Document:

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                                                               Exhibit 10.31

                          TRANSKARYOTIC THERAPIES, INC.

          2001 NON-OFFICER, NON-DIRECTOR EMPLOYEE STOCK INCENTIVE PLAN

1.       PURPOSE

         The purpose of this 2001 Non-Officer, Non-Director Employee Stock
Incentive Plan (the "Plan") of Transkaryotic Therapies, Inc., a Delaware
corporation (the "Company"), is to advance the interests of the Company's
stockholders by enhancing the Company's ability to attract, retain and
motivate persons who make (or are expected to make) important contributions
to the Company by providing such persons with equity ownership opportunities
and performance-based incentives and thereby better aligning the interests of
such persons with those of the Company's stockholders. Except where the
context otherwise requires, the term "Company" shall include any of the
Company's present or future parent or subsidiary corporations as defined in
Sections 424(e) or (f) of the Internal Revenue Code of 1986, as amended, and
any regulations promulgated thereunder (the "Code"), and any other business
venture (including, without limitation, joint venture or limited liability
company) in which the Company has a significant interest, as determined by
the Board of Directors of the Company (the "Board").

2.       ELIGIBILITY

         All of the Company's employees (and any individuals who have accepted
an offer for employment), other than those employees who are also officers or
directors of the Company, and all of the Company's consultants and advisors are
eligible to be granted options or restricted stock awards (each, an "Award")
under the Plan. Each person who has been granted an Award under the Plan shall
be deemed a "Participant."

3.       ADMINISTRATION AND DELEGATION

         (a) ADMINISTRATION BY BOARD OF DIRECTORS. The Plan will be administered
by the Board. The Board shall have authority to grant Awards and to adopt, amend
and repeal such administrative rules, guidelines and practices relating to the
Plan as it shall deem advisable. The Board may correct any defect, supply any
omission or reconcile any inconsistency in the Plan or any Award in the manner
and to the extent it shall deem expedient to carry the Plan into effect and it
shall be the sole and final judge of such expediency. All decisions by the Board
shall be made in the Board's sole discretion and shall be final and binding on
all persons having or claiming any interest in the Plan or in any Award. No
director or person acting pursuant to the authority delegated by the Board shall
be liable for any action or determination relating to or under the Plan made in
good faith.

         (b) APPOINTMENT OF COMMITTEES. To the extent permitted by applicable
law, the Board may delegate any or all of its powers under the Plan to one or
more committees or subcommittees of the Board (a "Committee"). All references in
the Plan to the "Board" shall mean the Board or a Committee of the Board to the
extent that the Board's powers or authority under the Plan have been delegated
to such Committee.

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         (c) Subject to the provisions of the Plan, the Board shall have
complete authority, in its sole discretion, to make or to select the manner of
making any and all determinations required for the operation of the Plan, and
without limiting the generality of the foregoing, shall have the authority to:

                  (1) grant Awards to eligible individuals, pursuant to the
terms of the Plan;

                  (2) determine whether and to what extent Awards are to be
granted hereunder;

                  (3) determine the number of shares of Common Stock to be
covered by each Award granted hereunder;

                  (4) determine the terms and conditions, not inconsistent with
the terms of the Plan, of any Award (which need not be identical in every case),
including, but not limited to, the share price and any restriction or
limitation, or any vesting acceleration or forfeiture waiver regarding any Award
and the shares of Common Stock relating thereto, based on such factors as the
Board shall determine;

                  (5) determine whether and under what circumstances an Option
may be settled, as provided in Section 5(e);

                  (6) determine whether and under what circumstances an Option
may be exercised without a payment of cash as provided in Section 5(e); and

                  (7) determine whether, to what extent and under what
circumstances Common Stock and other amounts payable with respect to an Award
under this Plan shall be deferred either automatically or at the election of the
Participant.

         In making such determinations, the Board may take into account the
nature of the services rendered by the respective individuals, their present and
potential contributions to the success of the Company, and such other factors as
the Board in its discretion shall deem relevant. Subject to the provisions of
the Plan, the Board shall also have complete authority, in its sole discretion,
to interpret the Plan, to prescribe, amend and rescind rules and regulations
relating to it, to determine the terms and provisions of any Award issued under
the Plan (and any agreements relating thereto), to resolve all disputes arising
under the Plan, and to make all other determinations necessary or advisable for
the administration of the Plan.

4.       STOCK AVAILABLE FOR AWARDS

         Subject to adjustment under Section 7, Awards may be made under the
Plan for up to 2,000,000 shares of common stock, $0.01 par value per share,
of the Company (the "Common Stock"). If (i) any Award expires or is
terminated, surrendered or canceled without having been fully exercised, (ii)
any Award is forfeited in whole or in part, (iii) any Award results in any
shares of Common Stock not being issued or (iv) the shares of Common Stock
issued pursuant to any Award are repurchased by the Company (including
without limitation shares of Common Stock issued upon exercise of an Option
(as hereinafter defined) that are subsequently

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repurchased by the Company pursuant to a contractual repurchase right or
otherwise), the unused shares of Common Stock covered by such Award shall again
be available for the grant of Awards under the Plan. Shares issued under the
Plan may consist in whole or in part of authorized but unissued shares or
treasury shares.

5.       NON-STATUTORY STOCK OPTIONS

         (a) GENERAL. The Board may grant non-statutory stock options to
purchase Common Stock (each, an "Option") and determine the number of shares of
Common Stock to be covered by each Option, the exercise price of each Option and
the conditions and limitations applicable to the exercise of each Option,
including conditions relating to applicable federal or state securities laws, as
it considers necessary or advisable. No Option granted under the Plan shall be
intended to be an "incentive stock option" as defined in Section 422 of the
Code.

         (b) EXERCISE PRICE. The Board shall establish the exercise price at the
time each Option is granted and specify it in the applicable instrument
evidencing the grant of the Option.

         (c) DURATION OF OPTIONS. Each Option shall be exercisable at such times
and subject to such terms and conditions as the Board may specify in the
applicable instrument evidencing the grant of the Option.

         (d) EXERCISE OF OPTION. Options may be exercised by delivery to the
Company of a written notice of exercise signed by the proper person or by any
other form of notice (including electronic notice) approved by the Board
together with payment in full as specified in Section 5(e) for the number of
shares for which the Option is exercised.

