Document:

FORM OF EXECUTIVE INCENTIVE AWARD AGREEMENT

 Exhibit 10.3 
 ORGANOVO HOLDINGS, INC. 
 EQUITY INCENTIVE AWARD AGREEMENT

 This Equity Incentive Award Agreement (the “Agreement”) is made effective as of
            , 2012 (the “Effective Date”), by and between Organovo Holdings, Inc., a Delaware corporation (the “Company”) and
            (“Executive”). 
 WHEREAS, the Board of
Directors of the Company (the “Board”) has the authority to issue awards pursuant to the terms, conditions and limitations of the Organovo Holdings, Inc. 2012 Equity Incentive Plan (the “Plan”); 

WHEREAS, the Company awarded the Executive a restrictive stock unit award for up to
            shares of Company common stock (“Common Stock”) on             , 2012 (“Initial Award”).

 WHEREAS, the Initial Award provides that Executive may return shares of vested restricted stock the Executive receives
pursuant to the Initial Award to the Company for cancellation on each Vesting Date of the Initial Award to satisfy federal and state tax withholding taxes. 
 WHEREAS, the Board believes it is in the Company’s best interest to agree to grant Executive additional option awards on the terms and conditions contained herein. 

NOW, THEREFORE, in consideration for the promises and mutual agreements hereinafter set forth, it is agreed by and between the
undersigned as follows: 
 1. Award Grants. The Company hereby agrees to grant the Executive a fully-vested option Award
(each a “New Award” and collectively, the “New Awards”) on each Vesting Date of the Initial Award for a number of shares to be determined in accordance with the provisions of Section 2 if the Executive elects on a Vesting
Date to return Shares of vested Common Stock to the Company for cancellation on Vesting Date to satisfy federal and state tax withholding taxes. 
 2. Number of Shares Subject to Award. The number of shares subject to a New Award shall be equal to the number of Shares of Common Stock returned to the Company on the applicable Vesting Date in
order to satisfy the federal and state withholding taxes. 
 3. Option Terms. The New Awards (i) shall be issued on
the applicable Vesting Date; (ii) shall be fully-vested; (iii) shall have an Exercise Price equal to fair market value on the applicable Vesting Date, and (iv) shall be in the form attached hereto as Exhibit A. 

4. Non-Guarantee of Employment or Service Relationship. Nothing in this Agreement shall alter Executive’s at-will or other
employment status or other service relationship with the Company, nor be construed as a contract of employment or service relationship between the Company and Executive, or as a contractual right of Executive to continue in the employ of, or in a
service relationship with, the Company for any period of time, or as a limitation of the right of the Company to discharge Executive at any time with or without cause or notice and whether or not such discharge results in the forfeiture of any Award
Shares. 

 5. The Company’s Rights. The existence of the Award shall not affect in any way
the right or power of the Company or its stockholders to make or authorize any or all adjustments, recapitalizations, reorganizations or other changes in the Company’s capital structure or its business, or any merger or consolidation of the
Company, or any issue of bonds, debentures, preferred or other stocks with preference ahead of or convertible into, or otherwise affecting the Common Stock or the rights thereof, or the dissolution or liquidation of the Company, or any sale or
transfer of all or any part of the Company’s assets or business, or any other corporate act or proceeding, whether of a similar character or otherwise. 
 6. Notices. All notices and other communications made or given pursuant to this Agreement shall be in writing and shall be sufficiently made or given if hand delivered or mailed by certified mail,
addressed to Executive at the address contained in the records of the Company, or addressed to the Administrator, care of the Company for the attention of its Corporate Secretary at its principal executive office or, if the receiving party consents
in advance, transmitted and received via telecopy or via such other electronic transmission mechanism as may be available to the parties. 
 7. Entire Agreement. This Agreement contains the entire agreement between the parties with respect to the Award granted hereunder. Any oral or written agreements, representations, warranties,
written inducements, or other communications made prior to the execution of this Agreement with respect to the Award granted hereunder shall be void and ineffective for all purposes. 

8. Effect of Administrator’s Decision. All actions taken and decisions and determinations made by the Administrator on all
matters relating to this Agreement and the Award pursuant to the powers vested in it under the Plan shall be in the Administrator’s sole and absolute discretion and shall be conclusive and binding on all parties concerned, including Executive,
the Company, its stockholders, director and officers, and Executive’s and its respective successors in interest. 
 9.
Amendment. This Agreement may be amended from time to time by the Plan Administrator in its discretion; provided, however, that this Agreement may not be modified in a manner that would have a materially adverse effect on Executive’s
rights with respect to the Award as determined in the discretion of the Administrator, except as provided in a written document signed by each of the parties hereto. 
 10. Governing Law. The validity, construction and effect of this Agreement, and of any determinations or decisions made by the Administrator relating to this Agreement, and the rights of any and
all persons having or claiming to have any interest under this Agreement, shall be determined exclusively in accordance with the laws of the State of California, without regard to its provisions concerning the applicability of laws of other
jurisdictions. As a condition of this Agreement, Executive agrees that he will not bring any action arising under, as a result of, pursuant to or relating to, this Agreement in any court other than a federal or state court in the districts which
include San Diego, California, and Executive hereby agrees and submits to the personal jurisdiction of any federal court located in the district which includes San Diego, California or any state court in the district which includes San Diego,
California. Executive further agrees that he will not deny or attempt to defeat such personal jurisdiction or object to venue by motion or other request for leave from any such court. 

