page_title: Sexual intercourse
section_title: Other animals
hierarchical_section_title: Sexual intercourse / Other animals
context_page_description: Sexual intercourse is sexual activity typically involving the insertion and thrusting of the penis into the vagina for sexual pleasure, reproduction, or both. This is also known as vaginal intercourse or vaginal sex. Other forms of penetrative sexual intercourse include anal sex, oral sex, fingering, and penetration by use of a dildo. These activities involve physical intimacy between two or more individuals and are usually used among humans solely for physical or emotional pleasure and can contribute to human bonding.
  There are different views on what constitutes sexual intercourse or other sexual activity, which can impact on views on sexual health. Although sexual intercourse, particularly the variant coitus, generally denotes penile–vaginal penetration and the possibility of creating offspring, it also commonly denotes penetrative oral sex and penile–anal sex, especially the latter. It usually encompasses sexual penetration, while non-penetrative sex has been labeled "outercourse", but non-penetrative sex may also be considered sexual intercourse. Sex, often a shorthand for sexual intercourse, can mean any form of sexual activity.
context_section_description: In zoology, copulation often means the process in which a male introduces sperm into the female's body, especially directly into her reproductive tract. Spiders have separate male and female sexes. Before mating and copulation, the male spider spins a small web and ejaculates on to it. He then stores the sperm in reservoirs on his large pedipalps, from which he transfers sperm to the female's genitals. The females can store sperm indefinitely.
  Many animals that live in water use external fertilization, whereas internal fertilization may have developed from a need to maintain gametes in a liquid medium in the Late Ordovician epoch. Internal fertilization with many vertebrates (such as reptiles, some fish, and most birds) occur via cloacal copulation (see also hemipenis), while mammals copulate vaginally, and many basal vertebrates reproduce sexually with external fertilization.
  For primitive insects, the male deposits spermatozoa on the substrate, sometimes stored within a special structure; courtship involves inducing the female to take up the sperm package into her genital opening, but there is no actual copulation. In groups that have reproduction similar to spiders, such as dragonflies, males extrude sperm into secondary copulatory structures removed from their genital opening, which are then used to inseminate the female. In dragonflies, it is a set of modified sternites on the second abdominal segment. In advanced groups of insects, the male uses its aedeagus, a structure formed from the terminal segments of the abdomen, to deposit sperm directly (though sometimes in a capsule called a spermatophore) into the female's reproductive tract.
  Bonobos, chimpanzees and dolphins are species known to engage in heterosexual behaviors even when the female is not in estrus, which is a point in her reproductive cycle suitable for successful impregnation. These species are also known to engage in same-sex sexual behaviors. In these animals, the use of sexual intercourse has evolved beyond reproduction to apparently serve additional social functions (such as bonding).
caption: A pair of lions copulating in the Maasai Mara, Kenya
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