previously believed, and the animals may have had to move their whole bodies around to better access areas where they could browse vegetation. As such, they might have spent more time foraging to meet their minimum energy needs. The conclusions of Cobley et al. are disputed by Taylor, who analyzed the amount and positioning of intervertebral cartilage to determine the flexibility of the neck of Apatosaurus and Diplodocus. He found that the neck of Apatosaurus was very flexible.
caption: Artistic interpretation of an individual of A. louisae arching its neck down to drink
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