         (e) PAYMENT UPON EXERCISE. Common Stock purchased upon the exercise of
an Option granted under the Plan shall be paid for as follows:

                  (1) in cash or by check, payable to the order of the Company;

                  (2) except as the Board may, in its sole discretion, otherwise
provide in an instrument evidencing the grant of an Option, by (i) delivery of
an irrevocable and unconditional undertaking by a creditworthy broker to deliver
promptly to the Company sufficient funds to pay the exercise price and any
required tax withholding or (ii) delivery by the Participant to the Company of a
copy of irrevocable and unconditional instructions to a creditworthy broker to
deliver promptly to the Company cash or a check sufficient to pay the exercise
price and any required tax withholding;

                  (3) except as the Board may, in its sole discretion, otherwise
provide in an instrument evidencing the grant of an Option, when the Common
Stock is registered under the Securities Exchange Act of 1934, as amended (the
"Exchange Act"), by delivery of shares of Common Stock owned by the Participant
valued at their fair market value as determined by (or in a manner approved by)
the Board in good faith ("Fair Market Value"), provided (i) such method of
payment is then permitted under applicable law and (ii) such Common Stock, if
acquired directly from the Company, was owned by the Participant at least six
months prior to such delivery;

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                  (4) to the extent permitted by the Board, in its sole
discretion, by (i) delivery of a full recourse promissory note of the
Participant to the Company on terms determined by the Board, or (ii) payment of
such other lawful consideration as the Board may determine; or

                  (5) by any combination of the above-permitted forms of
payment.

         (f) SUBSTITUTE OPTIONS. In connection with a merger or consolidation of
an entity with and into the Company or the acquisition by the Company of
property or stock of an entity, the Board may grant Options in substitution for
any options or other stock or stock-based awards granted by such entity or an
affiliate thereof. Substitute Options may be granted on such terms as the Board
deems appropriate in the circumstances, notwithstanding any limitations on
Options contained in the other sections of this Section 5 or in Section 2
hereof.

6.       RESTRICTED STOCK

         (a) GRANTS. The Board may grant Awards entitling Participants to
acquire shares of Common Stock, subject to the right of the Company to
repurchase all or part of such shares at their issue price or other stated or
formula price (or to require forfeiture of such shares if issued at no cost)
from the Participant in the event that conditions specified by the Board in the
applicable Award are not satisfied prior to the end of the applicable
restriction period or periods established by the Board for such Award (each, a
"Restricted Stock Award").

         (b) TERMS AND CONDITIONS. The Board shall determine the terms and
conditions of any such Restricted Stock Award, including the conditions for
repurchase (or forfeiture) and the issue price, if any.

7.       ADJUSTMENTS FOR CHANGES IN COMMON STOCK AND CERTAIN OTHER EVENTS

         (a) CHANGES IN CAPITALIZATION. In the event of any stock split, reverse
stock split, stock dividend, recapitalization, combination of shares,
reclassification of shares, spin-off or other similar change in capitalization
or event, or any distribution to holders of Common Stock other than a normal
cash dividend, (i) the number and class of securities available under this Plan,
(ii) the number and class of securities and exercise price per share subject to
each outstanding Option, and (iii) the repurchase price per share subject to
each outstanding Restricted Stock Award shall be appropriately adjusted by the
Company (or substituted Awards may be made, if applicable) to the extent the
Board shall determine, in good faith, that such an adjustment (or substitution)
is necessary and appropriate. If this Section 7(a) applies and Section 7(c) also
applies to any event, Section 7(c) shall be applicable to such event, and this
Section 7(a) shall not be applicable.

         (b) LIQUIDATION OR DISSOLUTION. In the event of a proposed liquidation
or dissolution of the Company, the Board shall upon written notice to the
Participants provide that all then unexercised Options will (i) become
exercisable in full as of a specified time at least 10 business days prior to
the effective date of such liquidation or dissolution and (ii) terminate
effective upon such liquidation or dissolution, except to the extent exercised
before such effective date. The Board may specify the effect of a liquidation or
dissolution on any Restricted Stock Award granted under the Plan at the time of
the grant.

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         (c) REORGANIZATION EVENTS.

                  (1) DEFINITION. A "Reorganization Event" shall mean: (a) any
merger or consolidation of the Company with or into another entity as a result
of which all of the Common Stock of the Company is converted into or exchanged
for the right to receive cash, securities or other property or (b) any exchange
of all of the Common Stock of the Company for cash, securities or other property
pursuant to a share exchange transaction.

                  (2) CONSEQUENCES OF A REORGANIZATION EVENT ON OPTIONS. Upon
the occurrence of a Reorganization Event, or the execution by the Company of any
agreement with respect to a Reorganization Event, the Board shall provide that
all outstanding Options shall be assumed, or equivalent options shall be
substituted, by the acquiring or succeeding corporation (or an affiliate
thereof). For purposes hereof, an Option shall be considered to be assumed if,
following consummation of the Reorganization Event, the Option confers the right
to purchase, for each share of Common Stock subject to the Option immediately
prior to the consummation of the Reorganization Event, the consideration
(whether cash, securities or other property) received as a result of the
Reorganization Event by holders of Common Stock for each share of Common Stock
held immediately prior to the consummation of the Reorganization Event (and if
holders were offered a choice of consideration, the type of consideration chosen
by the holders of a majority of the outstanding shares of Common Stock);
provided, however, that if the consideration received as a result of the
Reorganization Event is not solely common stock of the acquiring or succeeding
corporation (or an affiliate thereof), the Company may, with the consent of the
acquiring or succeeding corporation, provide for the consideration to be
received upon the exercise of Options to consist solely of common stock of the
acquiring or succeeding corporation (or an affiliate thereof) equivalent in fair
market value to the per share consideration received by holders of outstanding
shares of Common Stock as a result of the Reorganization Event.

