龍蝨科

龍蝨科（学名：Dytiscidae），亦作龍虱科，為水甲蟲（英语：water beetle）的一種，隸屬於鞘翅目之下的肉食亞目。生活在田野、水溝、小溪等水體中，是一種掠食性水生昆蟲。其學名源於希臘文δυτικός（dytikos），意思就是「肉食性的潛水甲蟲」。本科物種幾乎見於全世界所有淡水棲境，但亦有部分於落葉堆中棲息。成蟲體長一般在1和2.5 cm（0.4—1.0英寸）之間，儘管物種間的差異會比較大。當中見於歐洲的Dytiscus latissimus（英语：Dytiscus latissimus）及巴西的Megadytes ducalis（英语：Megadytes ducalis）有最大的體型，可分別長至4.5 cm（1.8英寸）到4.75 cm（1.9英寸）。與之相反，擁有最小體型的物種應當為棲息於澳大利亞地下水中的Limbodessus（英语：Limbodessus） atypicali，只有約0.9 mm（0.035英寸）長。 大多數物種的顏色都在深棕色、黑色或深橄榄绿，某些亚科的物種會在其邊沿帶有金黃色的條狀斑紋。 本科物種的成蟲，特别是Cybister屬的物種，均為可食用昆蟲。在美國内华达州洪堡乾湖（英语：Humboldt Sink）的一個山洞内，還在當地史前人類遺下的粪化石内發現有Cybister explanatus（英语：Cybister explanatus）被食用的殘骸。在墨西哥，他們在吃C. explanatus時會以墨西哥夹饼夾着來吃。在日本多個地區（例如：長野縣）有以日本龍蝨（Cybister japonicus）為食的習慣；在中华人民共和国广东省，除了日本龍蝨以外，亦會食用C. bengalensis、C. guerini、C. limbatus、C. sugillatus、C. tripunctatus等物種。D. marginalis（英语：Great diving beetle）很可能也包括在內，但這個物種的肉很少。也有報告指在台灣、泰国及新幾內亞亦有食用本科物種的習慣。 大但在陸地上很慢，成年後也不是特別兇猛，它們也被許多中型鳥類、哺乳動物和其他大型捕食者津津有味地吃掉。幼蟲通常比較安全，由於它們的保護色和通過水柱逃跑的能力；它們可能相當難抓，可能成為小池塘的頂級掠食者。 本科包括下列各屬： Acilius Leach, 1817 Aethionectes Sharp, 1882 Africodytes Biström, 1988 Africophilus Guignot, 1948 Agabetes Crotch, 1873 Agabinus Crotch, 1873 豆龍蝨屬 Agabus Leach, 1817，端毛龙虱属 Agametrus Sharp, 1881 Agaporomorphus Zimmermann, 1921 Aglymbus Sharp, 1880 Agnoshydrus Biström, Nilsson & Wewalka, 1997 Allodessus Guignot, 1953 彩斑龍蝨屬 Allopachria Zimmermann, 1924 Amarodytes Régimbart, 1900 Amurodytes Fery & Petrov, 2013 Andex Sharp, 1882 Andonectes Guéorguiev, 1971 Anginopachria Wewalka, Balke & Hendrich, 2001 Anodocheilus Babington, 1841 Antiporus Sharp, 1882 Australphilus Watts, 1978 Austrelatus Shaverdo, Hájek, Hendrich, Surbakti, Panjaitan & Balke, 2023 Austrodytes Watts, 1978 Barrethydrus Barretthydrus Lea, 1927 Barretthydrus geminatus Lea, 1927 Barretthydrus stepheni Watts, 1978 Barretthydrus tibialis Lea, 1927 Batrachomatus Clark, 1863 Belladessus K.B.Miller & Short, 2015 Bidessini Sharp, 1880 Bidessodes Régimbart, 1895 Bidessonotus Régimbart, 1895 Bidessus Sharp, 1882 Boreonectes (Fery & Ribera, 2018) Fery & Ribera, 2018 Boreonectes Angus, 2010 Borneodessus Balke, Hendrich, Mazzoldi & Biström, 2002 Brachyvatus Zimmermann, 1919 Brancuporus Hendrich, Toussaint & Balke, 2014 Bunites Spangler, 1972 Calicovatellus K.B.Miller & Lubkin, 2001 Canthyporus Zimmermann, 1919 Capelatus Turner & Bilton, 2015 Caperhantus Balke, Hájek & Hendrich, 2017 Carabdytes Balke, Hendrich & Wewalka, 1992 Carabhudrus Carabhydrus Watts, 1978 Carrhydrus Celina Aubé, 1837 Chostonectes Sharp, 1880 Clarkhydrus Fery & Ribera, 2018 Clemnius Villastrigo, Ribera, Manuel, Millán & Fery, 2017 Clypeodytes Régimbart, 1894 Coelambus Thomson, 1860 Coelambus cribrarius (Scudder, 1900) Coelambus derelictus (Scudder, 1900) Coelambus disjectus (Scudder, 1900) Coelambus dzieduszyckii (Lomnicki, 1894) Coelambus ganglbaueri (Lomnicki, 1894) Coelambus infernalis (Scudder, 1900) Coelambus latefasciatus (Lomnicki, 1894) Coelambus miocenus (Wickham, 1912) Coelambus niedzwiedzkii (Lomnicki, 1894) Coelambus ozokeriticus (Lomnicki, 1894) Coelambus picipoides (Lomnicki, 1894) Coelhydrus Sharp, 1882 Colymbetes Clairville, 1806 Colymbetes Schellenberg, 1806 Comaldessus Spangler & Barr, 1995 Copelatus Erichson, 1832 Copelatus abonnenci Guignot, 1939 Copelatus acamas Guignot, 1955 Copelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017 Copelatus advena Sharp, 1882 Copelatus aemulus Bilardo & Rocchi, 1995 Copelatus aequatorius Régimbart, 1899 Copelatus aethiopicus Régimbart, 1906 Copelatus agaroensis Balke, 2008 Copelatus agrias Guignot, 1954 Copelatus alternatus Sharp, 1882 Copelatus amaroides Guignot, 1952 Copelatus amatolensis Omer-Cooper, 1965 Copelatus amazonicus Régimbart, 1889 Copelatus ambiguus Bertrand & Legros, 1975 Copelatus amphibius Ranarilalatiana & Bergsten, 2019 Copelatus anastomosans Guignot, 1952 Copelatus andobonicus Guignot, 1960 Copelatus andreinii Régimbart, 1905 Copelatus andrewesi J.Balfour-Browne, 1939 Copelatus angolensis Peschet, 1924 Copelatus angouei Bilardo & Rocchi, 2011 Copelatus angustus Gschwendtner, 1932 Copelatus ankaratra Ranarilalatiana & Bergsten, 2019 Copelatus annobomensis J.Balfour-Browne, 1939 Copelatus anthracinus Régimbart, 1895 Copelatus antoniorum Hájek & Brancucci, 2011 Copelatus aphroditae Balke, 2003 Copelatus apicinotatus Bilardo & Rocchi, 2018 Copelatus assalyi Bilardo & Rocchi, 2010 Copelatus assimilis Régimbart, 1895 Copelatus ateles Guignot, 1955 Copelatus atrosulcatus Régimbart, 1906 Copelatus babyrousa Hájek, Hendrich & Balke, 2018 Copelatus bacchusi Wewalka, 1981 Copelatus bacillifer Guignot, 1955 Copelatus baculiformis Guignot, 1955 Copelatus badeni Sharp, 1882 Copelatus bakewelli J.Balfour-Browne, 1939 Copelatus baoulicus Bilardo & Pederzani, 1978 Copelatus bapounouensis Bilardo & Rocchi, 2019 Copelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021 Copelatus barbouri Young, 1942 Copelatus basifasciatus Régimbart, 1895 Copelatus basilewskyi Bilardo & Pederzani, 1979 Copelatus bechynei Guignot, 1953 Copelatus befasicus Guignot, 1956 Copelatus bengalensis Guignot, 1955 Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996 Copelatus betampona Ranarilalatiana & Bergsten, 2019 Copelatus bezdeki Sheth, Ghate & Hájek, 2018 Copelatus bibulus Guignot, 1948 Copelatus bicolor (Fabricius, 1792) Copelatus biformis Sharp, 1882 Copelatus bimaculatus (Resende & Vanin, 1991) Copelatus binaghii Bilardo & Pederzani, 1978 Copelatus biolleyi Guignot, 1952 Copelatus biroi Guignot, 1956 Copelatus birougouensis Bilardo & Rocchi, 2018 Copelatus biseriatus J.Balfour-Browne, 1939 Copelatus biswasi Mukherjee & Sengupta, 1986 Copelatus blancasi Guignot, 1958 Copelatus blatchleyi Young, 1953 Copelatus bolivianus Guignot, 1957 Copelatus bombycinus Guignot, 1952 Copelatus bonvouloiri Sharp, 1882 Copelatus bottegoi Régimbart, 1895 Copelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021 Copelatus boukali Hendrich & Balke, 1998 Copelatus brancuccii Rocchi, 1979 Copelatus brasiliensis Zimmermann, 1921 Copelatus brendelli Hájek, Hendrich & Balke, 2018 Copelatus brevicornis (Sharp, 1882) Copelatus brevistrigatus Guignot, 1959 Copelatus brivioi Rocchi, 1976 Copelatus bromeliarum (Scott, 1912) Copelatus brullei Aubé, 1838 Copelatus brunneus J.Balfour-Browne, 1939 Copelatus buqueti Aubé, 1838 Copelatus burgeoni Gschwendtner, 1930 Copelatus caelatipennis Aubé, 1838 Copelatus caelatipennis Chevrolat, 1863 Copelatus caelatus Guignot, 1952 Copelatus caffer J.Balfour-Browne, 1939 Copelatus calaquei Bilardo & Rocchi, 2008 Copelatus camerunensis Guignot, 1941 Copelatus capensis Sharp, 1882 Copelatus carayoni Legros, 1949 Copelatus carinatus Sharp, 1882 Copelatus celinoides Guignot, 1952 Copelatus cessaima Caetano, Bená & Vanin, 2013 Copelatus chevrolati Aubé, 1838 Copelatus chibcha Guignot, 1952 Copelatus chinensis Regimbart, 1899 Copelatus chipiriricus Guignot, 1957 Copelatus chloroticus Régimbart, 1899 Copelatus cinnamomeus Régimbart, 1895 Copelatus clarki Sharp, 1882 Copelatus collarti Gschwendtner, 1932 Copelatus compertus Guignot, 1956 Copelatus concii Bilardo, 1982 Copelatus concolor Sharp, 1882 Copelatus concolorans J.Balfour-Browne, 1939 Copelatus confinis Bilardo & Rocchi, 1999 Copelatus congo Gschwendtner, 1938 Copelatus consimilis Bilardo & Rocchi, 2002 Copelatus consors Sharp, 1882 Copelatus cooperae Bilardo & Pederzani, 1972 Copelatus cordovai Megna & Epler, 2012 Copelatus cordylinoides J.Balfour-Browne, 1938 Copelatus coxalis Sharp, 1882 Copelatus crassus Régimbart, 1895 Copelatus