page_title: Anhanguera (pterosaur)
section_title: Former species and nomina dubia
hierarchical_section_title: Anhanguera (pterosaur) / Classification / Former species and nomina dubia
caption_attribution_description: Specimen AMNH 22555, a partial anhanguerid skeleton. Some selected elements are figured in detail. (A) pelvic region in dorsal view; (B) torso in dorsal view; (C, D, E) sixth cervical vertebrae in, respectively, anterior, dorsal and right lateral views; (F, G) right mandibular ramus in, respectively, medial and lateral views; (H) left scapula in dorsal view; (I) left coracoid in lateral view; (J) distal carpals in distal view; (K) proximal carpals in distal view. Scale bars equal to 50 mm. Line drawings of some bones were modified from Witton (2013).
context_page_description: Anhanguera is a genus of pterodactyloid pterosaur known from the Early Cretaceous Romualdo Formation of Brazil. This pterosaur is closely related to Ornithocheirus, but belongs in the family Anhangueridae. The generic name comes from the tupi words añanga, meaning "spirit protector of the animals" + wera "bygone".
context_section_description: Various other species have been assigned to Anhanguera, but all of them are either dubious or have been transferred to other genera. A 2017 review found A. araripensis, A. robustus, and A. santanae to be based on non-diagnostic material, and therefore were dubious.
  A. araripensis (Wellnhofer 1985) = Santanadactylus araripensis Wellnhofer 1985 [also classified as Coloborhynchus]
  A. cuvieri (Bowerbank 1851) = Ornithocheirus cuvieri = Pterodactylus cuvieri Bowerbank 1851 [now classified as Cimoliopterus]
  A. fittoni (Owen 1858) = Pterodactylus fittoni Owen 1858
  A. robustus (Wellnhofer 1987) = Tropeognathus robustus Wellnhofer 1987 [also classified as Coloborhynchus]
  A. ligabuei (Dalla Vecchia 1993) = Cearadactylus ligabuei Dalla Vecchia 1993 [also classified as Coloborhynchus or Cearadactylus]
  A. santanae (Wellnhofer 1985) = Araripesaurus santanae Wellnhofer 1985
caption: Anhanguerid skeleton AMNH 22555, which was formerly assigned to A. santanae
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