  
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 11. Resolution of Disputes. Any dispute or disagreement which shall arise under, or
as a result of, or pursuant to or relating to, this Agreement shall be determined by the Administrator in good faith in its absolute and uncontrolled discretion, and any such determination or any other determination by the Administrator under or
pursuant to this Agreement and any interpretation by the Administrator of the terms of this Agreement, will be final, binding and conclusive on all persons affected thereby. Executive agrees that before he may bring any legal action arising under,
as a result of, pursuant to or relating to, this Agreement he will first exhaust his administrative remedies before the Administrator. Executive further agrees that in the event that the Administrator does not resolve any dispute or disagreement
arising under, as a result of, pursuant to or relating to, this Agreement to Executive’s satisfaction, no legal action may be commenced or maintained relating to this Agreement more than twenty-four (24) months after the
Administrator’s decision. 
 12. Headings. The headings in this Agreement are for reference purposes only and shall
not affect the meaning or interpretation of this Agreement. 
 13. Counterparts. This Agreement may be executed in
counterparts, each of which shall be deemed an original, but all of which together shall constitute one and the same instrument. 
 14. Electronic Delivery of Documents. By Executive’s signing this Agreement, he (i) consents to the electronic delivery of this Agreement, all information with respect to the Award Shares
and any reports of the Company provided generally to the Company’s stockholders; (ii) acknowledges that he may receive from the Company a paper copy of any documents delivered electronically at no cost to him by contacting the Company by
telephone or in writing; (iii) further acknowledges that he may revoke Executive’s consent to the electronic delivery of documents at any time by notifying the Company of such revoked consent by telephone, postal service or electronic
mail; and (iv) further acknowledges that he understands that he is not required to consent to electronic delivery of documents. 
 15. No Future Entitlement. By Executive’s signing this Agreement, he acknowledges and agrees that: (i) the grant of this Award is a one-time benefit which does not create any contractual
or other right to receive future grants of stock, or compensation in lieu of stock grants, even if stock grants have been granted repeatedly in the past; (ii) all determinations with respect to any such future grants, including, but not limited
to, the times when stock grants shall be granted, the maximum number of shares subject to each stock grant, and the times or conditions under which restrictions on such stock grants shall lapse, will be at the sole discretion of the Administrator;
(iii) the value of this stock grant is an extraordinary item of compensation which is outside the scope of Executive’s employment contract, if any; (iv) the value of this stock grant is not part of normal or expected compensation or
salary for any purpose, including, but not limited to, calculating any termination, severance, resignation, redundancy, end of service payments or similar payments, or bonuses, long-service awards, pension or retirement benefits; (v) the
vesting of these Award Shares ceases upon termination of employment with the Company or transfer of employment from the Company, or other cessation of eligibility for any 

  
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reason, except as may otherwise be explicitly provided in this Agreement; (vi) the Company does not guarantee any future value of these Award Shares; and (vii) no claim or entitlement
to compensation or damages arises if these Award Shares do not increase in value and Executive irrevocably releases the Company from any such claim that does arise. 
 16. Personal Data. For purposes of the implementation, administration and management of the Award or the effectuation of any acquisition, equity or debt financing, joint venture, merger,
reorganization, consolidation, recapitalization, business combination, liquidation, dissolution, share exchange, sale of stock, sale of material assets or other similar corporate transaction involving the Company (a “Corporate
Transaction”), Executive consents, by execution of the Notice, to the collection, receipt, use, retention and transfer, in electronic or other form, of Executive’s personal data by and among the Company and its third party vendors or any
potential party to a potential Corporate Transaction. Executive understands that personal data (including but not limited to, name, home address, telephone number, employee number, employment status, social security number, tax identification
number, date of birth, nationality, job and payroll location, data for tax withholding purposes and shares awarded, cancelled, vested and unvested) may be transferred to third parties assisting in the implementation, administration and management of
the stock grant or the effectuation of a Corporate Transaction and Executive expressly authorizes such transfer as well as the retention, use, and the subsequent transfer of the data by the recipient(s). Executive understands that these recipients
may be located in his country or elsewhere, and that the recipient’s country may have different data privacy laws and protections than Executive’s country. Executive understands that data will be held only as long as is necessary to
implement, administer and manage the stock grant or effect a Corporate Transaction. Executive understands that he may, at any time, request a list with the names and addresses of any potential recipients of the personal data, view data, request
additional information about the storage and processing of data, require any necessary amendments to data or refuse or withdraw the consents herein, in any case without cost, by contacting in writing the Company’s Secretary. Executive
understands, however, that refusing or withdrawing Executive’s consent may affect Executive’s ability to accept a stock grant. 
 17. No Transfers. The Executive’s rights under this Agreement are not transferable other than by the Company pursuant to the terms and conditions of the Plan. 

18. Plan Governs. Any capitalized terms not defined herein shall have the meaning set forth in the Plan. In the event of any
discrepancy between the terms of this Agreement and the terms of the Plan, the terms of the Plan shall govern. 

  
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	 ORGANOVO HOLDINGS, INC.

		
	 By:
	 	  

	 Date:
	 	  

 The undersigned hereby acknowledges that he/she has carefully read this Agreement and agrees to be
bound by all of the provisions set forth herein. The undersigned also consents to electronic delivery of all notices or other information with respect to the Award Shares or the Company. 