cryptarchoides Régimbart, 1899 Copelatus cubaensis Schaeffer, 1908 Copelatus curtistriatus Bilardo & Rocchi, 1995 Copelatus curvispinis Bilardo & Rocchi, 2008 Copelatus cynthiae Bilardo & Rocchi, 2008 Copelatus daemeli Sharp, 1882 Copelatus danyi Megna & Epler, 2012 Copelatus darlingtoni Young, 1942 Copelatus davidi Wewalka, 2017 Copelatus debilis Sharp, 1882 Copelatus deccanensis Sheth, Ghate & Hájek, 2018 Copelatus decellei Bilardo, 1982 Copelatus decemsulcatus Régimbart, 1895 Copelatus deceptor Bilardo & Rocchi, 1995 Copelatus dentatipenis Jiang, Hajek & Jia, 2022 Copelatus depressus Sharp, 1882 Copelatus descarpentriesi Bertrand & Legros, 1975 Copelatus diffisus Guignot, 1939 Copelatus dimorphus Sharp, 1882 Copelatus distinctus Aubé, 1838 Copelatus distinguendus Régimbart, 1903 Copelatus diversistriatus Jiang, Hajek & Jia, 2022 Copelatus djiboutensis Wewalka & Jäch, 2017 Copelatus dolosus Guignot, 1956 Copelatus doriae Sharp, 1882 Copelatus doudouensis Bilardo & Rocchi, 2015 Copelatus duodecimstriatus Aubé, 1838 Copelatus duponti Aubé, 1838 Copelatus efoutensis Bilardo & Rocchi, 1995 Copelatus ejactus Omer-Cooper, 1965 Copelatus ellai Bilardo & Rocchi, 1995 Copelatus elutus Guignot, 1958 Copelatus enganensis Guignot, 1940 Copelatus epactus Guignot, 1948 Copelatus erichsonii Guérin-Méneville, 1847 Copelatus espinhasso Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024 Copelatus esteriensis Bilardo & Pederzani, 1978 Copelatus eucritus Guignot, 1952 Copelatus evanidus Bilardo & Rocchi, 1995 Copelatus exaratus Sharp, 1882 Copelatus externus Kirsch, 1873 Copelatus falciformis Pederzani & Schizzerotto, 2017 Copelatus fasciatus Bilardo & Rocchi, 1995 Copelatus fastidiosus Guignot, 1959 Copelatus feae Régimbart, 1888 Copelatus felicis Hajek, Jiang & Jia, 2022 Copelatus fernandensis Guignot, 1952 Copelatus ferruginicollis Régimbart, 1895 Copelatus ferus Guignot, 1953 Copelatus festae Régimbart, 1899 Copelatus fidschiensis Zimmermann, 1928 Copelatus fijiensis Guignot, 1955 Copelatus filiformis Sharp, 1882 Copelatus fizpaci Bilardo & Rocchi, 2013 Copelatus flavicans Guignot, 1952 Copelatus flavidus Régimbart, 1895 Copelatus florae Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024 Copelatus fluviaticus Guignot, 1957 Copelatus fontanus J.Balfour-Browne, 1950 Copelatus fossilis Říha, 1974 Copelatus fractistriatus Bilardo & Rocchi, 1995 Copelatus freudei Guignot, 1955 Copelatus fryi J.Balfour-Browne, 1939 Copelatus fulviceps J.Balfour-Browne, 1938 Copelatus fuscipennis Sharp, 1882 Copelatus fuscomaculatus Guignot, 1952 Copelatus gabonicus Bilardo & Pederzani, 1978 Copelatus galapagoensis G.R.Waterhouse, 1845 Copelatus garambanus Guignot, 1955 Copelatus gardineri Scott, 1912 Copelatus geayi Régimbart, 1904 Copelatus geniculatus Sharp, 1882 Copelatus gentilis Sharp, 1882 Copelatus gestroi (Sharp, 1882) Copelatus gigas Bilardo & Rocchi, 2019 Copelatus glyphicus (Say, 1823) Copelatus griffinii Régimbart, 1899 Copelatus gschwendtneri Guignot, 1939 Copelatus guadelupensis Legros, 1948 Copelatus guerini Aubé, 1838 Copelatus guineensis Guignot, 1955 Copelatus haemorrhoidalis Régimbart, 1883 Copelatus hararensis Guignot, 1952 Copelatus hardenbergi J.Balfour-Browne, 1950 Copelatus hebeter Guignot, 1953 Copelatus heterogynus Régimbart, 1899 Copelatus hydroporoides (Murray, 1859) Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000 Copelatus iguelaensis Bilardo & Rocchi, 2002 Copelatus ilybioides Régimbart, 1895 Copelatus imitator Bilardo & Rocchi, 2002 Copelatus inaequalis Sharp, 1882 Copelatus incognitus Sharp, 1882 Copelatus indicus Sharp, 1882 Copelatus inopinatus Bilardo & Rocchi, 1995 Copelatus inornatus Sharp, 1882 Copelatus insidiosus Bilardo & Rocchi, 1995 Copelatus