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end
</PDF>Exhibit 4.5

EXECUTION COPY

FIRST SUPPLEMENTAL
INDENTURE

FIRST SUPPLEMENTAL INDENTURE, dated as of June 14,
2007 (this “Supplemental Indenture”), among JANUS CAPITAL GROUP INC. (formerly
known as Stilwell Financial Inc. (“Stilwell”)), a Delaware corporation (the “Company”),
and THE BANK OF NEW YORK TRUST COMPANY, N.A. (as successor to The Chase Manhattan
Bank) (the “Trustee”).

W I T N E S S E T H:

WHEREAS, Stilwell and the Trustee heretofore executed
and delivered an Indenture, dated as of November 6, 2001 (the “Indenture”),
providing for unlimited issuance of senior debt securities;

WHEREAS, on July 2, 2002, Stilwell issued $200,000,000
aggregate principal amount of 7.75% Notes due 2009 (the “2009 Notes”) under the
Indenture and established the terms and provisions of the 2009 Notes pursuant
to the Officers’ Certificate, dated July 2, 2002 (the “2009 Notes Certificate”),
pursuant to Sections 201, 203 and 301 of the Indenture;

WHEREAS, on September 18, 2006, the Company issued
$275,000,000 aggregate principal amount of 5.875% Notes due 2011 (the “2011
Notes” and, together with the 2009 Notes, the “Outstanding Notes”) under the
Indenture and established the terms and provisions of the 2011 Notes pursuant
to the Officers’ Certificate, dated September 18, 2006 (the “2011 Notes
Certificate” and, together with the 2009 Notes, the “Outstanding Notes
Certificates”), pursuant to Sections 201, 203 and 301 of the Indenture;

WHEREAS, on the date hereof, the Company has issued
$750,000,000 aggregate principal amount of senior notes (the “New Notes”)
pursuant to the Indenture and as part of the terms thereof has agreed to a
covenant (the “Interest Rate Adjustment Covenant”) whereby the applicable
interest rate on the New Notes shall be adjusted upon the occurrence of certain
specified changes in the rating of the Company’ debt securities;

WHEREAS, the Company desires to enter into the First
Supplemental Indenture in order to add the same Interest Rate Adjustment Covenant
for the benefit of the Holders of all the Outstanding Notes, and pursuant to
Section 901(2) of the Indenture, the Trustee and the Company are authorized to
enter into a supplemental indenture, without notice to or consent of any
Holders of the Outstanding Notes, for the purpose of adding to the covenants of
the Company solely for the benefit of the Holders of all the Outstanding Notes;

WHEREAS, this First Supplemental Indenture has been
duly authorized by all necessary corporate action on the part of the Company;
and

NOW THEREFORE, in consideration of the foregoing and
for other good and valuable consideration, the receipt of which is hereby
acknowledged, the Company and the Trustee mutually covenant and agree for the
equal and ratable benefit of the Holders of the Outstanding Notes as follows:

The amendment set forth below shall become effective
upon the execution and delivery of this First Supplemental Indenture by the
Company and the Trustee.

ARTICLE I

The following covenant shall be added to, and shall be
deemed a part of, Article Ten of the Indenture:

1008. Interest Rate Adjustment. The interest
rate payable on the Notes shall be subject to adjustments from time to time if
either Moody’s Investors Service, Inc., or any successor thereto (“Moody’s”) or
Standard & Poor’s Ratings Services, a division of McGraw-Hill, Inc., or any
successor thereto (“S&P”) downgrades (or subsequently upgrades) the debt
rating assigned to the Notes, as set forth below.

