page_title: Cleridae
section_title: Systematics
hierarchical_section_title: Cleridae / Systematics
caption_attribution_description: Opilus Fasciatus = Tarsostenus univittatus English: Shillingford's Opilus
context_page_description: Cleridae are a family of beetles of the superfamily Cleroidea. They are commonly known as checkered beetles. The family Cleridae has a worldwide distribution, and a variety of habitats and feeding preferences.
  Cleridae have many niches and feeding habits. Most genera are predaceous and feed on other beetles and larvae; however other genera are scavengers or pollen feeders. Clerids have elongated bodies with bristly hairs, are usually bright colored, and have variable antennae. Checkered beetles range in length between 3 millimeters and 24 millimeters. Cleridae can be identified based on their 5–5–5 tarsal formula, division of sternites, and the absence of a special type of vesicle. Female Cleridae lay between 28–42 eggs at a time predominately under the bark of trees. Larvae are predaceous and feed vigorously before pupation and subsequently emergence as adults.
  Clerids have a minor significance in forensic entomology. Some species are occasionally found on carrion in the later dry stages of decay. Also, some species are pests and are found infesting various food products. Research efforts related to Cleridae have focused primarily on using certain species as biological controls.
context_section_description: The genera of Cleridae are divided among several subfamilies, though some genera still defy easy classification. Several taxonomic schemes exist, recognizing for example a group around Neorthopleura as distinct subfamily Neorthopleurinae, or splitting off the Thaneroclerinae as distinct family, or circumscribing the Korynetinae sensu stricto or sensu lato. The following list of tribes and selected genera is thus preliminary. Some notable species are also listed.
  
  
   
  Clerinae
  Allonyx Jacquelin du Val, 1860
  Anthicoclerus Schenkling, 1906
  Aphelochroa Quedenfeldt, 1885
  Apopempsis Schenkling, 1903
  Apteroclerus Wollaston, 1867
  Aptinoclerus Kuwert, 1893
  Aradamicula Sedlacek & Winkler, 1975
  †Arawakis (fossil)
  Astigmus Kuwert, 1894
  Aulicus Spinola, 1841
  Axina Kirby, 1818
  Balcus
  Barriella Opitz, 2003
  Barrotillus Rifkind, 1996
  Blaxima Gorham, 1882
  Bousquetoclerus Menier, 1997
  Burgeoneus Pic, 1950
  Caestron Dupont in Spinola, 1844
  Calendyma Lacordaire, 1857
  Canariclerus Winkler, 1982
  Cardiostichus Quedenfeldt, 1885
  Caridopus Schenkling in Sjöstedt, 1908
  Cleromorpha Gorham, 1876
  Cleropiestus Fairmaire, 1889
  Clerus Fabricius, 1775
  Clytomadius Corporaal, 1949
  Colyphus Spinola, 1841
  Coptoclerus Chapin, 1924
  Cormodes Pascoe, 1860
  Corynommadius Schenkling, 1899
  Ctenaxina Schenkling, 1906
  Ctenoclerus Solervicens, 1997
  Dasyceroclerus Kuwert, 1894
  Dasyteneclines Pic, 1941
  Dieropsis Gahan, 1908
  Dologenitus Opitz, 2009
  Dozocolletus Chevrolat, 1842
  Eburiphora Spinola, 1841
  Eburneoclerus Pic, 1950
  Ekisius Winkler, 1987
  Eleale Newman, 1840
  Enoclerus Gahan, 1910
  Epiclines Chevrolat in Guérin-Ménéville, 1839
  Eunatalis Schenkling, 1909
  Eunatalis porcata
  Erymanthus Spinola, 1841
  Eurymetomorphon Pic, 1950
  Falsomadius Gerstmeier, 2002
  Falsoorthrius Pic, 1940
  Graptoclerus Gorham, 1901
  Gyponyx Gorham, 1883
  Hemitrachys Gorham, 1876
  Homalopilo Schenkling, 1915
  Inhumeroclerus Pic, 1955
  Jenjouristia Fursov, 1936
  Languropilus Pic, 1940
  Lissaulicus C.O.Waterhouse, 1879
  Memorthrius Pic, 1940
  Metademius Schenkling, 1899
  Microclerus Wollaston, 1867
  Micropteroclerus Chapin, 1920
  Microstigmatium Kraatz, 1899
  Mimolesterus Gerstmeier, 1991
  Mitrandiria Kolibac, 1997
  Myrmecomaea Fairmaire, 1886
  Natalis Laporte de Castelnau, 1836
  Neogyponyx Schenkling, 1906
  Neoscrobiger Blackburn, 1900
  Ohanlonella Rifkind, 2008
  Olesterus Spinola, 1841
  Omadius Laporte de Castelnau, 1836
  Oodontophlogistus Elston, 1923
  Operculiphorus Kuwert, 1894
  Opilo Latreille, 1802
  Orthrius Gorham, 1876
  Oxystigmatium Kraatz, 1899
  Phlogistomorpha Hintz, 1908
  Phlogistus Gorham, 1876
  Phloiocopus Spinola, 1841
  Phonius Chevrolat, 1843
  Pieleus Pic, 1940
  Placocerus Klug, 1837
  Placopterus Wolcott, 1910
  Plathanocera Schenkling, 1902
  Platyclerus Spinola, 1841
  Priocera Kirby, 1818
  Priocleromorphus Pic, 1950
  Prioclerus Hintz, 1902
  Pseudolesterus Miyatake, 1968
  Pseudomadius Chapin, 1924
  Pujoliclerus Pic, 1947
  Sallea Chevrolat, 1874
  Scrobiger Spinola, 1841
  Sedlacekius Winkler, 1972
  Sikorius Kuwert, 1893
  Stigmatium Gray in Griffith, 1832
  Systenoderes Spinola, 1841
  Tanocleria Hong, 2002
  Thalerocnemis Lohde, 1900
  Thanasimodes Murray, 1867
  Thanasimus Latreille, 1806
  Thanasimus formicarius – Ant Beetle
  Tillicera Spinola, 1841
  Trichodes Herbst, 1792
  Trichodes alvearius
  Trichodes apiarius
  Trichodes leucopsideus
  Trogodendron Spinola, 1841
  Trogodendron fasciculatum – Yellow-horned Clerid
  Wilsonoclerus
  Winklerius Menier, 1986
  Wittmeridecus Winkler, 1981
  Xenorthrius Gorham, 1892
  Zahradnikius Winkler, 1992
  Zenithicola Spinola, 1841
   
  Enopliinae (sometimes in Korynetinae)
  Antygodera
  Apolopha Spinola, 1841
  Corinthiscus Fairmaire & Germain, 1861
  Cregya LeConte, 1861
  Curacavi
  Enoplium Latreille, 1802
  Exochonotus
  Hublella
  Lasiodera Gray in Griffith, 1832
  Neopylus Solervicens, 1989
  Paracregya
  Pelonium
  Phymatophaea Pascoe, 1876
  Platynoptera Chevrolat, 1834
  Pseudichnea Schenkling, 1900
  Pylus Newman, 1840
  Pyticara Spinola, 1841 (including Pelonides)
  Teneroides Gahan, 1910
  Tenerus Laporte de Castelnau, 1836
  Thriocerodes Wolcott & Dybas, 1947
  Epiphloeinae (sometimes in Korynetinae)
  Acanthocollum
  Amboakis
  Decaphloeus
  Decorosa Opitz, 2008
  Dia
caption: Tarsostenus univittatus (Tarsosteninae)
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