insolitus Chevrolat, 1863 Copelatus instabilis Régimbart, 1897 Copelatus instriatus Zimmermann, 1921 Copelatus insuetus Guignot, 1941 Copelatus insulanus Guignot, 1939 Copelatus integer Sharp, 1882 Copelatus internus Régimbart, 1904 Copelatus interstriatus J.Balfour-Browne, 1939 Copelatus inuber Guignot, 1952 Copelatus ipiformis Régimbart, 1895 Copelatus irinus Régimbart, 1899 Copelatus irregularis W.J.MacLeay, 1871 Copelatus ischius Guignot, 1956 Copelatus jactator Guignot, 1955 Copelatus jaechi Bilardo & Rocchi, 2016 Copelatus jamaicensis Young, 1942 Copelatus japonicus Sharp, 1884 Copelatus jarrigei Legros, 1954 Copelatus javanus Régimbart, 1883 Copelatus jocosus Guignot, 1955 Copelatus johannis Guignot, 1954 Copelatus kalaharii Gschwendtner, 1935 Copelatus kammuriensis Tamu & Tsukamoto, 1955 Copelatus kaszabi Guignot, 1956 Copelatus kely Ranarilalatiana & Bergsten, 2019 Copelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021 Copelatus kilimandjarensis Bilardo & Pederzani, 1972 Copelatus kindianus Guignot, 1954 Copelatus kongouensis Bilardo & Rocchi, 1999 Copelatus koreanus Mori, 1932 Copelatus laccophilinus Sharp, 1882 Copelatus laeticulus Sharp, 1882 Copelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021 Copelatus lamottei Legros, 1954 Copelatus lanzai Bilardo & Rocchi, 1995 Copelatus laraensis Bilardo & Rocchi, 1995 Copelatus lasckonyi Bilardo & Rocchi, 1995 Copelatus latens Guignot, 1952 Copelatus laticollis Régimbart, 1899 Copelatus latifasciatus Bilardo & Rocchi, 1999 Copelatus latipes Sharp, 1882 Copelatus latus J.Balfour-Browne, 1950 Copelatus leonardii Bilardo & Rocchi, 1999 Copelatus leonensis Legros, 1954 Copelatus lepersonneae Gschwendtner, 1943 Copelatus lignosus Guignot, 1955 Copelatus lineatipennis Guignot, 1955 Copelatus lineatus (Guérin-Méneville, 1838) Copelatus longicornis Sharp, 1882 Copelatus lootensi Guignot, 1955 Copelatus louayensis Bilardo & Rocchi, 2004 Copelatus luctuosus Guignot, 1939 Copelatus luridescens Régimbart, 1889 Copelatus luteocinctus Guignot, 1955 Copelatus luteomaculatus Guignot, 1956 Copelatus luzonicus Guignot, 1952 Copelatus macellus Guignot, 1950 Copelatus madoni Guignot, 1952 Copelatus mahajanga Pederzani & Hájek, 2005 Copelatus mahleri Holmen & Vazirani, 1990 Copelatus makokouensis Bilardo & Rocchi, 1995 Copelatus malaisei Guignot, 1954 Copelatus malavergnorum Manuel & Ramahandrison, 2022 Copelatus mancus Sharp, 1887 Copelatus marginatus Sharp, 1882 Copelatus marginipennis (Laporte, 1835) Copelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019 Copelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018 Copelatus masculinus Régimbart, 1899 Copelatus massaicus Guignot, 1941 Copelatus mathani Guignot, 1952 Copelatus maushomi Sheth, Ghate & Hájek, 2018 Copelatus mbokoensis Bilardo & Rocchi, 2006 Copelatus melanogrammus Régimbart, 1883 Copelatus mendax Bilardo & Rocchi, 2011 Copelatus miaowangi Hajek, Jiang & Jia, 2022 Copelatus minimus Bilardo & Rocchi, 1995 Copelatus minor Bilardo & Pederzani, 1978 Copelatus minutissimus J.Balfour-Browne, 1939 Copelatus mocquerysi Régimbart, 1895 Copelatus mohelicus Guignot, 1959 Copelatus monticola Guignot, 1951 Copelatus montivagus Young, 1942 Copelatus mopanshanensis Jiang, Zhao & Hajek, 2022 Copelatus mulangensis Bameul, 2003 Copelatus mundus Sharp, 1882 Copelatus mutabilis Bilardo & Rocchi, 1999 Copelatus mvoungensis Bilardo & Rocchi, 2004 Copelatus mysorensis Vazirani, 1970 Copelatus nakamurai Guéorguiev, 1970 Copelatus nangaensis Guignot, 1952 Copelatus nanlingensis Hajek, Jiang & Jia, 2022 Copelatus nauclerus Guignot, 1939 Copelatus neavei J.Balfour-Browne, 1950 Copelatus neelumae Vazirani, 1973 Copelatus neogestroi Balke, 2008 Copelatus neoguineensis