 

							
	 WITNESS:
	 		 	GRANTEE
			
	  
	 		 	  

				
		 		 	Date:	 	  

  
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 EXHIBIT A 

ORGANOVO HOLDINGS, INC. 
 2012 EQUITY INCENTIVE PLAN 
 STOCK OPTION AWARD AGREEMENT 

(EXECUTIVE) 
 Unless otherwise defined herein, the terms defined in the Organovo Holdings, Inc. 2012 Equity Incentive Plan (the “Plan”) will have the same defined meanings in this Stock Option Award Agreement
(the “Award Agreement”). 
  

	 	•	 	 NOTICE OF STOCK OPTION GRANT 

 Participant Name: 
 You have been granted an Option to purchase Common
Stock of Organovo Holdings, Inc. (the “Company”) pursuant to the Executive Incentive Award Agreement, dated             , subject to the terms and conditions of the Plan and this
Award Agreement, as follows: 
  

							
	Date of Grant	 	  
	 	
				
	Exercise Price per Share	 	$	 	  
	 	
			
	Total Number of Shares Granted	 	  
	 	
				
	Total Exercise Price	 	$	 	  
	 	
				
	Type of Option	 		 	        Incentive Stock Option	 	
				
		 		 	        Nonstatutory Stock Option	 	
			
	Term/Expiration Date:	 	  
	 	

 Vesting and Exercise Schedule: The Option shall be fully vested and exercisable on the Date of
Grant. 
 Termination Period: This Option will be exercisable for three (3) months after Participant ceases to be a
Service Provider, unless such termination is due to Participant’s death or Disability, in which case this Option will be exercisable for twelve (12) months after Participant ceases to be Service Provider. Notwithstanding the foregoing, in
no event may this Option be exercised after the Term/Expiration Date as provided above and may be subject to earlier termination as provided in Section 15 of the Plan. 
 By Participant’s signature and the signature of the Company’s representative below, Participant and the Company agree that this Option is granted under and governed by the terms and conditions
of the Plan, the Executive Incentive Award Agreement and this Award 

  
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Agreement, including the Terms and Conditions of Stock Option Grant, attached hereto as Exhibit A-1, all of which are made a part of this document. Participant has reviewed the Plan, the
Executive Incentive Award Agreement and this Award Agreement in their entirety, has had an opportunity to obtain the advice of counsel prior to executing this Award Agreement and fully understands all provisions of the Plan and Award Agreement.
Participant hereby agrees to accept as binding, conclusive and final all decisions or interpretations of the Administrator upon any questions relating to the Plan, the Executive Incentive Award Agreement and the Award Agreement. Participant further
agrees to notify the Company upon any change in the residence address indicated below. 
  

					
	PARTICIPANT	 		 	ORGANOVO HOLDINGS, INC.
			
	  
	 		 	  

	Signature	 		 	By
			
	  
	 		 	  

	Print Name	 		 	Title
			
	Residence Address:	 		 	
			
	  
	 		 	
			
	  
	 		 	

  
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 EXHIBIT A-1 

TERMS AND CONDITIONS OF STOCK OPTION GRANT 
 1. Grant of Option. The Company hereby grants to the Participant named in the Notice of Stock Option Grant (“Notice of Grant”) attached as Part I of this Award Agreement (the
“Participant”) an option (the “Option”) to purchase the number of Shares, as set forth in the Notice of Grant, at the exercise price per Share set forth in the Notice of Grant (the “Exercise Price”), subject to all of
the terms and conditions in this Award Agreement, the Executive Incentive Award Agreement and the Plan, which is incorporated herein by reference. Subject to Section 20 of the Plan, in the event of a conflict between the terms and conditions of
the Plan and the terms and conditions of this Award Agreement, the terms and conditions of the Plan will prevail. 
 If
designated in the Notice of Grant as an Incentive Stock Option (“ISO”), this Option is intended to qualify as an ISO under Section 422 of the Internal Revenue Code of 1986, as amended (the “Code”). However, if this Option is
intended to be an ISO, to the extent that it exceeds the $100,000 rule of Code Section 422(d) it will be treated as a Nonstatutory Stock Option (“NSO”). Further, if for any reason this Option (or portion thereof) will not qualify as
an ISO, then, to the extent of such nonqualification, such Option (or portion thereof) shall be regarded as a NSO granted under the Plan. In no event will the Administrator, the Company or any Parent or Subsidiary or any of their respective
employees or directors have any liability to Participant (or any other person) due to the failure of the Option to qualify for any reason as an ISO. 
 2. Vesting Schedule. The Option is fully vested on the Grant Date. 
 3.
Exercise of Option. 
 (a) Right to Exercise. This Option may be exercised only within the term set out in the
Notice of Grant, and may be exercised during such term only in accordance with the Plan and the terms of this Award Agreement. 

