page_title: Cecidomyiidae
section_title: Description
hierarchical_section_title: Cecidomyiidae / Description
caption_attribution_description: English: Gall midge (Vitisiella) larva in gall in Vitis. Josie Porter Farm - Stroud Township, Monroe County, Pennsylvania, USA. https://bugguide.net/node/view/659410
context_page_description: Cecidomyiidae is a family of flies known as gall midges or gall gnats. As the name implies, the larvae of most gall midges feed within plant tissue, creating abnormal plant growths called galls. Cecidomyiidae are very fragile small insects usually only 2–3 mm in length; many are less than 1 mm long. They are characterised by hairy wings, unusual in the order Diptera, and have long antennae. Cecidomyiids are also known for the strange phenomenon of paedogenesis in which the larval stage reproduces without maturing first. In some species, the daughter larvae consume the mother, while in others, reproduction occurs later on in the egg or pupa.
  More than 6,000 species and 783 genera are described worldwide, but since 1,100 are from well-studied North America, this is certainly an underestimate, and a recent DNA barcoding study estimated the fauna of Canada alone to be in excess of 16,000 species, hinting at a staggering global count of over 1 million cecidomyiid species that have yet to be described, which would make it the most speciose single family in the entire animal kingdom.
context_section_description: Cecidomyiidae are minute to small (0.5–3.0 mm), rarely larger (up to 8 mm, wing length 15 mm) flies with a delicate appearance. Except for a few genera with reduced wings, the eyes are holoptic. The mouthparts are reduced. Cecidomyiid antennae are notably long, with 12–14 segments, (sometimes fewer and up to 40 in some genera). The antennal segments either consist of a basal thickening and petiole or they are binodal, with a proximal node, an intermediate petiole and a distal node. Basal, medial, and apical whorls of hairs occur on the antennal segments. In some species, whorls of loop-shaped sensory filaments are also found, the basal or medial one sometimes being reduced. Some gall flies have only one (basal) whorl of hairs on the antennal segments, and the sensoria (transparent sensory appendages) differ in size and shape. The filaments are thread-like in the Porricondylinae and in all the Cecidomyiinae and take the form of long loops in the supertribe Cecidomyiidi). Ocelli are present only in the Lestremiinae. The wings are usually clear, rarely patterned. The wing bears microtrichia, often as scales, and some species have macrotrichia. The number of longitudinal veins is reduced. Only veins R1, R4+5, M3+4 and Cu1 are well developed in most species. The medial veins M1 and M2 are developed only in primitive groups, and the costa usually has a break just beyond vein R5. The legs are long and slender, without apical bristles. 
  The genitalia of males consist of gonocoxites, gonostyles, aedeagus, and tergites 9 and 10. Lower (in the evolutionary sense) gall flies often have sclerotized parameres and a more or less transparent plate (the tegmen) located above theaedeagus-the tegmen. In higher gall flies, the parameres and tegmen are not developed. In these, instead, close to the aedeagus, is a triangular basal outgrowth of the gonocoxites called the gonosterna. Supporting structures called apodema are located near the base of the genitalia in males; these are often equipped with two outgrowths. The ovipositor is short, lamelliform, or long, mobile, and in some species, acicular.
  The larva is peripneustic. The head is tiny, cone-shaped, and has two posterolateral extensions. The mouthparts are reduced, with minute styliform mandibles. The relatively prominent antennae are two-segmented. Integumental setae or papillae are important in taxonomy since they are constant in number within groups. The prothorax has sclerotized sternal spatula (most). The anus is terminal in the Lestremiinae and paedogenetic in the Porricondylinae and ventral in other groups. The pupa is exarate (in a few species it is enclosed within the last instar larval integument). The anterior spiracle and anterior angle of antennal bases is prominent (most).
caption: Vitisiella larva
exif: {"Image Make": "Canon", "Image Model": "Canon PowerShot S3 IS", "Image Orientation": "Horizontal (normal)", "Image XResolution": "180", "Image YResolution": "180", "Image ResolutionUnit": "Pixels/Inch", "Image DateTime": "2012:06:17 20:04:47", "Image YCbCrPositioning": "Centered", "Image ExifOffset": "186", "Thumbnail Compression": "JPEG (old-style)", "Thumbnail XResolution": "180", "Thumbnail YResolution": "180", "Thumbnail ResolutionUnit": "Pixels/Inch", "Thumbnail JPEGInterchangeFormat": "2948", "Thumbnail JPEGInterchangeFormatLength": "3177", "EXIF ExposureTime": "1/40", "EXIF FNumber": "8", "EXIF ExifVersion": "0220", "EXIF DateTimeOriginal": "2012:06:17 20:04:47", "EXIF DateTimeDigitized": "2012:06:17 20:04:47", "EXIF ComponentsConfiguration": "YCbCr", "EXIF CompressedBitsPerPixel": "5", "EXIF ShutterSpeedValue": "85/16", "EXIF ApertureValue": "6", "EXIF ExposureBiasValue": "0", "EXIF MaxApertureValue": "29/8", "EXIF MeteringMode": "CenterWeightedAverage", "EXIF Flash": "Flash did not fire, compulsory flash mode", "EXIF FocalLength": "72", "EXIF FlashPixVersion": "0100", "EXIF ColorSpace": "sRGB", "EXIF ExifImageWidth": "2816", "EXIF ExifImageLength": "2112", "Interoperability InteroperabilityIndex": "R98", "Interoperability InteroperabilityVersion": "[48, 49, 48, 48]", "Interoperability RelatedImageWidth": "2816", "Interoperability RelatedImageLength": "2112", "EXIF InteroperabilityOffset": "2774", "EXIF FocalPlaneXResolution": "112640/9", "EXIF FocalPlaneYResolution": "2112000/169", "EXIF FocalPlaneResolutionUnit": "2", "EXIF SensingMethod": "One-chip color area", "EXIF FileSource": "Digital Camera", "EXIF CustomRendered": "Normal", "EXIF ExposureMode": "Auto Exposure", "EXIF WhiteBalance": "Manual", "EXIF DigitalZoomRatio": "1", "EXIF SceneCaptureType": "Standard", "EXIF OffsetSchema": "2"}