If the rating from Moody’s of the Notes is decreased
to a rating set forth in the immediately following table, the interest rate on
the Notes shall increase from the interest rate payable on the Notes on the
date of their issuance (the “Original Interest Rate”) by the percentage set
forth opposite that rating:

	
  Rating

  	
   

  	
  Percentage

  	
   

  
	
  Ba1

  	
   

  	
  0.25

  	
  %

  
	
  Ba2

  	
   

  	
  0.50

  	
  %

  
	
  Ba3

  	
   

  	
  0.75

  	
  %

  
	
  B1 or below

  	
   

  	
  1.00

  	
  %

  

 

If the rating from S&P of the Notes is decreased
to a rating set forth in the immediately following table, the interest rate on
the Notes shall increase from the Original Interest Rate by the percentage set
forth opposite that rating: 

	
  Rating

  	
   

  	
  Percentage

  	
   

  
	
  BB+

  	
   

  	
  0.25

  	
  %

  
	
  BB

  	
   

  	
  0.50

  	
  %

  
	
  BB-

  	
   

  	
  0.75

  	
  %

  
	
  B+ or below

  	
   

  	
  1.00

  	
  %

  

 

Notwithstanding the foregoing, if at any time the
interest rate on the Notes has been adjusted upward and either Moody’s or
S&P, as the case may be, subsequently increases its rating of the Notes to
any of the threshold ratings set forth in the tables above, the interest rate
on the Notes shall be decreased such that the interest rate for the Notes
equals the Original Interest Rate plus the percentages set forth opposite the
ratings from the tables above in effect immediately following the increase. If
Moody’s 

 2
 

subsequently increases its rating of the Notes to Baa3
or higher and S&P increases its rating to BBB- or higher the interest rate
on the Notes shall be decreased to the Original Interest Rate.

Each adjustment required by any decrease or increase
in a rating set forth above, whether occasioned by the action of Moody’s or
S&P, shall be made independent of any and all other adjustments. In no
event shall (1) the interest rate for the Notes be reduced to below the
Original Interest Rate or (2) the total increase in the interest rate on the
Notes exceed 2.00% above the Original Interest Rate.

If either Moody’s or S&P ceases to provide a
rating of the Notes, any subsequent increase or decrease in the interest rate
of the Notes necessitated by a reduction or increase in the rating by the
agency continuing to provide the rating shall be twice the percentage set forth
in the applicable table above. No adjustments in the interest rate of the Notes
shall be made solely as a result of either Moody’s or S&P ceasing to
provide a rating. If both Moody’s and S&P cease to provide a rating of the
Notes, the interest rate on the Notes shall increase to, or remain at, as the
case may be, 2.00% above the Original Interest Rate.

Any interest rate increase or decrease described above
shall take effect from the first day of the interest period during which a
rating change requires an adjustment in the interest rate.

The interest rate on the Notes shall permanently cease
to be subject to any adjustment described above (notwithstanding any subsequent
decrease in the ratings by either or both rating agencies) and, if applicable,
shall be decreased to the Original Interest Rate, if the Notes become rated
Baa2 and BBB or higher by Moody’s and S&P, respectively (or one of these
ratings if only rated by one rating agency), with a stable or positive outlook
by each of the rating agencies.

ARTICLE II

Miscellaneous

Section 2.1.            Effect
of First Supplemental Indenture. Upon the execution and delivery of this
First Supplemental Indenture by the Company and the Trustee, the Indenture and
each of the Outstanding Notes Certificates shall be supplemented in accordance
herewith, and this First Supplemental Indenture shall form a part of the
Indenture for all purposes, and every Holder of 2009 Notes or 2011 Notes, as
the case may be, heretofore or hereafter authenticated and delivered under the
Indenture shall be bound thereby.

Section 2.2.            Indenture
and Outstanding Notes Certificates Remain in Full Force and Effect. Except
as supplemented hereby, all provisions in the Indenture and each of the Outstanding
Notes Certificates shall remain in full force and effect.

Section 2.3.            Indenture,
Outstanding Notes Certificates and First Supplemental Indenture Construed
Together. This First Supplemental Indenture is an 

 3
 

indenture supplemental to and in implementation of the
Indenture, and the Indenture, the applicable Outstanding Notes Certificate and
this First Supplemental Indenture shall henceforth be read and construed
together.