Notwithstanding the foregoing, if the acquiring or succeeding corporation (or an
affiliate thereof) does not agree to assume, or substitute for, such Options,
then the Board shall, upon written notice to the Participants, provide that all
then unexercised Options will become exercisable in full as of a specified time
prior to the Reorganization Event and will terminate immediately prior to the
consummation of such Reorganization Event, except to the extent exercised by the
Participants before the consummation of such Reorganization Event; provided,
however, that in the event of a Reorganization Event under the terms of which
holders of Common Stock will receive upon consummation thereof a cash payment
for each share of Common Stock surrendered pursuant to such Reorganization Event
(the "Acquisition Price"), then the Board may instead provide that all
outstanding Options shall terminate upon consummation of such Reorganization
Event and that each Participant shall receive, in exchange therefor, a cash
payment equal to the amount (if any) by which (A) the Acquisition Price
multiplied by the number of shares of Common Stock subject to such outstanding
Options (whether or not then exercisable), exceeds (B) the aggregate exercise
price of such Options. To the extent all or any portion of an Option becomes
exercisable solely as a result of the first sentence of this paragraph, upon
exercise of such Option the Participant shall receive shares subject to a right
of repurchase by the Company or its successor at the Option exercise price. Such
repurchase right (1) shall lapse at the same rate as the Option would have
become exercisable under its terms and (2) shall

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not apply to any shares subject to the Option that were exercisable under its
terms without regard to the first sentence of this paragraph.

                  (3) CONSEQUENCES OF A REORGANIZATION EVENT ON RESTRICTED STOCK
AWARDS. Upon the occurrence of a Reorganization Event, the repurchase and other
rights of the Company under each outstanding Restricted Stock Award shall inure
to the benefit of the Company's successor and shall apply to the cash,
securities or other property which the Common Stock was converted into or
exchanged for pursuant to such Reorganization Event in the same manner and to
the same extent as they applied to the Common Stock subject to such Restricted
Stock Award.

8.       GENERAL PROVISIONS APPLICABLE TO AWARDS

         (a) TRANSFERABILITY OF AWARDS. Except as the Board may otherwise
determine or provide in an Award, Awards shall not be sold, assigned,
transferred, pledged or otherwise encumbered by the person to whom they are
granted, either voluntarily or by operation of law, except by will or the laws
of descent and distribution, and, during the life of the Participant, shall be
exercisable only by the Participant. References to a Participant, to the extent
relevant in the context, shall include references to authorized transferees.

         (b) DOCUMENTATION. Each Award shall be evidenced in such form (written,
electronic or otherwise) as the Board shall determine. Each Award may contain
terms and conditions in addition to those set forth in the Plan.

         (c) BOARD DISCRETION. Except as otherwise provided by the Plan, each
Award may be made alone or in addition or in relation to any other Award. The
terms of each Award need not be identical, and the Board need not treat
Participants uniformly.

         (d) TERMINATION OF STATUS. The Board shall determine the effect on an
Award of the disability, death, retirement, authorized leave of absence or other
change in the employment or other status of a Participant and the extent to
which, and the period during which, the Participant, the Participant's legal
representative, conservator, guardian or Designated Beneficiary may exercise
rights under the Award.

         (e) WITHHOLDING. Each Participant shall pay to the Company, or make
provision satisfactory to the Board for payment of, any taxes required by law to
be withheld in connection with Awards to such Participant no later than the date
of the event creating the tax liability. Except as the Board may otherwise
provide in an Award, when the Common Stock is registered under the Exchange Act,
Participants may satisfy such tax obligations in whole or in part by delivery of
shares of Common Stock, including shares retained from the Award creating the
tax obligation, valued at their Fair Market Value; provided, however, that the
total tax withholding where stock is being used to satisfy such tax obligations
cannot exceed the Company's minimum statutory withholding obligations (based on
minimum statutory withholding rates for federal and state tax purposes,
including payroll taxes, that are applicable to such supplemental taxable
income). The Company may, to the extent permitted by law, deduct any such tax
obligations from any payment of any kind otherwise due to a Participant.

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         (f) AMENDMENT OF AWARD. The Board may amend, modify or terminate any
outstanding Award, including but not limited to, substituting therefor another
Award of the same or a different type and changing the date of exercise or
realization, provided that the Participant's consent to such action shall be
required unless the Board determines that the action, taking into account any
related action, would not materially and adversely affect the Participant.

         (g) CONDITIONS ON DELIVERY OF STOCK. The Company will not be obligated
to deliver any shares of Common Stock pursuant to the Plan or to remove
restrictions from shares previously delivered under the Plan until (i) all
conditions of the Award have been met or removed to the satisfaction of the
Company, (ii) in the opinion of the Company's counsel, all other legal matters
in connection with the issuance and delivery of such shares have been satisfied,
including any applicable securities laws and any applicable stock exchange or
stock market rules and regulations, and (iii) the Participant has executed and
delivered to the Company such representations or agreements as the Company may
consider appropriate to satisfy the requirements of any applicable laws, rules
or regulations.

         (h) ACCELERATION. The Board may at any time provide that any Award
shall become immediately exercisable in full or in part, free of some or all
restrictions or conditions, or otherwise realizable in full or in part, as the
case may be.

9.       MISCELLANEOUS

         (a) NO RIGHT TO EMPLOYMENT OR OTHER STATUS. No person shall have any
claim or right to be granted an Award, and the grant of an Award shall not be
construed as giving a Participant the right to continued employment or any other
relationship with the Company. The Company expressly reserves the right at any
time to dismiss or otherwise terminate its relationship with a Participant free
from any liability or claim under the Plan, except as expressly provided in the
applicable Award.

         (b) NO RIGHTS AS STOCKHOLDER. Subject to the provisions of the
applicable Award, no Participant or Designated Beneficiary shall have any rights
as a stockholder with respect to any shares of Common Stock to be distributed
with respect to an Award until becoming the record holder of such shares.
Notwithstanding the foregoing, in the event the Company effects a split of the
Common Stock by means of a stock dividend and the exercise price of and the
number of shares subject to such Option are adjusted as of the date of the
distribution of the dividend (rather than as of the record date for such
dividend), then an optionee who exercises an Option between the record date and
the distribution date for such stock dividend shall be entitled to receive, on
the distribution date, the stock dividend with respect to the shares of Common
Stock acquired upon such Option exercise, notwithstanding the fact that such
shares were not outstanding as of the close of business on the record date for
such stock dividend.

         (c) EFFECTIVE DATE AND TERM OF PLAN. The Plan is effective as of
March 14, 2001, the date on which it was adopted by the Board (the "Effective
Date"). No Awards shall be granted under the Plan after the completion of ten
years from the Effective Date, but Awards previously granted may extend beyond
that date.

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         (d) AMENDMENT OF PLAN. The Board may amend, suspend or terminate the
Plan or any portion thereof at any time.

