page_title: Frog
section_title: Respiration and circulation
hierarchical_section_title: Frog / Morphology and physiology / Respiration and circulation
caption_attribution_description: Frog anatomy display in NMH Washington DC, United States. Right atrium Liver Aorta Egg mass Colon Left atrium Ventricle Stomach Left lung Spleen Small intestine Cloaca
context_page_description: A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura. The oldest fossil "proto-frog" appeared in the early Triassic of Madagascar, but molecular clock dating suggests their origins may extend further back to the Permian, 265 million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. There are over 6,300 recorded species, accounting for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from taxonomy or evolutionary history. All toads are frogs, but not all frogs are toads.
  An adult frog has a stout body, protruding eyes, anteriorly-attached tongue, limbs folded underneath, and no tail. Frogs have glandular skin, with secretions ranging from distasteful to toxic. Their skin varies in colour from well-camouflaged dappled brown, grey and green to vivid patterns of bright red or yellow and black to show toxicity and ward off predators.
context_section_description: The skin of a frog is permeable to oxygen and carbon dioxide, as well as to water. There are blood vessels near the surface of the skin and when a frog is underwater, oxygen diffuses directly into the blood. When not submerged, a frog breathes by a process known as buccal pumping. Its lungs are similar to those of humans, but the chest muscles are not involved in respiration, and no ribs or diaphragm exist to help move air in and out. Instead, it puffs out its throat and draws air in through the nostrils, which in many species can then be closed by valves. When the floor of the mouth is compressed, air is forced into the lungs. The fully aquatic Bornean flat-headed frog (Barbourula kalimantanensis) is the first frog known to lack lungs entirely.
  Frogs have three-chambered hearts, a feature they share with lizards. Oxygenated blood from the lungs and de-oxygenated blood from the respiring tissues enter the heart through separate atria. When these chambers contract, the two blood streams pass into a common ventricle before being pumped via a spiral valve to the appropriate vessel, the aorta for oxygenated blood and pulmonary artery for deoxygenated blood. The ventricle is partially divided into narrow cavities which minimizes the mixing of the two types of blood. These features enable frogs to have a higher metabolic rate and be more active than would otherwise be possible.
  Some species of frog have adaptations that allow them to survive in oxygen deficient water. The Titicaca water frog (Telmatobius culeus) is one such species and has  wrinkly skin that increases its surface area to enhance gas exchange. It normally makes no use of its rudimentary lungs but will sometimes raise and lower its body rhythmically while on the lake bed to increase the flow of water around it.
caption: Anatomical model of a dissected frog: 1 Right atrium, 2 Lungs, 3 Aorta, 4 Egg mass, 5 Colon, 6 Left atrium, 7 Ventricle, 8 Stomach, 9 Liver, 10 Gallbladder, 11 Small intestine, 12 Cloaca
exif: {}