page_title: List of non-avian dinosaur species preserved with evidence of feathers
hierarchical_section_title: List of non-avian dinosaur species preserved with evidence of feathers
caption_attribution_description: English: Cast of the fossil dromaeosaur specimen NGMC 91 (nicknamed "Dave", cf. Sinornithosaurus), at the American Museum of Natural History in New York.
context_page_description: Several non-avian dinosaurs were feathered. Direct evidence of feathers exists for the following species, listed in the order currently accepted evidence was first published. In all examples, the evidence described consists of feather impressions, except those genera inferred to have had feathers based on skeletal or chemical evidence, such as the presence of quill knobs or a pygostyle.
  Ostromia crassipes
  Avimimus portentosus
  Sinosauropteryx prima
  Fulicopus lyellii, an ichnotaxon, possible squatting Dilophosaurus or similar
  Protarchaeopteryx robusta
  GMV 2124
  Caudipteryx zoui
  Rahonavis ostromi
  Shuvuuia deserti
  Beipiaosaurus inexpectus
  Sinornithosaurus millenii
  Caudipteryx dongi
  Caudipteryx sp.
  Microraptor zhaoianus
  Nomingia gobiensis
  Psittacosaurus sp.?
  Scansoriopteryx heilmanni
  IVPP V13476
  Yixianosaurus longimanus
  Dilong paradoxus
  Pedopenna daohugouensis
  Jinfengopteryx elegans
  Juravenator starki
  Sinocalliopteryx gigas
  Velociraptor mongoliensis
  Epidexipteryx hui
  Similicaudipteryx yixianensis
  Anchiornis huxleyi
  Tianyulong confuciusi?
  Incisivosaurus sp.
  Concavenator corcovatus?
  Xiaotingia zhengi
  Yutyrannus huali
  Sciurumimus albersdoerferi
  Ornithomimus edmontonicus
context_section_description: Several non-avian dinosaurs were feathered. Direct evidence of feathers exists for the following species, listed in the order currently accepted evidence was first published. In all examples, the evidence described consists of feather impressions, except those genera inferred to have had feathers based on skeletal or chemical evidence, such as the presence of quill knobs (the anchor points for wing feathers on the forelimb) or a pygostyle (the fused vertebrae at the tail tip which often supports large feathers).
  Ostromia crassipes (1970; possibly avialan)
  Avimimus portentosus (inferred 1987: ulnar ridge)
  Sinosauropteryx prima (1996)
  Fulicopus lyellii, an ichnotaxon, possible squatting Dilophosaurus or similar (1996)
  Protarchaeopteryx robusta (1997)
  GMV 2124 (1997)
  Caudipteryx zoui (1998)
  Rahonavis ostromi (inferred 1998: quill knobs; possibly avialan)
  Shuvuuia deserti (1999)
  Beipiaosaurus inexpectus (1999)
  Sinornithosaurus millenii (1999)
  Caudipteryx dongi (2000)
  Caudipteryx sp. (2000)
  Microraptor zhaoianus (2000)
  Nomingia gobiensis (inferred 2000: pygostyle)
  Psittacosaurus sp.? (2002)
  Scansoriopteryx heilmanni (2002; possibly avialan)
  IVPP V13476 (2003)
  Yixianosaurus longimanus (2003; possibly avialan)
  Dilong paradoxus (2004)
  Pedopenna daohugouensis (2005; possibly avialan)
  Jinfengopteryx elegans (2005)
  Juravenator starki (2006)
  Sinocalliopteryx gigas (2007)
  Velociraptor mongoliensis (inferred 2007: quill knobs)
  Epidexipteryx hui (2008; possibly avialan)
  Similicaudipteryx yixianensis (inferred 2008: pygostyle)
  Anchiornis huxleyi (2009; possibly avialan)
  Tianyulong confuciusi? (2009)
  Incisivosaurus sp. (2010)
  Concavenator corcovatus? (inferred 2010: quill knobs?)
  Xiaotingia zhengi (2011; possibly avialan)
  Yutyrannus huali (2012)
  Sciurumimus albersdoerferi (2012)
  Ornithomimus edmontonicus (2012)
  Ningyuansaurus wangi (2012)
  Eosinopteryx brevipenna (2013; possibly avialan)
  Jianchangosaurus yixianensis (2013)
  Aurornis xui (2013; possibly avialan)
  Changyuraptor yangi (2014)
  Kulindadromeus zabaikalicus? (2014)
  Citipati osmolskae (inferred 2014: pygostyle)
  Conchoraptor gracilis (inferred 2014: pygostyle)
  Deinocheirus mirificus? (inferred 2014: pygostyle)
  Yi qi (2015; possibly avialan)
  Ornithomimus sp. (2015)
  Zhenyuanlong suni (2015)
  Dakotaraptor steini (inferred 2015: quill knobs)
  Apatoraptor pennatus (inferred 2016: quill knobs)
  DIP-V-15103 (2016; possibly avialan)
  Jianianhualong tengi (2017)
  Serikornis sungei (2017; possibly avialan)
  Caihong juji (2018; possibly avialan)
  Xingtianosaurus ganqi (2019)
  Ambopteryx longibrachium (2019; possibly avialan)
  Wulong bohaiensis (2020)
  Chirostenotes pergracilis? (2020)
  Dineobellator notohesperus (inferred 2020: quill knobs)
  Note that the filamentous structures in some ornithischian dinosaurs (Psittacosaurus, Tianyulong and Kulindadromeus) and the pycnofibres found in some pterosaurs may or may not be homologous with the feathers of theropods.
caption: Fossil cast of a Sinornithosaurus millenii
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