Section 2.4.            Confirmation
and Preservation of Indenture. The Indenture and the applicable Outstanding
Notes Certificates, as supplemented by this First Supplemental Indenture, shall
in all respects remain in full force and effect.

Section 2.5.            Conflict
with Trust Indenture Act. If any provision of this First Supplemental
Indenture limits, qualifies or conflicts with any provision of the TIA that is
required under the TIA to be part of and govern any provision of this First
Supplemental Indenture, the provision of the TIA shall control.  If any provision of this First Supplemental
Indenture modifies or excludes any provision of the TIA that may be so modified
or excluded, the provision of the TIA shall be deemed to apply to the Indenture
as so modified or to be excluded by this First Supplemental Indenture, as the
case may be.

Section 2.6.            Severability.
If any court of competent jurisdiction shall determine that any provision in
this First Supplemental Indenture shall be invalid, illegal or unenforceable,
the validity, legality and enforceability of the remaining provisions shall not
in any way be affected or impaired thereby.

Section 2.7.            Terms
Defined in the Indenture. All capitalized terms not otherwise defined
herein shall have the meanings ascribed to them in the Indenture.

Section 2.8.            Headings.
The Article and Section headings of this First Supplemental Indenture have been
inserted for convenience of reference only, are not to be considered a part of
this First Supplemental Indenture and shall in no way modify or restrict any of
the terms or provisions hereof.

Section 2.9.            Benefits
of First Supplemental Indenture. 
Nothing in this First Supplemental Indenture, the Outstanding Notes
Certificates, the 2009 Notes or the 2011 Notes, express or implied, shall give
to any Person, other than the parties hereto and thereto and their successors
hereunder and thereunder and the Holders of the 2009 Notes and the 2011 Notes,
any benefit of any legal or equitable right, remedy or claim under the
Indenture, the applicable Outstanding Notes Certificate, this First Supplemental
Indenture or the 2009 Notes or 2011 Notes, as the case may be.

Section 2.10.          Successors.
All agreements of the Company in this First Supplemental Indenture shall bind
its successors.  All agreements of the
Trustee in this First Supplemental Indenture shall bind its successors.

Section 2.11.          Trustee
Not Responsible for Recitals. The recitals contained herein shall be taken
as the statements of the Company and the Trustee assumes no responsibility for
their correctness.

Section 2.12.          Certain
Duties and Responsibilities of the Trustee. 
In entering into this First Supplemental Indenture, the Trustee shall be
entitled to the benefit 

 4
 

of every provision of the Indenture, each of the Outstanding
Notes Certificates, the 2009 Notes and the 2011 Notes relating to the conduct
or affecting the liability or affording protection to the Trustee, whether or
not elsewhere herein so provided.

Section 2.13.          Governing
Law. This First Supplemental Indenture shall be governed by, and construed
in accordance with, the laws of the State of New York but without giving effect
to applicable principles of conflicts of law to the extent that the application
of the laws of another jurisdiction would be required thereby.

Section 2.14.          Counterpart
Originals. The parties may sign any number of copies of this First
Supplemental Indenture.  Each signed copy
shall be an original, but all of them together represent the same agreement.

 5
 

IN WITNESS WHEREOF, the parties hereto have caused
this First Supplemental Indenture to be duly executed as of the date first
above written.

	
  

  	
  JANUS CAPITAL GROUP INC.

  
	
   

  	
   

  	
   

  
	
   

  	
  By:

  	
  /s/ Gregory A. Frost

  
	
   

  	
  Name:

  	
  Gregory A. Frost

  
	
   

  	
  Title:

  	
  Senior Vice President and Chief Financial Officer

  
	
   

  	
   

  	
   

  
	
   

  	
   

  	
   

  
	
   

  	
  THE BANK OF NEW YORK TRUST COMPANY, N.A. as Trustee

  
	
   

  	
   

  
	
   

  	
  By:

  	
  /s/ Lawrence M. Kusch

  
	
   

  	
  Name:

  	
  Lawrence M. Kusch

  
	
   

  	
  Title:

  	
  AVP

  

 

 6

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