Zimmermann, 1919 Copelatus nguemai Bilardo & Rocchi, 2011 Copelatus nigricans Sharp, 1882 Copelatus nigricollis Paulino de Oliveira, 1880 Copelatus nigrolineatus Sharp, 1882 Copelatus nigropennis Zimmermann, 1927 Copelatus nigrostriatus Régimbart, 1895 Copelatus nilssoni Wewalka, 1982 Copelatus nimbaensis Legros, 1958 Copelatus nitens Bilardo & Rocchi, 1999 Copelatus nitidus Sharp, 1882 Copelatus nodieri Régimbart, 1895 Copelatus normalis Erichson, 1847 Copelatus notius Omer-Cooper, 1965 Copelatus nzei Bilardo & Rocchi, 1999 Copelatus oblitus Sharp, 1882 Copelatus obscurus Sharp, 1882 Copelatus occultus Bilardo & Rocchi, 1995 Copelatus ogasawarensis Kamiya, 1932 Copelatus onorei Pederzani & Rocchi, 1982 Copelatus ornatipennis Zimmermann, 1926 Copelatus owas Régimbart, 1895 Copelatus pachys Guignot, 1955 Copelatus pallidus Régimbart, 1895 Copelatus paludorum Guignot, 1952 Copelatus pandanorum Scott, 1912 Copelatus panguana Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024 Copelatus pantosi Guignot, 1958 Copelatus papuensis J.Balfour-Browne, 1939 Copelatus parabaptus Guignot, 1955 Copelatus parallelipipedus Régimbart, 1895 Copelatus parallelus Zimmermann, 1920 Copelatus pardii Rocchi, 1990 Copelatus parisii Guignot, 1959 Copelatus parumstriatus Gschwendtner, 1934 Copelatus paryphes Guignot, 1955 Copelatus pederzanii Bilardo & Rocchi, 1995 Copelatus pereirai Guignot, 1955 Copelatus peridinus Guignot, 1955 Copelatus pinnifer Guignot, 1952 Copelatus piriensis Omer-Cooper, 1965 Copelatus platynotus Régimbart, 1895 Copelatus polystrigus Sharp, 1882 Copelatus ponomarenkoi Říha, 1974 Copelatus portior Guignot, 1956 Copelatus posticatus (Fabricius, 1801) Copelatus poungai Bilardo & Rocchi, 2006 Copelatus predaveterus Miller, 2003 Copelatus prolixus Sharp, 1882 Copelatus prolongatus Sharp, 1882 Copelatus propino Guignot, 1960 Copelatus propinquus Régimbart, 1895 Copelatus proximus Sharp, 1882 Copelatus pseudostriatus Ranarilalatiana & Bergsten, 2019 Copelatus pulchellus (Klug, 1834) Copelatus pulicarius Régimbart, 1895 Copelatus pumilus Régimbart, 1895 Copelatus punctatus Bilardo & Rocchi, 1995 Copelatus punctulatus Aubé, 1838 Copelatus puzhelongi Jiang, Hajek & Jia, 2022 Copelatus quadrisignatus Régimbart, 1877 Copelatus quadristriatus Pederzani & Schizzerotto, 2017 Copelatus racenisi Guignot, 1951 Copelatus ragazzii Régimbart, 1887 Copelatus regimbarti Branden, 1885 Copelatus restrictus Sharp, 1882 Copelatus rimosus Guignot, 1952 Copelatus rivalis Guignot, 1952 Copelatus rocchii Bilardo, 1982 Copelatus romani Zimmermann, 1924 Copelatus rosulae Hajek, Jiang & Jia, 2022 Copelatus royi Legros, 1958 Copelatus rubiginosus Guignot, 1954 Copelatus ruficapillus Régimbart, 1895 Copelatus ruteri Legros, 1954 Copelatus saegeri Guignot, 1955 Copelatus safiotra Ranarilalatiana & Bergsten, 2019 Copelatus sahlbergi J.Balfour-Browne, 1939 Copelatus sallaei Sharp, 1882 Copelatus sanfilippoi Bilardo & Pederzani, 1972 Copelatus saverii Pederzani & Schizzerotto, 2017 Copelatus scalptus Guignot, 1954 Copelatus scaphites Guignot, 1955 Copelatus schaefferi Young, 1942 Copelatus schereri Wewalka, 1981 Copelatus schuhi Hendrich & Balke, 1998 Copelatus scutatus Guignot, 1952 Copelatus scytalotus Guignot, 1956 Copelatus senegalensis Lagar & Hernando, 1991 Copelatus sexstriatus Sharp, 1882 Copelatus sharpi Branden, 1885 Copelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010 Copelatus silvestrii Régimbart, 1903 Copelatus simoni Régimbart, 1889 Copelatus singularis Bilardo & Rocchi, 1995 Copelatus sociennus J.Balfour-Browne, 1952 Copelatus solitarius Sharp, 1882 Copelatus sordidipennis Régimbart, 1895 Copelatus speciosus Régimbart, 1892 Copelatus spoliatus Guignot, 1955 Copelatus stavropolitanus