page_title: Imprinting (psychology)
section_title: Filial imprinting
hierarchical_section_title: Imprinting (psychology) / Filial imprinting
caption_attribution_description: Christian Moullec flying with geese in Dahlem, Germany
context_page_description: In psychology and ethology, imprinting is any kind of phase-sensitive learning that is rapid and apparently independent of the consequences of behaviour. It was first used to describe situations in which an animal or person learns the characteristics of some stimulus, which is therefore said to be "imprinted" onto the subject. Imprinting is hypothesized to have a critical period.
context_section_description: The best-known form of imprinting is filial imprinting, in which a young animal narrows its social preferences to an object (typically a parent) as a result of exposure to that object. It is most obvious in nidifugous birds, which imprint on their parents and then follow them around. It was first reported in domestic chickens, by Sir Thomas More in 1516 as described in his treatise Utopia, 350 years earlier than by the 19th-century amateur biologist Douglas Spalding. It was rediscovered by the early ethologist Oskar Heinroth, and studied extensively and popularized by his disciple Konrad Lorenz working with greylag geese.
  Lorenz demonstrated how incubator-hatched geese would imprint on the first suitable moving stimulus they saw within what he called a "critical period" between 13–16 hours shortly after hatching. For example, the goslings would imprint on Lorenz himself (to be more specific, on his wading boots), and he is often depicted being followed by a gaggle of geese who had imprinted on him. Lorenz also found that the geese could imprint on inanimate objects. In one notable experiment, they followed a box placed on a model train in circles around the track.  Filial imprinting is not restricted to non-human animals that are able to follow their parents, however.
  The filial imprinting of birds was a primary technique used to create the movie Winged Migration (Le Peuple Migrateur), which contains a great deal of footage of migratory birds in flight. The birds imprinted on handlers, who wore yellow jackets and honked horns constantly. The birds were then trained to fly along with a variety of aircraft, primarily ultralights.
  The Italian hang-glider pilot Angelo d'Arrigo extended this technique. D'Arrigo noted that the flight of a non-motorised hang-glider is very similar to the flight patterns of migratory birds; both use updrafts of hot air (thermal currents) to gain altitude that then permits soaring flight over distance. He used this to reintroduce threatened species of raptors. Because birds hatched in captivity have no mentor birds to teach them traditional migratory routes, D'Arrigo hatched chicks under the wing of his glider and imprinted on him. Then, he taught the fledglings to fly and to hunt. The young birds followed him not only on the ground (as with Lorenz) but also in the air as he took the path of various migratory routes. He flew across the Sahara and over the Mediterranean Sea to Sicily with eagles, from Siberia to Iran (5,500 km) with a flock of Siberian cranes, and over Mount Everest with Nepalese eagles. In 2006, he worked with a condor in South America.
  In a similar project, orphaned Canada geese were trained to their normal migration route by the Canadian ultralight enthusiast Bill Lishman, as shown in the fact-based movie drama Fly Away Home.
  Chicks of domestic chickens prefer to be near large groups of objects that they have imprinted on. This behaviour was used to determine that very young chicks of a few days old have rudimentary counting skills. In a series of experiments, they were made to imprint on plastic balls and could figure out which of two groups of balls hidden behind screens had the most balls.
  American coot mothers have the ability to recognize their chicks by imprinting on cues from the first chick that hatches. This allows mothers to distinguish their chicks from parasitic chicks.
  The peregrine falcon has also been known to imprint on specific structures for their breeding grounds such as cliff sides and bridges and thus will favour that location for breeding.
caption: Imprinted geese and cranes flying with an ultralight aircraft
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