(b) Method of Exercise. This Option is exercisable by delivery of an exercise notice, in the form attached as
Exhibit B (the “Exercise Notice”) or in a manner and pursuant to such procedures as the Administrator may determine, which will state the election to exercise the Option, the number of Shares in respect of which the Option is
being exercised (the “Exercised Shares”), and such other representations and agreements as may be required by the Company pursuant to the provisions of the Plan. The Exercise Notice will be completed by Participant and delivered to the
Company. The Exercise Notice will be accompanied by payment of the aggregate Exercise Price as to all Exercised Shares together with any applicable tax withholding. This Option will be deemed to be exercised upon receipt by the Company of such fully
executed Exercise Notice accompanied by such aggregate Exercise Price. 
 4. Method of Payment. Payment of the aggregate
Exercise Price will be by any of the following, or a combination thereof, at the election of Participant. 
 (a) cash;

  
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 (b) check; 
 (c) consideration received by the Company under a formal cashless exercise program adopted by the Company in connection with the Plan; or 

(d) surrender of other Shares which have a Fair Market Value on the date of surrender equal to the aggregate Exercise Price of the
Exercised Shares, provided that accepting such Shares, in the sole discretion of the Administrator, will not result in any adverse accounting consequences to the Company. 
 5. Tax Obligations. 
 (a) Withholding Taxes. Notwithstanding any
contrary provision of this Award Agreement, no certificate representing the Shares will be issued to Participant, unless and until satisfactory arrangements (as determined by the Administrator) will have been made by Participant with respect to the
payment of income, employment and other taxes which the Company determines must be withheld with respect to such Shares. To the extent determined appropriate by the Company in its discretion, it will have the right (but not the obligation) to
satisfy any tax withholding obligations by reducing the number of Shares otherwise deliverable to Participant. If Participant fails to make satisfactory arrangements for the payment of any required tax withholding obligations hereunder at the time
of the Option exercise, Participant acknowledges and agrees that the Company may refuse to honor the exercise and refuse to deliver Shares if such withholding amounts are not delivered at the time of exercise. 

(b) Notice of Disqualifying Disposition of ISO Shares. If the Option granted to Participant herein is an ISO, and if Participant
sells or otherwise disposes of any of the Shares acquired pursuant to the ISO on or before the later of (i) the date two (2) years after the Date of Grant, or (ii) the date one (1) year after the date of exercise, Participant
will immediately notify the Company in writing of such disposition. Participant agrees that Participant may be subject to income tax withholding by the Company on the compensation income recognized by Participant. 

(c) Code Section 409A. Under Code Section 409A, an option that vests after December 31, 2004 (or that vested on or
prior to such date but which was materially modified after October 3, 2004) that was granted with a per Share exercise price that is determined by the Internal Revenue Service (the “IRS”) to be less than the Fair Market Value of a
Share on the date of grant (a “Discount Option”) may be considered “deferred compensation.” A Discount Option may result in (i) income recognition by Participant prior to the exercise of the option, (ii) an additional
twenty percent (20%) federal income tax, and (iii) potential penalty and interest charges. The Discount Option may also result in additional state income, penalty and interest charges to the Participant. Participant acknowledges that the
Company cannot and has not guaranteed that the IRS will agree that the per Share exercise price of this Option equals or exceeds the Fair Market Value of a Share on the Date of Grant in a later examination. Participant agrees that if the IRS
determines that the Option was granted with a per Share exercise price that was less than the Fair Market Value of a Share on the date of grant, Participant will be solely responsible for Participant’s costs related to such a determination.

  
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 6. Rights as Stockholder. Neither Participant nor any person claiming under or
through Participant will have any of the rights or privileges of a stockholder of the Company in respect of any Shares deliverable hereunder unless and until certificates representing such Shares will have been issued, recorded on the records of the
Company or its transfer agents or registrars, and delivered to Participant. After such issuance, recordation and delivery, Participant will have all the rights of a stockholder of the Company with respect to voting such Shares and receipt of
dividends and distributions on such Shares. 
 7. No Guarantee of Continued Service. PARTICIPANT ACKNOWLEDGES AND AGREES
THAT THE VESTING OF SHARES PURSUANT TO THE VESTING SCHEDULE HEREOF IS EARNED ONLY BY CONTINUING AS A SERVICE PROVIDER AT THE WILL OF THE COMPANY (OR THE PARENT OR SUBSIDIARY EMPLOYING OR RETAINING PARTICIPANT) AND NOT THROUGH THE ACT OF BEING HIRED,
BEING GRANTED THE OPTION OR ACQUIRING SHARES HEREUNDER. PARTICIPANT FURTHER ACKNOWLEDGES AND AGREES THAT THIS AWARD AGREEMENT, THE TRANSACTIONS CONTEMPLATED HEREUNDER AND THE VESTING SCHEDULE SET FORTH HEREIN DO NOT CONSTITUTE AN EXPRESS OR IMPLIED
PROMISE OF CONTINUED ENGAGEMENT AS A SERVICE PROVIDER FOR THE VESTING PERIOD, FOR ANY PERIOD, OR AT ALL, AND WILL NOT INTERFERE IN ANY WAY WITH PARTICIPANT’S RIGHT OR THE RIGHT OF THE COMPANY (OR THE PARENT OR SUBSIDIARY EMPLOYING OR RETAINING
PARTICIPANT) TO TERMINATE PARTICIPANT’S RELATIONSHIP AS A SERVICE PROVIDER AT ANY TIME, WITH OR WITHOUT CAUSE. 
 8.
Address for Notices. Any notice to be given to the Company under the terms of this Award Agreement will be addressed to the Company, in care of its Chief Financial Officer at Organovo Holdings, Inc., 5897 Oberlin Drive, Suite 217, San Diego,
CA 92121, or at such other address as the Company may hereafter designate in writing. 
 9. Non-Transferability of
Option. This Option may not be transferred in any manner otherwise than by will or by the laws of descent or distribution and may be exercised during the lifetime of Participant only by Participant. 

