page_title: Tapejara (pterosaur)
section_title: Species and classification
hierarchical_section_title: Tapejara (pterosaur) / Species and classification
caption_attribution_description: English: Cast of Tapejara wellnhoferi - Pterosaurs Flight in the Age of Dinosaurs
context_page_description: Tapejara is a genus of Brazilian pterosaur from the Cretaceous Period. Tapejara crests consisted of a semicircular crest over the snout, and a bony prong which extended back behind the head.
context_section_description: The type species and only one currently recognized as valid by most researchers, is T. wellnhoferi. The specific name honours German paleontologist Peter Wellnhofer. Two larger species, originally named Tapejara imperator and Tapejara navigans, were later classified in the genus Tapejara. However, several studies have shown that T. imperator and T. navigans are significantly different from T. wellnhoferi and therefore were reclassified into new genera. The species T. imperator was given its own genus, Tupandactylus, by Kellner and Campos. Unwin and Martill found that T. imperator and T. navigans belong in the same genus, and named them Ingridia imperator and I. navigans, respectively. This genus name honoured Wellnhofer's late wife Ingrid. 
  Because Tupandactylus was named first, it retained priority over the name Ingridia. To complicate matters, both the name Tupandactylus and Ingridia used the former Tapejara imperator as their type species. The scientists who described Tupandactylus did not name a Tupandactylus navigans (but instead suggested it was synonymous to Tupandactylus imperator), and Tapejara navigans was not formally reclassified as a distinct species of Tupandactylus until 2011.
   
  The cladogram below follows the 2014 analysis Andres and colleagues.
caption: Cast
exif: {"Image Make": "NIKON CORPORATION", "Image Model": "NIKON D7100", "Image XResolution": "600", "Image YResolution": "600", "Image ResolutionUnit": "Pixels/Inch", "Image Software": "Adobe Photoshop Lightroom 5.7.1 (Windows)", "Image DateTime": "2018:11:06 23:35:38", "Image ExifOffset": "218", "Thumbnail Compression": "JPEG (old-style)", "Thumbnail XResolution": "72", "Thumbnail YResolution": "72", "Thumbnail ResolutionUnit": "Pixels/Inch", "Thumbnail JPEGInterchangeFormat": "978", "Thumbnail JPEGInterchangeFormatLength": "15999", "EXIF ExposureTime": "1/30", "EXIF FNumber": "19/5", "EXIF ExposureProgram": "Unidentified", "EXIF ISOSpeedRatings": "1250", "EXIF SensitivityType": "Recommended Exposure Index", "EXIF ExifVersion": "0230", "EXIF DateTimeOriginal": "2018:08:09 13:21:36", "EXIF DateTimeDigitized": "2018:08:09 13:21:36", "EXIF ShutterSpeedValue": "4906891/1000000", "EXIF ApertureValue": "3851999/1000000", "EXIF ExposureBiasValue": "0", "EXIF MaxApertureValue": "39/10", "EXIF MeteringMode": "Pattern", "EXIF LightSource": "Unknown", "EXIF Flash": "Flash did not fire, compulsory flash mode", "EXIF FocalLength": "22", "EXIF SubSecTimeOriginal": "80", "EXIF SubSecTimeDigitized": "80", "EXIF FocalPlaneXResolution": "83841555/32768", "EXIF FocalPlaneYResolution": "83841555/32768", "EXIF FocalPlaneResolutionUnit": "3", "EXIF SensingMethod": "One-chip color area", "EXIF FileSource": "Digital Camera", "EXIF SceneType": "Directly Photographed", "EXIF CVAPattern": "[2, 0, 2, 0, 0, 1, 1, 2]", "EXIF CustomRendered": "Normal", "EXIF ExposureMode": "Auto Exposure", "EXIF WhiteBalance": "Auto", "EXIF DigitalZoomRatio": "1", "EXIF FocalLengthIn35mmFilm": "33", "EXIF SceneCaptureType": "Standard", "EXIF GainControl": "High gain up", "EXIF Contrast": "Normal", "EXIF Saturation": "Normal", "EXIF Sharpness": "Normal", "EXIF SubjectDistanceRange": "0", "EXIF BodySerialNumber": "2534693", "EXIF LensSpecification": "[18, 55, 7/2, 28/5]", "EXIF LensModel": "18.0-55.0 mm f/3.5-5.6"}