page_title: Feathered dinosaur
section_title: List of dinosaurs with evidence of feathers
hierarchical_section_title: Feathered dinosaur / Current knowledge / List of dinosaurs with evidence of feathers
caption_attribution_description: English: The holotype of Microraptor gui, IVPP V 13352 under normal light. This shows the preserved feathers (white arrow) and the 'halo' around the specimen where they appear to be absent (black arrows). Scale bar at 5 cm. Polski: Holotyp Microraptor gui, IVPP V 13352, widziany w normalnym świetle. Białą strzałką zaznaczono zachowane pióra, a czarnymi – "halo" wokół okazu, gdzie pióra wydają się nieobecne. Pasek skali wynosi 5 cm.
context_page_description: A feathered dinosaur is a dinosaur with feathers. It is now thought that all coelurosaurs, and perhaps all theropods, had feathers.
  The possibility that dinosaurs are closely related to birds raised the possibility of feathers. Fossils of Archaeopteryx include well-preserved feathers, but it was not until the early 1990s that clearly non-bird dinosaur fossils were discovered with preserved feathers. About 40 genera of theropods are now known to be feathered.
  Most of these fossils come from the Yixian formation in China. The fossil feathers of one specimen, Shuvuuia deserti, have been tested positive for beta-keratin in immunological tests. This is the main protein in bird feathers.
context_section_description: A number of non-avialan dinosaurs are now known to have been feathered. Direct evidence of feathers exists for the following genera, listed in the order currently accepted evidence was first published.
  The evidence consists of feather impressions, or convincing skeletal or chemical evidence. Skeletal evidence would be the presence of quill knobs (the anchor points for wing feathers on the forelimb) or a pygostyle (the fused vertebrae at the tail tip which often supports large feathers).
  For comparison: Archaeopteryx (1861; avialan) definitely had working flight feathers with off-centre rachis (stem). Otherwise, apart from the front limbs, it was a typical small carnivorous dinosaur.
  Avimimus portentosus (inferred 1987: quill knobs)
  Sinosauropteryx prima (1996)
  Protarchaeopteryx robusta (1997)
  GMV 2124 (1997).
  Caudipteryx zoui (1998)
  Shuvuuia deserti (1999)
  Sinornithosaurus millenii (1999)
  Beipiaosaurus inexpectus (1999)
  Caudipteryx dongi (2000)
  Caudipteryx sp. (2000)
  Microraptor zhaoianus (2000)
  Nomingia gobiensis (inferred 2000: pygostyle)
  Psittacosaurus sp.? (2002)
  Scansoriopteryx heilmanni (2002)
  Yixianosaurus longimanus (2003)
  Dilong paradoxus (2004)
  Jinfengopteryx elegans (2005)
  Juravenator starki (2006)
  Sinocalliopteryx gigas (2007)
  Velociraptor mongoliensis (inferred 2007: quill knobs)
  Epidexipteryx hui (2008)
  Similicaudipteryx yixianensis (inferred 2008: pygostyle; confirmed 2010)
  Zhongornis haoae (2008)
  Tianyulong confuciusi? (2009)
  Concavenator corcovatus? (inferred 2010: quill knobs?)
  Yutyrannus huali (2012)
  Microraptor hanqingi (2012)
  Ornithomimus edmontonicus (2012)
  Ningyuansaurus wangi (2012)
  Eosinopteryxbrevipenna (2013)
  Citipati osmolskae (inferred 2013: pygostyle)
  Conchoraptor gracilis (inferred 2013: pygostyle)
  Jianchangosaurus yixianensis (2013)
  Aurornis xui (2013; possibly avialan)
  Changyuraptor yangi (2014)
  Kulindadromeus zabaikalicus? (2014)
  Citipati osmolskae (inferred 2014: pygostyle)
  Conchoraptor gracilis (inferred 2014: pygostyle)
  Deinocheirus mirificus (inferred 2014: pygostyle)
  Yi qi (2015)
  Note, filamentous structures in some ornithischian dinosaurs (Psittacosaurus, Tianyulong) and pterosaurs may or may not be homologous with the feathers and protofeathers of theropods.
caption: Fossil of Microraptor gui impressions of feathered wings
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