         (e) GOVERNING LAW. The provisions of the Plan and all Awards made
hereunder shall be governed by and interpreted in accordance with the laws of
the State of Delaware, without regard to any applicable conflicts of law.

                                    * * * * *

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                                                                 EXHIBIT 10.32

                     TRANSKARYOTIC THERAPIES, INC. DEFERRED
                               COMPENSATION PLAN

                           EFFECTIVE - OCTOBER 1, 2000

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            TRANSKARYOTIC THERAPIES, INC. DEFERRED COMPENSATION PLAN

                                Table of Contents

ARTICLE I.................................................................    4
   1.1 Statement of Purpose...............................................    4

ARTICLE II................................................................    5

DEFINITIONS...............................................................    5
   2.1 Account............................................................    5
   2.2 Base Salary........................................................    5
   2.3 Beneficiary........................................................    5
   2.4 Board..............................................................    5
   2.5 Change in Control..................................................    6
   2.6 Code...............................................................    6
   2.7 Committee..........................................................    6
   2.8 Compensation.......................................................    7
   2.9 Company............................................................    7
   2.10 Credited Service..................................................    7
   2.11 Deferral Account..................................................    7
   2.12 Deferral Benefit..................................................    7
   2.13 Deferral Election.................................................    7
   2.14 Disability........................................................    7
   2.15 Early Retirement..................................................    7
   2.16 Eligible Employee.................................................    7
   2.17 Employer..........................................................    8
   2.18 Hardship Withdrawal - "Haircut" Provisions........................    8
   2.19 Investment Return Rate............................................    8
   2.20 Participant.......................................................    8
   2.21 Participation Agreement...........................................    8
   2.22 Plan..............................................................    8
   2.23 Plan Year.........................................................    8
   2.24 Retirement........................................................    8
   2.25 Valuation Date....................................................    9

ELIGIBILITY AND PARTICIPATION.............................................   10
   3.1 Eligibility........................................................   10
   3.2 Participation......................................................   10
   3.3 Change in Participation Status.....................................   10
   3.4 Termination of Participation.......................................   10
   3.5 Ineligible Participant.............................................   11

ARTICLE IV................................................................   12

DEFERRAL OF COMPENSATION; COMPANY CREDITS.................................   12
   4.1 Amount of Deferral.................................................   12
   4.2 Crediting Deferred Compensation....................................   12
   4.3 Company Contribution Account.......................................   12

ARTICLE V.................................................................   13

BENEFIT ACCOUNTS..........................................................   13

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   5.1 Valuation of Account...............................................   13
   5.2 Crediting of Investment Return.....................................   13
   5.3 Statement of Accounts..............................................   13
   5.4 Vesting of Account.................................................   14
   5.5 Investment Vehicles................................................   14

ARTICLE VI................................................................   15

PAYMENT OF BENEFITS.......................................................   15
   6.1 Payment of Deferral Benefit upon Disability or Retirement..........   15
   6.2 Payment of Deferral Benefit upon Termination.......................   15
   6.3 Payments to Beneficiaries upon Participant Death...................   15
   6.4 Hardship Withdrawal - "Haircut" Provisions.........................   15
   6.5 In-Service Distribution............................................   16
   6.6  Form of Payment...................................................   16
   6.7 Commencement of Payments...........................................   16
   6.8 Small Benefit......................................................   16

BENEFICIARY DESIGNATION...................................................   17
   7.1 Beneficiary Designation............................................   17
   7.2 Change of Beneficiary Designation..................................   17
   7.3 No Designation.....................................................   17
   7.4 Effect of Payment..................................................   17

ARTICLE VIII..............................................................   18

ADMINISTRATION............................................................   18
    8.1 Committee.........................................................   18
    8.2 Agents............................................................   18
    8.3 Binding Effect of Decisions.......................................   18
    8.4 Indemnification of Committee......................................   18

ARTICLE IX................................................................   19

AMENDMENT AND TERMINATION OF PLAN.........................................   19
    9.1 Amendment.........................................................   19
    9.2 Termination.......................................................   19
    9.3 Change in Control.................................................   19

ARTICLE X.................................................................   20

MISCELLANEOUS.............................................................   20
    10.1 Funding..........................................................   20
    10.2 Nonassignability.................................................   20
    10.3 Captions.........................................................   21
    10.4 Governing Law....................................................   21
    10.5 Successors.......................................................   21
    10.6 Right to Continued Service.......................................   21

EXHIBIT A.................................................................   22

EXHIBIT B.................................................................   23

EXHIBIT C.................................................................   24

EXHIBIT D.................................................................   25

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                                    ARTICLE I

1.1 STATEMENT OF PURPOSE

This is the  Transkaryotic  Therapies,  Inc.  Deferred  Compensation  Plan  (the
"Plan")  made in the form of this Plan and in  related  agreements  between  the
Employer and certain management or highly compensated employees.  The purpose of
the Plan is to  provide  management  and  highly  compensated  employees  of the
Employer with the option to defer the receipt of portions of their  compensation
payable for services rendered to the Employer. It is intended that the Plan will
assist in attracting  and retaining  qualified  individuals to serve as officers
and managers of the Employer. The Plan is effective as of October 1, 2000.

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                                   ARTICLE II

DEFINITIONS

When used in this Plan and initially capitalized, the following words and
phrases shall have the meanings indicated:

2.1 ACCOUNT.

"Account" means the sum of a Participant's Deferral Account.

2.2 BASE SALARY.

"Base Salary" means a Participant's base earnings paid by an Employer to a
Participant without regard to any increases or decreases in base earnings as a
result of (i) an election to defer base earnings under this Plan or (ii) an
election between benefits or cash provided under a Plan of an Employer
maintained pursuant to Section 125 or 401(k) of the Code and as limited in
Exhibit B attached hereto.

2.3 BENEFICIARY.

"Beneficiary" means the person or persons designated or deemed to be designated
by the Participant pursuant to Article VII to receive benefits payable under the
Plan in the event of the Participant's death.

2.4 BOARD.

"Board" means the Board of Directors of the Company.

                                                                               5
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2.5 CHANGE IN CONTROL.

