page_title: Alfred Russel Wallace
section_title: Biogeography and ecology
hierarchical_section_title: Alfred Russel Wallace / Other scientific contributions / Biogeography and ecology
caption_attribution_description: English: The original drawing of the 'Wallace Line' showing the division between Asian and Australasian flora and fauna reproduced at the internet archive
context_page_description: Alfred Russel Wallace OM FRS was a British naturalist, explorer, geographer, anthropologist, biologist and illustrator. He is best known for independently conceiving the theory of evolution through natural selection; his paper on the subject was jointly published with some of Charles Darwin's writings in 1858. This prompted Darwin to publish On the Origin of Species.
  Like Darwin, Wallace did extensive fieldwork; first in the Amazon River basin, and then in the Malay Archipelago, where he identified the faunal divide now termed the Wallace Line, which separates the Indonesian archipelago into two distinct parts: a western portion in which the animals are largely of Asian origin, and an eastern portion where the fauna reflect Australasia.
  He was considered the 19th century's leading expert on the geographical distribution of animal species and is sometimes called the "father of biogeography". Wallace was one of the leading evolutionary thinkers of the 19th century and made many other contributions to the development of evolutionary theory besides being co-discoverer of natural selection.
context_section_description: In 1872, at the urging of many of his friends, including Darwin, Philip Sclater, and Alfred Newton, Wallace began research for a general review of the geographic distribution of animals. Initial progress was slow, in part because classification systems for many types of animals were in flux. He resumed the work in earnest in 1874 after the publication of a number of new works on classification. Extending the system developed by Sclater for birds—which divided the earth into six separate geographic regions for describing species distribution—to cover mammals, reptiles and insects as well, Wallace created the basis for the zoogeographic regions still in use today. He discussed all of the factors then known to influence the current and past geographic distribution of animals within each geographic region.
  These factors included the effects of the appearance and disappearance of land bridges (such as the one currently connecting North America and South America) and the effects of periods of increased glaciation. He provided maps showing factors, such as elevation of mountains, depths of oceans, and the character of regional vegetation, that affected the distribution of animals. He also summarised all the known families and genera of the higher animals and listed their known geographic distributions. The text was organised so that it would be easy for a traveller to learn what animals could be found in a particular location. The resulting two-volume work, The Geographical Distribution of Animals, was published in 1876 and served as the definitive text on zoogeography for the next 80 years.
  The book included evidence from the fossil record to discuss the processes of evolution and migration that had led to the geographical distribution of modern species. For example, he discussed how fossil evidence showed that tapirs had originated in the Northern Hemisphere, migrating between North America and Eurasia and then, much more recently, to South America after which the northern species became extinct, leaving the modern distribution of two isolated groups of tapir species in South America and Southeast Asia. Wallace was very aware of, and interested in, the mass extinction of megafauna in the late Pleistocene. In The Geographical Distribution of Animals (1876) he wrote, "We live in a zoologically impoverished world, from which all the hugest, and fiercest, and strangest forms have recently disappeared". He added that he believed the most likely cause for the rapid extinctions was glaciation, but by the time he wrote World of Life (1911) he had come to believe those extinctions were "due to man's agency".
  In 1880, Wallace published the book Island Life as a sequel to The Geographical Distribution of Animals. It surveyed the distribution of both animal and plant species on islands. Wallace classified islands into oceanic and two types of continental islands.
  Oceanic islands, such as the Galapagos and Hawaiian Islands (then called Sandwich Islands) formed in mid-ocean and never part of any large continent. Such islands were characterised by a complete lack of terrestrial mammals and amphibians, and their inhabitants (except migratory birds and species introduced by humans) were typically the result of accidental colonisation and subsequent evolution.
  Continental islands were divided into those that were recently separated from a continent (like Britain) and those much less recently (like Madagascar). Wallace discussed how that difference affected flora and fauna. He discussed how isolation affected evolution and how that could result in the preservation of classes of animals, such as the lemurs of Madagascar that were remnants of once widespread continental faunas. He extensively discussed how changes of climate, particularly periods of increased glaciation, may have affected the distribution of flora and fauna on some islands, and the first portion of the book discusses possible causes of these great ice ages. Island Life was considered a very important work at the time of its publication. It was discussed extensively in scientific 
caption: The line separating the Indo-Malayan and the Austro-Malayan region in Wallace's On the Physical Geography of the Malay Archipelago (1863)
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