Říha, 1974 Copelatus stillicidicola Bilardo & Rocchi, 1995 Copelatus striatellus Boheman, 1848 Copelatus striaticollis Lucas, 1857 Copelatus striatopterus Aubé, 1838 Copelatus striatulus Aubé, 1838 Copelatus strigipennis (Laporte, 1835) Copelatus strigosulus (Fairmaire, 1878) Copelatus strinatii Guignot, 1958 Copelatus striolatus Peschet, 1917 Copelatus stygis Guignot, 1958 Copelatus subdeficiens Régimbart, 1902 Copelatus subsimilis Guignot, 1958 Copelatus substriatus Kirsch, 1873 Copelatus subterraneus Guéorguiev, 1978 Copelatus sudrei Bameul, 2003 Copelatus sulcatus Sharp, 1882 Copelatus sulcipennis (Laporte, 1835) Copelatus sumbawensis Régimbart, 1899 Copelatus suppar Guignot, 1956 Copelatus supplementaris Régimbart, 1895 Copelatus sylvaticoides Bilardo & Rocchi, 2013 Copelatus sylvaticus Guignot, 1952 Copelatus takakurai Satô, 1985 Copelatus tanaus Guignot, 1953 Copelatus taprobanicus Wewalka & Vazirani, 1985 Copelatus tenebrosis Regimbart, 1880 Copelatus tenebrosus Régimbart, 1880 Copelatus tengchongensis Hajek, Jiang & Jia, 2022 Copelatus teranishii Kamiya, 1938 Copelatus terminalis Sharp, 1882 Copelatus ternatensis Régimbart, 1899 Copelatus terzanii Bilardo & Saverio, 2016 Copelatus thiriouxi Peschet, 1917 Copelatus thrasys Guignot, 1952 Copelatus tibialis Sharp, 1882 Copelatus tinctor Guignot, 1954 Copelatus togoensis Régimbart, 1895 Copelatus tomokunii Satô, 1985 Copelatus tondangoyei Bilardo & Rocchi, 2013 Copelatus tostus J.Balfour-Browne, 1950 Copelatus trifilis Guignot, 1958 Copelatus triglyphus Guignot, 1955 Copelatus trilobatus Régimbart, 1895 Copelatus tschaga Bilardo & Pederzani, 1972 Copelatus tucuchiensis J.Balfour-Browne, 1939 Copelatus tulagicus Guignot, 1942 Copelatus uludanuensis Hendrich & Balke, 1995 Copelatus undecimstriatus Aubé, 1838 Copelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021 Copelatus usagarensis Zimmermann, 1926 Copelatus vagatus Guignot, 1952 Copelatus vagestriatus Zimmermann, 1919 Copelatus vagus Bilardo & Rocchi, 1995 Copelatus validus Sharp, 1882 Copelatus vancli Hájek, Hendrich & Balke, 2018 Copelatus vanninii Bilardo & Rocchi, 1999 Copelatus variegatus Régimbart, 1895 Copelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021 Copelatus venustus Bilardo & Rocchi, 1995 Copelatus vigintistriatus Fairmaire, 1869 Copelatus vigintisulcatus Régimbart, 1895 Copelatus villiersi Guignot, 1950 Copelatus virungaensis Bilardo, 1982 Copelatus vitraci Legros, 1948 Copelatus vivax Guignot, 1953 Copelatus vokoka Ranarilalatiana & Bergsten, 2019 Copelatus wallacei J.Balfour-Browne, 1939 Copelatus waltoni J.Balfour-Browne, 1950 Copelatus wayanadensis Manivannan & Madani, 2011 Copelatus wewalkai Holmen & Vazirani, 1990 Copelatus weymarni J.Balfour-Browne, 1947 Copelatus xanthocephalus Régimbart, 1899 Copelatus xiniudong Jiang, Hajek & Jia, 2022 Copelatus yacumensis Guignot, 1957 Copelatus yaguarete Scheers, 2018 Copelatus yombensis Bilardo & Rocchi, 2015 Copelatus zadiensis Bilardo & Rocchi, 1995 Copelatus zanabato Manuel & Ramahandrison, 2022 Copelatus zanatanensis Ranarilalatiana & Bergsten, 2019 Copelatus zimmermanni Gschwendtner, 1934 Copelatus Sheth, Ghate & Hájek, 2018 Coptotomus Say, 1830 Cretodytes Ponomarenko, 1977 Crinodessus K.B.Miller, 1997 大龍蝨屬 Cybister Curtis, 1827，真龙虱属 Darwinhydrus Sharp, 1882 Deronectes Sharp, 1882 Derovatellus Sharp, 1882 Desmopachria Babington, 1841 Deuteronectes Guignot, 1945 Deuteronectus Dimitshydrus Uéno, 1996 Dyschirioides Dyticus Geoffroy, 1762 Dytiscus Linnaeus, 1758 Electruphilus Balker, Toledo & Hendrich, 2019 Ereboporus K.B.Miller, Gibson & Alarie, 2009 Eretes Laporte, 1833 Etruscodytes Mazza, Cianferoni & Rocchi, 2013 Exocelina Broun, 1886 Fontidessus K.B.Miller & Spangler, 2008 