10. Binding Agreement. Subject to the limitation on the transferability of this grant contained herein, this Award Agreement will
be binding upon and inure to the benefit of the heirs, legatees, legal representatives, successors and assigns of the parties hereto. 
 11. Additional Conditions to Issuance of Stock. If at any time the Company will determine, in its discretion, that the listing, registration or qualification of the Shares upon any securities
exchange or under any state or federal law, or the consent or approval of any governmental regulatory authority is necessary or desirable as a condition to the issuance of Shares to Participant (or his or her estate), such issuance will not occur
unless and until such listing, registration, qualification, consent or approval will have been effected or obtained free of any conditions not acceptable to the Company. The Company will make all reasonable efforts to meet the requirements of any
such state or federal law or securities exchange and to obtain any such consent or approval of any such governmental authority. Assuming such compliance, for income tax purposes the Exercised Shares will be considered transferred to Participant on
the date the Option is exercised with respect to such Exercised Shares. 

  
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 12. Plan Governs. This Award Agreement is subject to all terms and provisions of the
Plan. In the event of a conflict between one or more provisions of this Award Agreement and one or more provisions of the Plan, the provisions of the Plan will govern. Capitalized terms used and not defined in this Award Agreement will have the
meaning set forth in the Plan. 
 13. Administrator Authority. The Administrator will have the power to interpret the
Plan and this Award Agreement and to adopt such rules for the administration, interpretation and application of the Plan as are consistent therewith and to interpret or revoke any such rules (including, but not limited to, the determination of
whether or not any Shares subject to the Option have vested). All actions taken and all interpretations and determinations made by the Administrator in good faith will be final and binding upon Participant, the Company and all other interested
persons. No member of the Administrator will be personally liable for any action, determination or interpretation made in good faith with respect to the Plan or this Award Agreement. 

14. Electronic Delivery. The Company may, in its sole discretion, decide to deliver any documents related to Options awarded under
the Plan or future options that may be awarded under the Plan by electronic means or request Participant’s consent to participate in the Plan by electronic means. Participant hereby consents to receive such documents by electronic delivery and
agrees to participate in the Plan through any on-line or electronic system established and maintained by the Company or another third party designated by the Company. 
 15. Captions. Captions provided herein are for convenience only and are not to serve as a basis for interpretation or construction of this Award Agreement. 

16. Agreement Severable. In the event that any provision in this Award Agreement will be held invalid or unenforceable, such
provision will be severable from, and such invalidity or unenforceability will not be construed to have any effect on, the remaining provisions of this Award Agreement. 
 17. Modifications to the Agreement. This Award Agreement constitutes the entire understanding of the parties on the subjects covered. Participant expressly warrants that he or she is not accepting
this Award Agreement in reliance on any promises, representations, or inducements other than those contained herein. Modifications to this Award Agreement or the Plan can be made only in an express written contract executed by a duly authorized
officer of the Company. Notwithstanding anything to the contrary in the Plan or this Award Agreement, the Company reserves the right to revise this Award Agreement as it deems necessary or advisable, in its sole discretion and without the consent of
Participant, to comply with Code Section 409A or to otherwise avoid imposition of any additional tax or income recognition under Code Section 409A in connection to this Option. 

18. Amendment, Suspension or Termination of the Plan. By accepting this Award, Participant expressly warrants that he or she has
received an Option under the Plan, and has received, read and understood a description of the Plan. Participant understands that the Plan is discretionary in nature and may be amended, suspended or terminated by the Company at any time. 

  
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 19. Governing Law. This Award Agreement will be governed by the laws of the State of
California, without giving effect to the conflict of law principles thereof. For purposes of litigating any dispute that arises under this Option or this Award Agreement, the parties hereby submit to and consent to the jurisdiction of the State of
California, and agree that such litigation will be conducted in the courts of San Diego County, California, or the federal courts for the United States for the Southern District of California, and no other courts, where this Option is made and/or to
be performed. 

  
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 EXHIBIT B 

ORGANOVO HOLDINGS, INC. 
 2012 EQUITY INCENTIVE PLAN 
 EXERCISE NOTICE 

Organovo Holdings, Inc. 
 6275 Nancy Ridge Dr.,
San Diego, CA 92121 
 San Diego, CA 92121 
 Attention: Chief Financial Officer 
 1. Exercise of Option. Effective as of
today,                     ,             , the undersigned (“Purchaser”) hereby
elects to purchase             shares (the “Shares”) of the Common Stock of Organovo Holdings, Inc. (the “Company”) under and pursuant to the 2012 Equity Incentive Plan
(the “Plan”) and the Stock Option Award Agreement dated             (the “Award Agreement”). The purchase price for the Shares will be
$            , as required by the Award Agreement. 
 2. Delivery
of Payment. Purchaser herewith delivers to the Company the full purchase price of the Shares and any required tax withholding to be paid in connection with the exercise of the Option. 