     A "Change in Control" of the Company shall occur or be deemed to have
occurred in the event that:

     (i)  any "person", as such term is used in Sections 13(d) and 14(d) of the
          Securities Exchange Act of 1934, as amended (the "Exchange Act") (a
          "Person"), other than the Company, any trustee or other fiduciary
          holding securities under an employee benefit plan of the Company, or
          any corporation owned directly or indirectly by the stockholders of
          the Company in substantially the same proportion as their ownership of
          stock of the Company, acquires "beneficial ownership" (as defined in
          Rule 13d-3 under the Exchange Act) of securities of the Company
          representing 50% or more of the combined voting power of the Company's
          then outstanding securities (other than through an acquisition of
          securities directly from the Company);

     (ii) individuals who, as of October 1, 2000, constitute the Board (the
          "Incumbent Board") cease for any reason to constitute at least a
          majority of the Board; provided, however, that any individual becoming
          a director subsequent to October 1, 2000, whose election, or
          nomination for election by the Company's stockholders, was approved by
          a vote of at least a majority of the directors then comprising the
          Incumbent Board shall be considered as though such individual were a
          member of the Incumbent Board, but excluding, for this purpose, any
          such individual whose initial assumption of office occurs as a result
          of an actual or threatened election contest with respect to the
          election or removal of directors or other actual or threatened
          solicitation of proxies or consents by or on behalf of a Person other
          than the Board;

     (iii) the stockholders of the Company approve a merger or consolidation of
          the Company with any other entity, other than (A) a merger or
          consolidation which would result in the voting securities of the
          Company outstanding immediately prior thereto continuing to represent
          (either by remaining outstanding or by being converted into voting
          securities of the surviving entity) more than 50% of the combined
          voting power of the voting securities of the Company or such surviving
          entity outstanding immediately after such merger or consolidation or
          (B) a merger or consolidation effected to implement a recapitalization
          of the Company (or similar transaction) in which no Person acquires
          more than 50% of the combined voting power of the Company's then
          outstanding securities; or

     (iv) the stockholders of the Company approve a plan of complete liquidation
          of the Company or an agreement for the sale or disposition by the
          Company of all or substantially all of the Company's assets.

2.6 CODE.

"Code" means the Internal Revenue Code of 1986, as amended.

2.7 COMMITTEE.

"Committee" has the meaning set forth in Section 8.1.

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2.8 COMPENSATION.

"Compensation" means the Base Salary and Bonus payable with respect to an
Eligible Employee for each plan year.

2.9 COMPANY.

"Company" means Transkaryotic Therapies, Inc. (TKT), affiliate companies and any
successor thereto.

2.10 CREDITED SERVICE.

"Credited Service" means the sum of all periods of a Participant's employment by
the Company or a Selected Affiliate for which service credit is given under the
Transkaryotic Therapies, Inc. 401(k) Plan.

2.11 DEFERRAL ACCOUNT.

"Deferral Account" means the account maintained on the books of the Employer for
the purpose of accounting for the amount of Compensation that each Participant
elects to defer under the Plan and for the amount of investment return credited
thereto for each Participant pursuant to Article V.

2.12 DEFERRAL BENEFIT.

"Deferral Benefit" means the benefit payable to a Participant or his or her
Beneficiary pursuant to Article VI.

2.13 DEFERRAL ELECTION.

"Deferral Election" means the written election made by a Participant to defer
Compensation pursuant to Article IV.

2.14 DISABILITY.

"Disability" means a Participant's Disability as defined under the Company's
Long Term Disability Plan or its successors.

2.15 EARLY RETIREMENT.

"Early Retirement" will be as granted by the Committee at its sole discretion.

2.16 ELIGIBLE EMPLOYEE.

"Eligible Employee" means a highly compensated or management employee of the
Company who is designated by the Committee, by name or group or description, in
accordance with Section 3.1 as eligible to participate in the Plan.

                                                                               7
<PAGE>

2.17 EMPLOYER.

"Employer" means, with respect to a Participant, the Company or the Selected
Affiliate which pays such Participant's Compensation.

2.18 HARDSHIP WITHDRAWAL - "HAIRCUT" PROVISIONS.

"Hardship Withdrawal - Haircut Provisions" has the meaning set forth in Section
6.4.

2.19 INVESTMENT RETURN RATE.

"Investment Return Rate" means:

     (a)  In the case of an investment named in Exhibit C of a fixed income
          nature, the interest deemed to be credited,

     (b)  In the case of an investment named in Exhibit C of an equity
          investment nature, the increase and decrease in deemed value
          and dividends deemed to be credited.

2.20 PARTICIPANT.

"Participant" means any Eligible Employee who elects to participate by filing a
Participation Agreement or who is automatically enrolled as provided in Section
3.2.

2.21 PARTICIPATION AGREEMENT.

"Participation Agreement" means the agreement filed by a Participant, in the
form prescribed by the Committee, pursuant to Section 3.2.

2.22 PLAN.

"Plan" means the Transkaryotic Therapies, Inc. Deferred Compensation Plan, as
amended from time to time.

2.23 PLAN YEAR.

"Plan Year" means a twelve-month period commencing January 1 and ending the
following December 31.

2.24 RETIREMENT

"Retirement" means the termination of employment of a Participant who has
reached age 55 and has completed ten (10) or more years of service.

                                                                               8
<PAGE>

2.25 VALUATION DATE.

"Valuation Date" means a date on which the amount of a Participant's Account is
valued as provided in Article V. The Valuation Date shall be the last day of
each month and any other date determined by the Committee.

                                                                               9
<PAGE>

                                   ARTICLE III

ELIGIBILITY AND PARTICIPATION

3.1 ELIGIBILITY.

Eligibility to participate in the Plan is limited to Eligible Employees. From
time to time, and subject to Section 3.4, the Committee shall prepare, and
attach to the Plan as Exhibit D, a complete list of the Eligible Employees, by
individual name or by reference to an identifiable group of persons or by
descriptions of the components of compensation of an individual which would
qualify individuals which are eligible to participate and all of whom shall be a
select group of management or highly compensated employees.

3.2 PARTICIPATION.

Participation in the Plan shall be limited to Eligible Employees who elect to
participate in the Plan by filing a Participation Agreement with the Committee.
An Eligible Employee shall commence participation in the Plan upon the first day
of his or her first payroll period commencing for the Plan Year following the
receipt of his or her Participation Agreement by the Committee. In the case of
an employee's initial year of participation in the Plan, the Participant may
make a Deferral Election with respect to compensation for services to be
performed subsequent to such Deferral Election, provided such election is made
no later than 30 days after the date the Participant first becomes eligible for
the Plan. An employee shall commence participation in the Plan for the current
Plan Year upon the first day of his or her first payroll period following the
receipt of his or her Participation Agreement by the Committee.

