page_title: Giant panda
section_title: Diet
hierarchical_section_title: Giant panda / Ecology / Diet
caption_attribution_description: Pandas!! (GIANT PANDA/WOLONG/SICHUAN/CHINA)
context_page_description: The giant panda, also known as the panda bear or simply the panda, is a bear native to south central China. It is characterised by large, black patches around its eyes, over the ears, and across its round body. The name "giant panda" is sometimes used to distinguish it from the red panda, a neighboring musteloid. Though it belongs to the order Carnivora, the giant panda is a folivore, with bamboo shoots and leaves making up more than 99% of its diet. Giant pandas in the wild will occasionally eat other grasses, wild tubers, or even meat in the form of birds, rodents, or carrion. In captivity, they may receive honey, eggs, fish, yams, shrub leaves, oranges, or bananas along with specially prepared food.
  The giant panda lives in a few mountain ranges in central China, mainly in Sichuan, but also in neighbouring Shaanxi and Gansu. As a result of farming, deforestation, and other development, the giant panda has been driven out of the lowland areas where it once lived, and it is a conservation-reliant vulnerable species. A 2007 report showed 239 pandas living in captivity inside China and another 27 outside the country.
context_section_description: Despite its taxonomic classification as a carnivoran, the giant panda's diet is primarily herbivorous, consisting almost exclusively of bamboo. However, the giant panda still has the digestive system of a carnivore, as well as carnivore-specific genes, and thus derives little energy and little protein from consumption of bamboo. Its ability to digest cellulose is ascribed to the microbes in its gut. Pandas are born with sterile intestines and require bacteria obtained from their mother's feces to digest vegetation. The giant panda is a highly specialised animal with unique adaptations, and has lived in bamboo forests for millions of years.
  The average giant panda eats as much as 9 to 14 kg (20 to 30 lb) of bamboo shoots a day to compensate for the limited energy content of its diet. Ingestion of such a large quantity of material is possible and necessary because of the rapid passage of large amounts of indigestible plant material through the short, straight digestive tract. It is also noted, however, that such rapid passage of digesta limits the potential of microbial digestion in the gastrointestinal tract, limiting alternative forms of digestion. Given this voluminous diet, the giant panda defecates up to 40 times a day. The limited energy input imposed on it by its diet has affected the panda's behavior. The giant panda tends to limit its social interactions and avoids steeply sloping terrain to limit its energy expenditures.
  Two of the panda's most distinctive features, its large size and round face, are adaptations to its bamboo diet. Anthropologist Russell Ciochon observed: "[much] like the vegetarian gorilla, the low body surface area to body volume [of the giant panda] is indicative of a lower metabolic rate. This lower metabolic rate and a more sedentary lifestyle allows the giant panda to subsist on nutrient poor resources such as bamboo." Similarly, the giant panda's round face is the result of powerful jaw muscles, which attach from the top of the head to the jaw. Large molars crush and grind fibrous plant material.
  The morphological characteristics of extinct relatives of the giant panda suggest that while the ancient giant panda was omnivorous 7 million years ago (mya), it only became herbivorous some 2–2.4 mya with the emergence of A. microta. Genome sequencing of the giant panda suggests that the dietary switch could have initiated from the loss of the sole T1R1/T1R3 umami taste receptor, resulting from two frameshift mutations within the T1R1 exons. Umami taste corresponds to high levels of glutamate as found in meat and may have thus altered the food choice of the giant panda. Although the pseudogenisation of the umami taste receptor in Ailuropoda coincides with the dietary switch to herbivory, it is likely a result of, and not the reason for, the dietary change. The mutation time for the T1R1 gene in the giant panda is estimated to 4.2 mya while fossil evidence indicates bamboo consumption in the giant panda species at least 7 mya, signifying that although complete herbivory occurred around 2 mya, the dietary switch was initiated prior to T1R1 loss-of-function.
  Pandas eat any of 25 bamboo species in the wild, such as Fargesia dracocephala and Fargesia rufa. Only a few bamboo species are widespread at the high altitudes pandas now inhabit. Bamboo leaves contain the highest protein levels; stems have less.
  Because of the synchronous flowering, death, and regeneration of all bamboo within a species, the giant panda must have at least two different species available in its range to avoid starvation. While primarily herbivorous, the giant panda still retains decidedly ursine teeth and will eat meat, fish, and eggs when available. In captivity, zoos typically maintain the giant panda's bamboo diet, though some will provide specially formulated biscuits or other dietary supplements.
  Pandas will travel between different habitats if they need to, so they can get the nutrients that they need and to balance their diet for reproduction. For six years, scientists studied six pandas tagged with GPS collars at the Fo
caption: Pandas eating bamboo.
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