3. Representations of Purchaser. Purchaser acknowledges that Purchaser has received, read and understood the Plan and the Award
Agreement and agrees to abide by and be bound by their terms and conditions. 
 4. Rights as Stockholder. Until the
issuance (as evidenced by the appropriate entry on the books of the Company or of a duly authorized transfer agent of the Company) of the Shares, no right to vote or receive dividends or any other rights as a stockholder will exist with respect to
the Shares subject to the Option, notwithstanding the exercise of the Option. The Shares so acquired will be issued to Purchaser as soon as practicable after exercise of the Option. No adjustment will be made for a dividend or other right for which
the record date is prior to the date of issuance, except as provided in Section 15 of the Plan. 
 5. Tax
Consultation. Purchaser understands that Purchaser may suffer adverse tax consequences as a result of Purchaser’s purchase or disposition of the Shares. Purchaser represents that Purchaser has consulted with any tax consultants Purchaser
deems advisable in connection with the purchase or disposition of the Shares and that Purchaser is not relying on the Company for any tax advice. 
 6. Entire Agreement; Governing Law. The Plan and Award Agreement are incorporated herein by reference. This Exercise Notice, the Plan and the Award Agreement constitute the entire agreement of the
parties with respect to the subject matter hereof and supersede in their entirety all prior undertakings and agreements of the Company and Purchaser 

 
with respect to the subject matter hereof, and may not be modified adversely to the Purchaser’s interest except by means of a writing signed by the Company and Purchaser. This agreement is
governed by the internal substantive laws, but not the choice of law rules, of the State of California. 
  

					
	 Submitted by
	 		 	Accepted by
			
	PURCHASER	 		 	ORGANOVO HOLDINGS, INC.
			
	  
	 		 	  

	Signature	 		 	By
			
	  
	 		 	  

	Print Name	 		 	Title
			
	Residence Address:	 		 	
			
	  
	 		 	
			
	  
	 		 	
			
		 		 	  

		 		 	Date ReceivedFourth Supplemental Indenture

 Exhibit 4.1 
 FOURTH SUPPLEMENTAL INDENTURE 
 Fourth Supplemental Indenture (this
“Fourth Supplemental Indenture”), dated as of August 9, 2012, among Nationstar Mortgage Holdings Inc., Nationstar Sub1 LLC, Nationstar Sub2 LLC (together with Nationstar Mortgage Holdings Inc. and Nationstar Sub1 LLC, the
“Guaranteeing Parent Entities,” and each a “Guaranteeing Parent Entity”) and Wells Fargo Bank, National Association, as trustee (the “Trustee”). 

W I T N E S S E T H 
 WHEREAS, each of the Guaranteeing Parent Entities is a direct or indirect parent of Nationstar Mortgage LLC, a Delaware limited liability company (the “Company”) and Nationstar Capital
Corporation, a Delaware corporation (the “Co-Issuer” and, together with the Company, the “Issuers”); 
 WHEREAS, the Issuers and each of the Guarantors (as defined in the Indenture referred to below) have heretofore executed and delivered to the Trustee an indenture (the “Indenture”), dated
as of March 26, 2010, providing for the issuance of 10.875% Senior Notes due 2015 (the “Notes”); 

WHEREAS, the Indenture provides that under certain circumstances any direct or indirect parent of the Company shall execute and deliver
to the Trustee a supplemental indenture pursuant to which such direct or indirect parent shall unconditionally guarantee all of the Issuers’ Obligations under the Notes and the Indenture on the terms and conditions set forth herein and under
the Indenture (the “Note Guarantees”); and 
 WHEREAS, pursuant to Section 9.01 of the Indenture, the
Trustee is authorized to execute and deliver this Fourth Supplemental Indenture. 
 NOW THEREFORE, in consideration of the
foregoing and for other good and valuable consideration, the receipt of which is hereby acknowledged, the parties mutually covenant and agree for the equal and ratable benefit of the Holders of the Notes as follows: 

(1)        Capitalized Terms. Capitalized terms used herein without definition shall have
the meanings assigned to them in the Indenture. 
 (2)        Agreement to
Guarantee. Each of the Guaranteeing Parent Entities hereby agrees as follows: 

(a)        Along with all other Guarantors named in the Indenture (including pursuant to any
supplemental indentures), to jointly and severally unconditionally guarantee to each Holder of a Note authenticated and delivered by the Trustee and to the Trustee and its respective successors and assigns, irrespective of the validity and
enforceability of the Indenture, the Notes or the obligations of the Issuers hereunder or thereunder, that: 

   (i)         the principal of, interest, premium, if any, and
Additional Interest, if any, on the Notes shall be promptly paid in full when due, whether at maturity, by acceleration, redemption or otherwise, and interest on the overdue principal of and interest on the Notes, if any, if lawful, and all other
obligations of the Issuers to the Holders or the Trustee hereunder or thereunder shall be promptly paid in full or performed, all in accordance with the terms hereof and thereof; and 

    (ii)        in
case of any extension of time of payment or renewal of any Notes or any of such other obligations, that same shall be promptly paid in full when due or performed in accordance with the terms of the extension or renewal, whether at stated maturity,
by acceleration or otherwise. 
 Failing payment when due of any amount so guaranteed or any performance so guaranteed for
whatever reason, the Guarantors and the Guaranteeing Parent Entities shall be jointly and severally obligated to pay the same immediately. This is a guarantee of payment and not a guarantee of collection. 