3.3 CHANGE IN PARTICIPATION STATUS.

During the election period each December, a Participant may change a previously
elected percentage of deferral of base salary and commission. Changes will only
become effective as of the beginning of the next Plan Year following receipt of
the change in election by the Committee and in accordance with the Company's
prevailing administrative procedures. With the exception of the initial plan
year, a Participant must make a Bonus deferral election by filing a written
notice thereof with the Committee prior to June 30th of the year preceding the
actual payment or deferral date of the Bonus.

3.4 TERMINATION OF PARTICIPATION.

A Participant may elect to terminate his or her active participation in the
Plan at any time by filing a written notice thereof with the Committee. Such
termination of active participation shall become effective as of the
beginning of the next full payroll period following receipt of such election
by the Committee. Amounts credited to the Participant's Account before the
effective date of such termination of active participation shall continue to
be payable, receive investment credits, and otherwise be governed in
accordance with the terms of the Plan as applied to all Participants. A
participant that has terminated active participation shall not be permitted
to resume his or her active participation in the Plan before the end of the
Plan Year within which he terminated participation.

                                                                              10
<PAGE>

3.5 INELIGIBLE PARTICIPANT.

Notwithstanding any other provisions of this Plan to the contrary, if the
Committee determines that any Participant may not qualify as a "management or
highly compensated employee" within the meaning of the Employee Retirement
Income Security Act of 1974, as amended ("ERISA"), or regulations thereunder,
the Committee may determine, in its sole discretion, that such Participant shall
cease to be eligible to participate in this Plan. Upon such determination, any
future Participant's contributions to the Plan will cease; however, the
Participant's existing account balance will be maintained in the same manner as
other Plan Participants.

                                                                              11
<PAGE>

                                   ARTICLE IV

DEFERRAL OF COMPENSATION; COMPANY CREDITS

4.1 AMOUNT OF DEFERRAL.

With respect to each Plan Year, a Participant may elect to defer a specified
percentage of his or her Compensation up to the percentage of compensation
defined and the terms described in Exhibit B attached hereto.

4.2 CREDITING DEFERRED COMPENSATION.

The amount of Compensation that a Participant elects to defer under the Plan
shall be credited by the Employer to the Participant's Deferral Account
periodically, the frequency of which will be determined by the Committee. To the
extent that the Employer is required to withhold any taxes or other amounts from
a Participant's deferred Compensation pursuant to any state, federal or local
law, such amounts shall be withheld only from the Participant's compensation
before such amounts are credited.

4.3 COMPANY CONTRIBUTION ACCOUNT.

The Company may, in its sole discretion, credit amounts to an account for a
Participant (the "Company Contribution Account"). The Company shall have no
obligation to treat each Participant the same with respect to such credits and
may credit different amounts or no amounts to different Participants. The
Company Contribution account and its terms are described in Exhibit B attached
hereto.

                                                                              12

<PAGE>

                                    ARTICLE V

BENEFIT ACCOUNTS

5.1 VALUATION OF ACCOUNT.

As of each Valuation Date, a Participant's Account shall consist of the balance
of the Participant's Account as of the immediately preceding Valuation Date,
plus the Participant's Deferred Compensation credited pursuant to Section 4.2
since the immediately preceding Valuation Date, plus investment return credited
as of such Valuation Date pursuant to Section 5.2, minus the aggregate amount of
distributions, if any, made from such Account since the immediately preceding
Valuation Date.

5.2 CREDITING OF INVESTMENT RETURN.

As of each Valuation Date, each Participant's Deferral Account shall be
increased by the amount of investment return earned since the immediately
preceding Valuation Date. Investment return shall be credited at the Investment
Return Rate as of such Valuation Date based on the average balance of the
Participant's Deferral Account, since the immediately preceding Valuation Date,
but after such Accounts have been adjusted for any contributions or
distributions to be credited or deducted for such period. Investment return for
the period prior to the first Valuation Date applicable to a Deferral Account
shall be deemed earned ratably over such period. Until a Participant or his or
her Beneficiary receives his or her entire Account, the unpaid balance thereof
shall earn an investment return as provided in this Section 5.2.

5.3 STATEMENT OF ACCOUNTS.

The Committee shall provide to each Participant, within 30 days after the close
of each calendar quarter, a statement setting forth the balance of such
Participant's Account as of the last day of the preceding calendar quarter and
showing all adjustments made thereto during such calendar quarter.

                                                                              13
<PAGE>

5.4 VESTING OF ACCOUNT.

Except as provided in Sections 10.1 and 10.2, a Participant shall be 100% vested
in his or her Deferral Account at all times. A Participant's interest in his or
her Company Contribution Account shall be 100% vested upon the occurrence of a
Change in Control. Prior to this event, a Participant's interest in his or her
Company Contribution Account shall vest under the following vesting schedule:

                  CREDITED SERVICE          VESTED PERCENTAGE
                  ----------------          -----------------
                       0-1 year                     0%
                       1-2 years                   20%
                       2-3 years                   40%
                       3-4 years                   60%
                       4-5 years                   80%
                  More than 5 years               100%

Any nonvested portion of a Participant's Company Contribution Account shall be
forfeited at termination. Forfeitures under the Plan shall be for the benefit of
the Employer and shall not be credited to other Participants.

5.5 INVESTMENT VEHICLES.

The Company may select investment vehicles owned as general assets by the
Company or as assets of a trust described in Section 10.1 to establish the
Investment Return Rate. The deemed investment vehicles are set forth in Exhibit
C, which the Company may amend from time to time in its sole discretion.

A Participant may request the Company to make deemed investments of the credit
balance of his Deferral Account in one or more of such investment vehicles. A
Participant may change the deemed investment of his Deferral Account or change
the deemed investment of future credits to his Deferral Account and the deemed
investment of his existing Deferral Account balance may differ from the deemed
investment of future amounts credited to the Deferral Account. Such changes
shall be made in accordance with procedures as the Committee may establish from
time to time. Such procedures may regulate the frequency of such changes and the
form of notice required to make such election or changes. The Committee may also
establish a deemed investment which shall apply if the Participant makes no
election.