Geodessus Brancucci, 1979 Gibbidessus Watts, 1978 Glareadessus Wewalka & Biström, 1998 Graphoderus Dejean, 1833 Graptodytes Seidlitz, 1887 Haideoporus Young & Longley, 1976 Hamadiana Benetti, Short & Michat, 2019 Hemibidessus Zimmermann, 1921 Heroceras Guignot, 1950 Heterhydrus Fairmaire, 1869 Heterosternuta Strand, 1935 Hintonia Young, 1980 Hoperius Fall, 1927 Hornectes Fery & Ribera, 2018 Hovahydrus Biström, 1982 Huxelhydrus Sharp, 1882 Hydaticus Leach, 1817 Hyderodes Hope, 1838 Hydrocolus Roughley & Larson, 2000 Hydrodessus J.Balfour-Browne, 1953 Hydrodytes K.B.Miller, 2001 短褶龙虱属 Hydroglyphus Motschulsky, 1853 Hydronebrius Jakovlev, 1897 Hydropeplus Sharp, 1882 Hydroporus Clairville, 1806 Hydroporus Schellenberg, 1806 Hydrotrupes Sharp, 1882 Hydrovatus Motschulsky, 1853 水龙虱属 Hygrotus Stephens, 1828 Hyphoporus Sharp, 1882 Hyphovatus Wewalka & Biström, 1994 异爪龙虱属 Hyphydrus Illiger, 1802 Hypodessus Guignot, 1939 Iberonectes Fery & Ribera, 2018 Iberoporus Castro & Delgado, 2001 Ilybiosoma Crotch, 1873 Ilybius Erichson, 1832 Incomptodessus K.B.Miller & García, 2011 Isonotus Houlbert, 1934 Japanolaccophilus Satô, 1972 Kakadudessus Hendrich & Balke, 2009 Kakadudessus tomweiri Hendrich & Balke, 2009 Kuschelydrus Ordish, 1976 Laccodytes Régimbart, 1895 Laccomimus Toledo & Michat, 2015 Lacconectus Motschulsky, 1855 粒龙虱属 Laccophilus Leach, 1815 Laccoporus J.Balfour-Browne, 1939 Laccoporus nigritulus (Gschwendtner, 1936) Laccornellus Roughley & Wolfe, 1987 Laccornis Gozis, 1914 Laccosternus Brancucci, 1983 Lancetes Sharp, 1882 Laodytes Queney, Lemaire & Ferrand, 2020 Larsonectes Fery & Ribera, 2018 Leconectes Fery & Ribera, 2018 Leiodytes Guignot, 1936 Leuronectes Sharp, 1882 Liadyxianus Prokin, Petrov, Wang & Ponomarenko, 2013 Limbodessus Guignot, 1939 Liodessus Guignot, 1939 Lioporeus Guignot, 1950 Liopterus Dejean, 1833 Madaglymbus Shaverdo & Balke, 2008 Matus Aubé, 1836 Megadytes Sharp, 1882 Megaporus Brinck, 1943 Meladema Laporte, 1835 Melanodytes Seidlitz, 1887 Meridiorhantus Balke, Hájek & Hendrich, 2017 Mesodytes Prokin, Petrov, Wang & Ponomarenko, 2013 Metaporus Guignot, 1945 Methles Sharp, 1882 Microdessus Young, 1967 Microdytes J.Balfour-Browne, 1946 Microhydrodytes K.B.Miller, 2002 Miodytiscus Wickham, 1911 Morimotoa Uéno, 1957 Mystonectes Fery & Ribera, 2018 Napodytes Steiner, 1981 Nartus Zaitzev, 1907 孔龙虱属 Nebrioporus Régimbart, 1906 Necterosoma Sharp, 1882 Necterosoma W.J.MacLeay, 1871 Nectoboreus Fery & Ribera, 2018 Nectomimus Fery & Ribera, 2018 Nectoporus Guignot, 1950 Neobidessodes Hendrich & Balke, 2009 Neobidessus Young, 1967 Neoclypeodytes Young, 1967 Neonectes J.Balfour-Browne, 1944 Neonectes Zimmermann, 1932 Neoporus Guignot, 1931 Neoscutopterus J.Balfour-Browne, 1943 彩裳粒龙虱属 Neptosternus Sharp, 1882 Notaticus Zimmermann, 1928 Novadessus K.B.Miller, 2016 Onychohydrus Schaum & White, 1847 Oreodytes Seidlitz, 1887 Pachydrus Sharp, 1882 Pachynectes Régimbart, 1903 Palaeodytes Ponomarenko, 1987 Papuadessus Balke, 2001 Paroster Sharp, 1882 Peschetius Guignot, 1942 Petrodessus K.B.Miller, 2012 Philaccolilus Guignot, 1937 Philaccolus Guignot, 1937 Philodytes J.Balfour-Browne, 1939 Phreatodessus Ordish, 1976 Platambus Thomson, 1859 Platydytes Biström, 1988 Platynectes Gueorguiev, 1972 Platynectes Régimbart, 1879 Porhydrus Guignot, 1945 Primospes Sharp, 1882 Procoelambus Théobald, 1937 Prodaticus Sharp, 1882 缝龙虱属 Pseuduvarus Biström, 1988 Psychopomporus Jean, Telles & K.B.Miller, 2012 Pteroporus Guignot, 1933 Queda Sharp, 1882 Regimbartina Chatanay, 1911 Rhantaticus Sharp, 1880 Rhantus Dejean, 1833 Rhithrodytes Bameul, 1989 Rompindessus Balke, Bergsten & Hendrich, 2017 Sandracottus Sharp, 1882 Sanfilippodytes Franciscolo, 1979 Scarodytes Gozis, 1914 Schistomerus