(b)        The obligations hereunder shall be unconditional, irrespective of the validity,
regularity or enforceability of the Notes or the Indenture, the absence of any action to enforce the same, any waiver or consent by any Holder of the Notes with respect to any provisions hereof or thereof, the recovery of any judgment against the
Issuers or any Guarantors, any action to enforce the same or any other circumstance which might otherwise constitute a legal or equitable discharge or defense of a guarantor. 
 (c)        Each Guaranteeing Parent Entity hereby waives: diligence, presentment, demand of payment, filing of claims with a court in the event of insolvency or
bankruptcy of the Issuers, any right to require a proceeding first against the Issuers, protest, notice and all demands whatsoever. 
 (d)        This Note Guarantee shall not be discharged except by full payment or complete performance of the obligations contained in the Notes, the Indenture and
this Fourth Supplemental Indenture, and each Guaranteeing Parent Entity accepts all obligations of a Guarantor under the Indenture, including Article X of the Indenture (which is deemed incorporated in this Fourth Supplemental Indenture and
applicable to this Guarantee). Each Guaranteeing Parent Entity acknowledges that by executing this Fourth Supplemental Indenture, it will become a Guarantor under the Indenture and subject to all the terms and conditions applicable to Guarantors
contained therein. 
 (e)        If any Holder or the Trustee is required by any court
or otherwise to return to the Issuers, the Guarantors (including the Guaranteeing Parent Entities), or any custodian, trustee, liquidator or other similar official acting in relation to either the Issuers or the Guarantors, any amount paid either to
the Trustee or such Holder, this Note Guarantee, to the extent theretofore discharged, shall be reinstated in full force and effect. 
 (f)        No Guaranteeing Parent Entity shall be entitled to any right of subrogation in relation to the Holders in respect of any obligations guaranteed hereby
until payment in full of all obligations guaranteed hereby. 
 (g)        As between any
Guaranteeing Parent Entity, on the one hand, and the Holders and the Trustee, on the other hand, (x) the maturity of the obligations guaranteed hereby may be accelerated as provided in Article VI of the Indenture for the purposes of this Note
Guarantee, notwithstanding any stay, injunction or other prohibition preventing such acceleration in respect of the obligations guaranteed hereby, and (y) in the event of any declaration of acceleration of such obligations as provided in
Article VI of the Indenture, such obligations (whether or not due and payable) shall forthwith become due and payable by such Guaranteeing Parent Entity for the purpose of this Note Guarantee. 

  
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 (h)        Each Guaranteeing Parent Entity shall
have the right to seek contribution from any non-paying Guarantor so long as the exercise of such right does not impair the rights of the Holders under this Note Guarantee. 
 (i)        Pursuant to Section 10.02 of the Indenture, the obligations of each Guaranteeing Parent Entities shall be limited to the maximum amount as will,
after giving effect to such maximum amount and all other contingent and fixed liabilities of such Guaranteeing Parent Entity that are relevant under any applicable Bankruptcy Law or fraudulent conveyance laws and after giving effect to any
collections from, rights to receive contribution from or payments made by or on behalf of any other Guarantor in respect of the obligations of such other Guarantor under Article X of the Indenture, result in the obligations of such Guaranteeing
Parent Entity under this Note Guarantee not constituting a fraudulent conveyance or fraudulent transfer under applicable law. 

(j)        This Note Guarantee shall remain in full force and effect and continue to be effective
should any petition be filed by or against the Issuers for liquidation, reorganization, should the Issuers become insolvent or make an assignment for the benefit of creditors or should a receiver or trustee be appointed for all or any significant
part of the Issuers’ assets, and shall, to the fullest extent permitted by law, continue to be effective or be reinstated, as the case may be, if at any time payment and performance of the Notes are, pursuant to applicable law, rescinded or
reduced in amount, or must otherwise be restored or returned by any obligee on the Notes and Note Guarantee, whether as a “voidable preference”, “fraudulent transfer” or otherwise, all as though such payment or performance had
not been made. In the event that any payment or any part thereof, is rescinded, reduced, restored or returned, the Note shall, to the fullest extent permitted by law, be reinstated and deemed reduced only by such amount paid and not so rescinded,
reduced, restored or returned. 
 (k)        In case any provision of this Note
Guarantee shall be invalid, illegal or unenforceable, the validity, legality, and enforceability of the remaining provisions shall not in any way be affected or impaired thereby. 

(l)        This Note Guarantee shall be a general unsecured senior obligation of such
Guaranteeing Parent Entity, ranking pari passu with any other future unsubordinated Indebtedness of the Guaranteeing Parent Entity, if any. 
 (m)        Each payment to be made by each Guaranteeing Parent Entity in respect of this Note Guarantee shall be made without set-off, counterclaim, reduction or
diminution of any kind or nature. 
 (3)        Execution and Delivery. Each
Guaranteeing Parent Entity agrees that the Note Guarantee shall remain in full force and effect notwithstanding the absence of the endorsement of any notation of such Note Guarantee on the Notes. 

(4)        Merger, Consolidation or Sale of All or Substantially All Assets. 

(a)        No Guaranteeing Parent Entity may sell or otherwise dispose of all or substantially
all of its assets to, or consolidate with or merge with or into (whether or not such Guaranteeing Parent Entity is the surviving Person), another Person, other than the Issuers or another Guarantor, unless: 

  
 3 

 (i)        except in the case of a merger entered
into solely for the purpose of reincorporating a Guaranteeing Parent Entity in another jurisdiction, immediately after giving effect to that transaction, no Default or Event of Default shall have occurred and be continuing; and 

(ii)        either: 
 (A)        the Person acquiring the property in any such sale or disposition or the Person formed by or surviving any such consolidation or merger (if not the
Guaranteeing Parent Entity) assumes all the obligations of that Guaranteeing Parent Entity under the Indenture, its Note Guarantee and the applicable Registration Rights Agreement pursuant to this Fourth Supplemental Indenture; or 