The effective date of any change shall be the date for which the appropriate
direction to the Company or its designee has been properly received in
accordance with the procedures established by the Committee. The Committee shall
have the right to refuse to honor any Participant direction related to
investments or withdrawals, including transfers among investment options, where
necessary or desirable to assure compliance with applicable law including U.S.
and other securities laws. However, neither the Company nor the Committee
assumes any responsibility for compliance by officers or others with any such
laws, and any failure by the Company or the Committee to delay or dishonor any
such direction shall not be deemed to increase the Company's legal obligations
to the Participant or third parties.

                                                                              14
<PAGE>

                                   ARTICLE VI

PAYMENT OF BENEFITS

6.1 PAYMENT OF DEFERRAL BENEFIT UPON DISABILITY OR RETIREMENT.

Upon the Disability or Retirement of a Participant, the Employer shall pay to
the Participant or his Beneficiary a Deferral Benefit equal to the balance of
his or her vested Account determined pursuant to Article V, less any amounts
previously distributed, based on his or her written election pursuant to Section
6.6 . However, in the event of an involuntary termination as a result of a
Change of Control of the Company, the participant will receive his or her
Deferral Benefit in a Lump Sum payment.

6.2 PAYMENT OF DEFERRAL BENEFIT UPON TERMINATION.

Upon the termination of service of the Participant as an employee of the
Employer and all Selected Affiliates for reasons other than death, Disability,
or Retirement, the Employer shall pay to the Participant a Deferral Benefit in a
lump sum equal to the balance of his or her vested Account determined pursuant
to Article V, less any amounts previously distributed, as soon as
administratively practical.

6.3 PAYMENTS TO BENEFICIARIES UPON PARTICIPANT DEATH.

In the event of the Participant's death prior to his or her receipt his account,
the Employer shall pay to the Participant's beneficiaries a Deferral Benefit in
a lump sum equal to the balance of his or her vested Account determined pursuant
to Article V, less any amounts previously distributed, as soon as
administratively practical.

6.4 HARDSHIP WITHDRAWAL - "HAIRCUT" PROVISIONS.

Notwithstanding any other provision of the Plan, a Participant at any time shall
be entitled to receive, upon written request to the Committee, a partial or full
lump sum distribution on the amount owed to the Participant under the Plan
subject to penalties as set forth below:

     (a)  The lump-sum will be subject to a 10% penalty and forfeited by the
          Participant, and;

     (b)  The Participant will not be eligible to recommence income deferrals
          until the first of the January following a one (1) year period
          commencing on the date of withdrawal, and then only if otherwise
          eligible to participate under the terms of the Plan.

The amount payable under this section of the Plan shall be paid within
forty-five (45) days following receipt of written notice by the Committee.

                                                                              15
<PAGE>

6.5 IN-SERVICE DISTRIBUTION

A participant may elect to receive an in-service distribution of all of his or
her Deferral Account. A Participant's election for an in-service distribution
must be filed in writing with the Committee at the same time his or her Deferral
Election is made. The Participant may elect to receive such Compensation as an
in-service distribution in lump sum only, the amount of which will be the lesser
of the distribution election for that year or the Deferral Account balance
attributable to that year's deferral. Any benefits paid to the Participant as an
in-service distribution shall reduce the amount of Deferral Benefit otherwise
payable to the Participant under the Plan.

6.6 FORM OF PAYMENT.

The Deferral Benefit payable pursuant to Section 6.1 shall be paid in one of the
following forms, as elected by the Participant in his or her Participant
Agreement on file as of one (1) year and one (1) day prior to the date of
termination or death:

     (a)  Annual payments of a substantially level amount which shall amortize
          the Account balance of the payment commencement date over a period of
          five (5) or ten (10) years. Each installment payment shall equal the
          quotient determined by dividing the Participant's remaining account
          balance at the time of the payment by the number of remaining
          installments (including the current installment).

     (b)  A lump sum as soon as administratively practical.

In the event a Participant fails to make a distribution election, his or her
Account Balance shall be distributed as a lump sum distribution as soon as
administratively practical after his or her termination, death or Disability.

6.7 COMMENCEMENT OF PAYMENTS.

Commencement of payments under Section 6.1 of the Plan shall begin within 60
days following receipt of written notice by the Committee of an event which
entitles a Participant (or a Beneficiary) to payments in lump sum under the Plan
or in the January following the event for annual payment.

6.8 SMALL BENEFIT.

In the event the Committee determines that the balance of a Participant's
Account is less than $3,500 at the time of commencement of payments, or the
portion of the balance of the Participant's Account payable to any Beneficiary
is less than $3,500 at the time of commencement of payments, the Committee may
inform the Employer and the Employer, in its discretion, may choose to pay the
benefit in the form of a lump sum payment, notwithstanding any provision of the
Plan or a Participant election to the contrary. Such lump sum payment shall be
equal to the balance of the Participant's Account or the portion thereof payable
to a Beneficiary.

                                                                              16
<PAGE>

                                   ARTICLE VII

BENEFICIARY DESIGNATION

7.1 BENEFICIARY DESIGNATION.

Each Participant shall have the sole right, at any time, to designate any person
or persons as his Beneficiary to whom payment under the Plan shall be made in
the event of his or her death prior to complete distribution to the Participant
of his or her Account. Any Beneficiary designation shall be made in a written
instrument provided by the Committee. All Beneficiary designations must be filed
with the Committee and shall be effective only when received in writing by the
Committee.

7.2 CHANGE OF BENEFICIARY DESIGNATION.

Any Beneficiary designation may be changed by a Participant by the filing of a
new Beneficiary designation, which will cancel all Beneficiary designations
previously filed. The designation of a Beneficiary may be made or changed at any
time without the consent of any person.

7.3 NO DESIGNATION.

If a Participant fails to designate a Beneficiary as provided above, or if all
designated Beneficiaries predecease the Participant, then the Participant's
designated Beneficiary shall be deemed to be the Participant's estate.

7.4 EFFECT OF PAYMENT.

Payment to a Participant's Beneficiary (or, upon the death of a primary
Beneficiary, to the contingent Beneficiary or, if none, to the Participant's
estate) shall completely discharge the Employer's obligations under the Plan.

                                                                              17
<PAGE>

                                  ARTICLE VIII

ADMINISTRATION

8.1 COMMITTEE.

Members of the Committee for the Transkaryotic Therapies, Inc. Deferred
Compensation Plan are listed on Exhibit A. The Committee shall have complete
discretion to i) supervise the administration and operation of the Plan, ii)
adopt rules and procedures governing the Plan from time to time and iii) shall
have authority to give interpretive rulings with respect to the Plan.