Palmer, 1957 Sekaliporus Watts, 1997 Sharphydrus Omer-Cooper, 1958 Siamoporus Spangler, 1996 Siettitia Abeille de Perrin, 1904 Sinodytes Spangler, 1996 Spanglerodessus K.B.Miller & García, 2011 Spencerhydrus Sharp, 1882 Sternhydrus Brinck, 1945 Sternopriscus Sharp, 1880 Stictonectes Brinck, 1943 Stictotarsus Zimmermann, 1919 Stygoporus Larson & LaBonte, 1994 Suphisellus Zimmermann, 1921 Tassilodytes Fery & Bouzid, 2016 Tepuidessus Spangler, 1981 Thermonectus Dejean, 1833 Thermonectus alfredi Griffini, 1898 Thermonectus basillaris (Harris, 1829) Thermonectus batesi Sharp, 1882 Thermonectus circumscriptus (Latreille, 1809) Thermonectus cuneatus Sharp, 1882 Thermonectus depictus Sharp, 1882 Thermonectus duponti (Aubé, 1838) Thermonectus intermedius Crotch, 1873 Thermonectus laporti (Aubé, 1838) Thermonectus leprieuri J.Balfour-Browne, 1944 Thermonectus margineguttatus (Aubé, 1838) Thermonectus marmoratus (Gray, 1831) Thermonectus nigrofasciatus (Aubé, 1838) Thermonectus nobilis Zimmermann, 1924 Thermonectus sibleyi Goodhue-McWilliams, 1981 Thermonectus simulator Sharp, 1882 Thermonectus succinctus (Aubé, 1838) Thermonectus tremouillesi Michat & Torres, 2016 Thermonectus variegatus (Dejean, 1836) Thermonectus variegatus (Laporte de Castelnau, 1835) Thermonectus zimmermani Goodhue-McWilliams, 1981 Tikoloshanes Omer-Cooper, 1956 Tiporus Watts, 1985 Trichonectes Guignot, 1941 Trogloguignotus Sanfilippo, 1958 Tyndallhydrus Sharp, 1882 Typhlodessus Brancucci, 1985 Uvarus Guignot, 1939 Vatellus Aubé, 1837 Yola Gozis, 1886 Yolina Guignot, 1936 Zaitzevhydrus Fery & Ribera, 2018 Zimpherus K.B.Miller & Wheeler, 2015 G.N. Foster; D.T. Bilton. The Conservation of Predaceous Diving Beetles: Knowns, Unknowns and Anecdotes. D.A. Yee (编). Ecology, Systematics, and the Natural History of Predaceous Diving Beetles (Coleoptera: Dytiscidae). 2014: 437–462. ISBN 978-94-017-9109-0.  Archived copy. [2015-05-19]. （原始内容存档于2015-05-21）.  Miller, Kelly; Bergsten, Johannes. Diving Beetles of the World: Systematics and Biology of the Dytiscidae. Baltimore: Johns Hopkins University Press. 3 October 2016: 20.  De Foliart (2002), Jäch (2003), CSIRO (2004) Dytiscidae Species List. Joel Hallan's Biology Catalog. Texas A&M University. [2012-05-07]. （原始内容存档于2012-10-23） （英语）.  Dytiscidae 龍蝨科. TaiBIF. [2019-03-13]. （原始内容存档于2019-07-28） （中文（繁體））.  Jiří Hájek, Yves Alarie, Cesar J. Benetti, Neusa Hamada, Monika Springer, Lars Hendrich, Adrián Villastrigo, Rodulfo Ospina Torres, Michael S. Basantes and Michael Balke. 2024. Underestimated Diversity and Range Size of Diving Beetles in Tank Bromeliads—Coleoptera of ‘hygrofloric’ lifestyle (Dytiscidae). Zoological Journal of the Linnean Society. 200(3); 720–735. DOI: 10.1093/zoolinnean/zlad093 Mariano C. Michat & Patricia L. M. Torres (2016) Thermonectus tremouillesi sp. nov. (Coleoptera: Dytiscidae: Aciliini): description of the adults and larvae and comparisons with other species of the genus, Journal of Natural History, 50:25-26, 1633-1648, DOI: 10.1080/00222933.2016.1145274 联邦科学与工业研究组织 (CSIRO) (2004): Water for a Healthy Country - Family Dytiscidae （页面存档备份，存于互联网档案馆）. Version of 2004-JUL-02. Retrieved 2008-AUG-04 De Foliart, Gene R. (2002): Chapter 26 - Eastern Asia: China, Japan, and other countries. In: The Human Use of Insects as a Food Resource: A Bibliographic Account in Progress. Jäch, Manfred A. (2003): Fried water beetles Cantonese style. American Entomologist 49(1): 34-37. PDF fulltext Larson, D.J., Alarie, Y., and Roughley, R.E. (2000): Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa. ISBN 978-0-660-17967-4. 维基共享资源上的相关多媒体资源：龍蝨科 維基物種上的相關信息：龍蝨科 YouTube上的Video of the dangerous mandibles of a Dytiscidae larva（英文） 聯邦科學與工業研究組織「家庭龍蝨」 （页面存档备份，存于互联网档案馆）