(B)        the Net Proceeds of such sale or other disposition are either (i) applied in
accordance with Section 4.10(d) of the Indenture or (ii) not required to be applied in accordance with any provision of the Indenture. 
 (5)        Releases. 
 The Note
Guarantee of a Guaranteeing Parent Entity shall be automatically and unconditionally released and discharged, and no further action by such Guaranteeing Parent Entity, the Issuers or the Trustee is required for the release of such Guaranteeing
Parent Entity’s Note Guarantee, in the following circumstances: 
 (a)        in
connection with any sale, transfer or other disposition of all or substantially all of the assets of that Guaranteeing Parent Entity (including by way of merger or consolidation) to a Person that is not (either before or after giving effect to such
transaction) the Company or a Restricted Subsidiary of the Company, if the sale or other disposition does not violate Section 4.10 of the Indenture; 
 (b)         in connection with any sale, transfer or other disposition of all of the Capital Stock of such Guaranteeing Parent Entity (including by way of merger or
consolidation) to a Person that is not (either before or after giving effect to such transaction) the Company or a Restricted Subsidiary of the Company, if the sale or other disposition does not violate Section 4.10 of the Indenture;

 (c)         if the Company designates any Restricted Subsidiary of the Company that
is a Guarantor to be an Unrestricted Subsidiary of the Company in accordance with Section 4.17 of the Indenture; or 

(d)        upon the exercise of Legal Defeasance by the Issuers or pursuant to Article XI of the
Indenture; and 
 in connection with such release, either of the Issuers shall deliver to the Trustee an Officers’ Certificate of such
Guarantor confirming the effective date of such release and stating that all conditions precedent provided for in this Indenture relating to such transaction have been complied with. 

(6)        No Recourse Against Others. No director, officer, employee, incorporator or
stockholder of any Guaranteeing Parent Entity shall have any liability for any obligations of the Issuers or the Guarantors (including such Guaranteeing Parent Entity), respectively, under the Notes, the Note Guarantees, the Indenture or this Fourth
Supplemental Indenture or for any claim based on, in respect of, 

  
 4 

 
or by reason of, such obligations or their creation; provided that the foregoing shall not limit any Guarantor’s obligations under its Note Guarantees. Each Holder by accepting Notes
waives and releases all such liability. The waiver and release are part of the consideration for issuance of the Notes. 

(7)        Governing Law. THIS FOURTH SUPPLEMENTAL INDENTURE WILL BE GOVERNED BY AND
CONSTRUED IN ACCORDANCE WITH THE LAWS OF THE STATE OF NEW YORK. 

(8)        Counterparts. The parties may sign any number of copies of this Fourth
Supplemental Indenture. Each signed copy shall be an original, but all of them together represent the same agreement. 

(9)        Effect of Headings. The Section headings herein are for convenience only and
shall not affect the construction hereof. 
 (10)        The Trustee. The Trustee
shall not be responsible in any manner whatsoever for or in respect of the validity or sufficiency of this Fourth Supplemental Indenture or for or in respect of the recitals contained herein, all of which recitals are made solely by the Guaranteeing
Parent Entities. 
 (11)        Subrogation. Each Guaranteeing Parent Entity
shall be subrogated to all rights of Holders of Notes against the Issuers in respect of any amounts paid by such Guaranteeing Parent Entity pursuant to the provisions of Section 2 hereof and Section 10.01 of the Indenture; provided
that, if an Event of Default has occurred and is continuing, no Guaranteeing Subsidiary shall be entitled to enforce or receive any payments arising out of, or based upon, such right of subrogation until all amounts then due and payable by the
Issuers under the Indenture or the Notes shall have been paid in full. 

(12)        Benefits Acknowledged. The Note Guarantee of each Guaranteeing Parent Entity
is subject to the terms and conditions set forth in the Indenture. The Guaranteeing Parent Entities acknowledge that they will receive direct and indirect benefits from the financing arrangements contemplated by the Indenture and this Fourth
Supplemental Indenture and that the guarantee and waivers made respectively by each pursuant to its Note Guarantee are knowingly made in contemplation of such benefits. 
 (13)        Successors. All agreements of each Guaranteeing Parent Entity in this Fourth Supplemental Indenture shall bind the Successors of such
Guaranteeing Parent Entity, except as otherwise set forth in this Fourth Supplemental Indenture. All agreements of the Trustee in this Fourth Supplemental Indenture shall bind its successors. 

(14)        Effectiveness. This Fourth Supplemental Indenture shall be effective and have
full force and effect from June 30, 2012. 
 [signature page follows] 

  
 5 

 IN WITNESS WHEREOF, the parties hereto have caused this Fourth Supplemental Indenture to be
duly executed, all as of the date first above written. 
  

			
	NATIONSTAR MORTGAGE HOLDINGS INC.
		
	By:	 	 /s/ Jay Bray

		 	Name: Jay Bray
		 	Title: Director and Chief Executive Officer
	
	NATIONSTAR SUB1 LLC
		
	By:	 	 /s/ Jay Bray

		 	Name: Jay Bray
		 	Title: Manager
	
	NATIONSTAR SUB2 LLC
		
	By:	 	 /s/ Jay Bray

		 	Name: Jay Bray
		 	Title: Manager
	
	 WELLS FARGO BANK, NATIONAL
ASSOCIATION,
 as Trustee

		
	By:	 	 /s/ Martin G. Reed

		 	Name: Martin G. Reed
		 	Title: Vice President

  
 6

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