8.2 AGENTS.

The Committee may appoint an individual, who may be an employee of the Company,
to be the Committee's agent with respect to the day-to-day administration of the
Plan. In addition, the Committee may, from time to time, employ other agents and
delegate to them such administrative duties as it sees fit, and may from time to
time consult with counsel who may be counsel to the Company.

8.3 BINDING EFFECT OF DECISIONS.

Any decision or action of the Committee with respect to any question arising out
of or in connection with the administration, interpretation and application of
the Plan shall be final and binding upon all persons having any interest in the
Plan.

8.4 INDEMNIFICATION OF COMMITTEE.

The Company shall indemnify and hold harmless the members of the Committee and
their duly appointed agents under Section 8.2 against any and all claims, loss,
damage, expense or liability arising from any action or failure to act with
respect to the Plan, except in the case of gross negligence or willful
misconduct by any such member or agent of the Committee.

                                                                              18
<PAGE>

                                   ARTICLE IX

AMENDMENT AND TERMINATION OF PLAN

9.1 AMENDMENT.

The Company, on behalf of itself and of each Selected Affiliate may at any time
amend, suspend or reinstate any or all of the provisions of the Plan, except
that no such amendment, suspension or reinstatement may adversely affect any
Participant's Account, as it existed as of the day before the effective date of
such amendment, suspension or reinstatement, without such Participant's prior
written consent. The Committee or its delegatee as the case may be, in its sole
discretion, may accelerate the date of payment of a Participant's Account.
Written notice of any amendment or other action with respect to the Plan shall
be given to each Participant.

9.2 TERMINATION.

The Company, on behalf of itself and of each Selected Affiliate, in its sole
discretion, may terminate this Plan at any time and for any reason whatsoever.
Upon termination of the Plan, the Committee shall take those actions necessary
to administer any Accounts existing prior to the effective date of such
termination; provided, however, that a termination of the Plan shall not
adversely affect the value of a Participant's Account, as it existed as of the
day before the effective date of such termination, or the timing or method of
distribution of a Participant's Account, without the Participant's prior written
consent. Except as provided in Section 9.3 , below, a termination of the Plan
shall not give rise to accelerated or automatic vesting of any Participant's
Account.

9.3 CHANGE IN CONTROL

Upon the occurrence of a Change in Control, the Plan shall terminate, all
accounts shall become fully vested and shall be distributed in a lump sum as
soon as practical after the occurrence of a Change in Control.

                                                                              19
<PAGE>

                                    ARTICLE X

MISCELLANEOUS

10.1 FUNDING.

Participants, their Beneficiaries, and their heirs, successors and assigns,
shall have no secured interest or claim in any property or assets of the
Employer. The Employer's obligation under the Plan shall be merely that of an
unsecured promise of the Employer to pay money in the future. Notwithstanding
the foregoing, in the event of a Change in Control, the Corporation shall create
an irrevocable trust, or before such time the Corporation may create an
irrevocable or revocable trust, to hold funds to be used in payment of the
obligations of Employers under the Plan. In the event of a Change in Control or
prior thereto, the Employers shall fund such trust in an amount equal to no less
than the total value of the Participants' Accounts under the Plan as of the
Determination Date immediately preceding the Change in Control, provided that
any funds contained therein shall remain liable for the claims of the respective
Employer's general creditors.

10.2 NONASSIGNABILITY.

No right or interest under the Plan of a Participant or his or her Beneficiary
(or any person claiming through or under any of them) shall be assignable or
transferable in any manner or be subject to alienation, anticipation, sale,
pledge, encumbrance or other legal process or in any manner be liable for or
subject to the debts or liabilities of any such Participant or Beneficiary. If
any Participant or Beneficiary shall attempt to or shall transfer, assign,
alienate, anticipate, sell, pledge or otherwise encumber his or her benefits
hereunder or any part thereof, or if by reason of his or her bankruptcy or other
event happening at any time such benefits would devolve upon anyone else or
would not be enjoyed by him or her, then the Committee, in its discretion, may
terminate his or her interest in any such benefit (including the Deferral
Account) to the extent the Committee considers necessary or advisable to prevent
or limit the effects of such occurrence. Termination shall be effected by filing
a written "termination declaration" with the Transkaryotic Therapies, Inc.
Deferred Compensation Plan Committee of the Company and making reasonable
efforts to deliver a copy to the Participant or Beneficiary whose interest is
adversely affected (the "terminated participant").

As long as the terminated participant is alive, any benefits affected by the
termination shall be retained by the Employer and, in the Committee's sole and
absolute judgment, may be paid to or expended for the benefit of the terminated
participant, his or her spouse, his or her children or any other person or
persons in fact dependent upon him or her in such a manner as the Committee
shall deem proper. Upon the death of the terminated participant, all benefits
withheld from him or her and not paid to others in accordance with the preceding
sentence shall be disposed of according to the provisions of the Plan that would
apply if he or she died prior to the time that all benefits to which he or she
was entitled were paid to him or her.

                                                                              20
<PAGE>

10.3 CAPTIONS.

The captions contained herein are for convenience only and shall not control or
affect the meaning or construction hereof.

10.4 GOVERNING LAW.

The provisions of the Plan shall be construed and interpreted according to the
laws of the State of Massachusetts.

10.5 SUCCESSORS.

The provisions of the Plan shall bind and inure to the benefit of the Company,
its Selected Affiliates, and their respective successors and assigns. The term
successors as used herein shall include any corporate or other business entity
which shall, whether by merger, consolidation, purchase or otherwise, acquire
all or substantially all of the business and assets of the Company or a Selected
Affiliate and successors of any such Company or other business entity.

10.6 RIGHT TO CONTINUED SERVICE.

Nothing contained herein shall be construed to confer upon any Eligible Employee
the right to continue to serve as an Eligible Employee of the Employer or in any
other capacity.

EXECUTED THIS 1st DAY OF OCTOBER, 2000

                                      TRANSKARYOTIC THERAPIES, INC.

                                      BY: /s/ Daniel E. Geffken
                                         --------------------------

                                      TITLE: Vice President, Finance
                                             and Chief Financial Officer

                                                                              21

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