page_title: Dodo
section_title: Evolution
hierarchical_section_title: Dodo / Taxonomy / Evolution
caption_attribution_description: NICOBAR PIGEON
context_page_description: The dodo is an extinct flightless bird that was endemic to the island of Mauritius, east of Madagascar in the Indian Ocean. The dodo's closest genetic relative was the also-extinct Rodrigues solitaire, the two forming the subfamily Raphinae of the family of pigeons and doves. The closest living relative of the dodo is the Nicobar pigeon. A white dodo was once thought to have existed on the nearby island of Réunion, but this is now thought to have been confusion based on the Réunion ibis and paintings of white dodos.
  Subfossil remains show the dodo was about 1 metre tall and may have weighed 10.6–17.5 kg in the wild. The dodo's appearance in life is evidenced only by drawings, paintings, and written accounts from the 17th century. As these vary considerably, and only some of the illustrations are known to have been drawn from live specimens, its exact appearance in life remains unresolved, and little is known about its behaviour. Though the dodo has historically been considered fat and clumsy, it is now thought to have been well-adapted for its ecosystem.
context_section_description: In 2002, American geneticist Beth Shapiro and colleagues analysed the DNA of the dodo for the first time. Comparison of mitochondrial cytochrome b and 12S rRNA sequences isolated from a tarsal of the Oxford specimen and a femur of a Rodrigues solitaire confirmed their close relationship and their placement within the Columbidae. The genetic evidence was interpreted as showing the Southeast Asian Nicobar pigeon (Caloenas nicobarica) to be their closest living relative, followed by the crowned pigeons (Goura) of New Guinea, and the superficially dodo-like tooth-billed pigeon (Didunculus strigirostris) from Samoa (its scientific name refers to its dodo-like beak). This clade consists of generally ground-dwelling island endemic pigeons. The following cladogram shows the dodo's closest relationships within the Columbidae, based on Shapiro et al., 2002:
  A similar cladogram was published in 2007, inverting the placement of Goura and Didunculus and including the pheasant pigeon (Otidiphaps nobilis) and the thick-billed ground pigeon (Trugon terrestris) at the base of the clade. The DNA used in these studies was obtained from the Oxford specimen, and since this material is degraded, and no usable DNA has been extracted from subfossil remains, these findings still need to be independently verified. Based on behavioural and morphological evidence, Jolyon C. Parish proposed that the dodo and Rodrigues solitaire should be placed in the subfamily Gourinae along with the Goura pigeons and others, in agreement with the genetic evidence. In 2014, DNA of the only known specimen of the recently extinct spotted green pigeon (Caloenas maculata) was analysed, and it was found to be a close relative of the Nicobar pigeon, and thus also the dodo and Rodrigues solitaire.
  The 2002 study indicated that the ancestors of the dodo and the solitaire diverged around the Paleogene-Neogene boundary, about 23.03 million years ago. The Mascarene Islands (Mauritius, Réunion, and Rodrigues), are of volcanic origin and are less than 10 million years old. Therefore, the ancestors of both birds probably remained capable of flight for a considerable time after the separation of their lineage. The Nicobar and spotted green pigeon were placed at the base of a lineage leading to the Raphinae, which indicates the flightless raphines had ancestors that were able to fly, were semi-terrestrial, and inhabited islands. This in turn supports the hypothesis that the ancestors of those birds reached the Mascarene islands by island hopping from South Asia. The lack of mammalian herbivores competing for resources on these islands allowed the solitaire and the dodo to attain very large sizes and flightlessness. Despite its divergent skull morphology and adaptations for larger size, many features of its skeleton remained similar to those of smaller, flying pigeons. Another large, flightless pigeon, the Viti Levu giant pigeon (Natunaornis gigoura), was described in 2001 from subfossil material from Fiji. It was only slightly smaller than the dodo and the solitaire, and it too is thought to have been related to the crowned pigeons.
caption: The Nicobar pigeon is the closest living relative of the dodo
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