page_title: Guadalupian
section_title: Biodiversity
hierarchical_section_title: Guadalupian / Biodiversity
caption_attribution_description: English: Onthogenetic stages of anteosaur dinocephalian Titanophoneus potens- young individual (1), described as T. potens and adult (2) described as Doliosauriscus ("Doliosaurus") yanshinovi English: Онтогенетические стадии диноцефала-антеозавра Titanophoneus potens- молодая особь (1), описанная как T. potens и взрослая (2) описанная как Doliosauriscus ("Doliosaurus") yanshinovi
context_page_description: The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and date between 272.3 ± 0.5 – 259.8 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson’s Extinction and a significant mass extinction called the end-Capitanian extinction event.
  The Guadalupian was previously known as the Middle Permian.
context_section_description: Therapsids became the dominant land animals in Guadalupian, displacing the pelycosaurs.  Therapsids evolved from a group of pelycosaurs called sphenacodonts.  Therapsida consists of four major clades: the dinocephalians, the herbivorous anomodonts, the carnivorous biarmosuchians, and the mostly carnivorous theriodonts.  After a brief burst of evolutionary diversity, the dinocephalians died out in the later Guadalupian.
  A mass extinction occurred 273 million years ago in the early Guadalupian before the larger Permian–Triassic extinction event.
  This extinction was originally called Olson's Gap because it was thought to be a problem in preservation of fossils. Since the 1990s it has been renamed Olson’s Extinction. This extinction event occurred near the beginning of the epoch and led to an extended period of low diversity when two-thirds of terrestrial vertebrate life was lost worldwide. Global diversity rose dramatically by the end probably the result of disaster taxa filling empty guilds, only to fall again when the end-Guadalupian event caused a diversity drop in the Wuchiapingian.
  There is no agreed cause for the Olson's Extinction. Climate change may be a possible cause. Extreme environments were observed from the Permian of Kansas which resulted from a combination of hot climate and acidic waters particularly coincident with Olson’s Extinction. Whether this climate change was a result of Earth's natural processes or exacerbated by another event is unknown.
caption: Titanophoneus, top of the food chain in the Guadalupian
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