步行蟲科

步行蟲科（Carabidae，Ground beetles）也稱為步甲科，是鞘翅目（甲蟲）中，世界性分佈的一個科。全世界有超過四萬種，北美洲有二千種，而欧洲有二千七百種。在2015年統計時，是動物界物種數最多的十個分科之一。如虎甲及放屁蟲等都屬於步行蟲科。 步行蟲科的外形及顏色極為多樣化，不過大部份是閃亮的黑色或帶有金屬光澤，而且有脊狀翼蓋（鞘翅）。有些物種的鞘翅無法分開（特別是較大型的物種），因此這類的步行蟲多數無法飛行。其中Mormolyce属因為其酷似小提琴的特殊鞘翅外形，也稱為提琴蟲（英语：violin beetle）（violin beetle）。除了非常原始的flanged bombardier beetle（英语：flanged bombardier beetle）外，所有步行蟲都在腿上有凹槽，其中有梳毛可以清潔昆蟲的觸角。 步行蟲科常見的棲地是在樹幹下、原木下、或是在池塘及河流邊的石頭或砂旁邊。大部份的步行蟲科是肉食性，會主動獵食的无脊椎动物獵物。有些步行蟲科會快速奔跑捕殺獵物，像虎甲蟲（Cicindelinae）可以維持時速9公里的速度，在考慮其相對體長的情形下，他們是陸地上跑得最快的動物。大部份的步行蟲科是夜行動物，虎甲晝夜都會捕食，身上有鮮明的顏色，虎甲有大眼睛，靠視力捕食。步行蟲科中的Promecognathus laevissimus（英语：Promecognathus laevissimus）會捕食黃斑馬陸（英语：Harpaphe haydeniana），由於他們可以對抗氰化氢-這種對大部份步行蟲有毒的物質。星步甲屬的步行蟲物種主要以毒蛾亞科（英语：Lymantriinae）、帶蛾科（英语：Thaumetopoeidae）、燈蛾亞科的毛蟲為食；這些毛蟲擁有螫毛，因此大部分食蟲動物並不會捕捉這些毛蟲。在 1905 年代時，大量來自歐洲的疆星步行蟲（英语：Calosoma sycophanta）被人工引進至紐西蘭進行生物防治，用以消滅當地的舞毒蛾。 本科包括以下属： 步行虫科的属 Abacaecus Allegro & Giachino, 2020 Abacaelostus Straneo, 1952 Abacetodes Straneo, 1939 Abacetus Dejean, 1828 Abacidus LeConte, 1863 Abacillius Straneo, 1949 Abacillodes Straneo, 1988 Abacoleptus Fauvel, 1903 Abacomorphus Chaudoir, 1878 Abacophrastus Will, 2011 Abacops Tschitscherine, 1902 Abaditicus Ball & Hilchie, 1983 Abaris Dejean, 1831 Abax Bonelli, 1810 Abrodiella Bousquet, 2002 Abropus G.R.Waterhouse, 1842 Abroscelis Hope, 1838 Acalathus Semenov, 1889 Acallistus Sharp, 1886 Acanthodes Basilewsky, 1953 Acanthoferonia Moore, 1965 Acanthoodes Basilewsky, 1953 Acanthoscelis Dejean, 1825 Acanthoscelis Latrielle, 1810 Accoella Ueno, 1990 Achaetocephala Habu, 1975 Achaetoprothorax Habu, 1978 Acheroniotes Lohaj & Lakota, 2010 Acinopus Dejean, 1821 Acinopus Latreille, 1829 Acrogenys W.J.MacLeay, 1864 Actenoncus Chaudoir, 1872 Actenonyx White, 1846 Actodus Alluaud, 1915 Acupalpus Dejean, 1829 Acupalpus Latreille, 1829 Acutosternus Lecordier & Girard, 1988 Adelopomorpha Heller, 1916 Adelotopus Hope, 1836 Adischissus Fedorenko, 2015 Adotela Laporte, 1867 Adriaphaenops Noesske, 1928 Adrimus Bates, 1872 Aenigma Newman, 1836 Aenigmatachys Baehr, 2020 Aeolodermus Andrewes, 1929 Aephnidius W.S.MacLeay, 1825 Aepiblemus Belousov & Kabak, 1993 Aepopsis Jeannel, 1922 Aepsera Chaudoir, 1874 Aepus Leach, 1819 Aepus Samouelle, 1819 Aethiopsis Quéinnec & Ollivier, 2021 Aethocarabus Ren & Lu&Guo, 1995 Afranillus Giachino, 2015 Afrodipnus Giachino, 2015 Afrodromius Basilewsky, 1958 Afrogehringia Baehr & Schüle&Lorenz, 2009 Afromizonus Basilewsky, 1947 Afrosyleter Basilewsky, 1959 Afrotachys Basilewsky, 1958 Afrotarus Jeannel, 1949 Afrotrechus Quéinnec & Ollivier, 2021 Afrozaena Jeannel, 1946 Agastillus Reitter, 1892 Agastus Schmidt-Goebel, 1846 Agatoides Motschulsky, 1856 Agelaea Gené, 1839 Agonica Sloane, 1920 Agonidium Jeannel, 1948 Agonidus Casey, 1914 Agonobembix Jeannel, 1948 Agonocheila Chaudoir, 1848 Agonoleptus Casey, 1914 Agonoriascus Basilewsky, 1985 Agonorites Jeannel, 1951 Agonotrechus Jeannel, 1923 Agonum Bonelli, 1810 Agostenus Fischer von Waldheim, 1829 Agostinia Jeannel, 1928 Agra Fabricius, 1801 Agrogenys Akephorus LeConte, 1852 Alachnothorax Liebke, 1929 Alanorites Belousov, 1998 Albanotrechus Casale & V.B.Gueorguiev, 1994 Alkestis Liebke, 1939 Allardina Basilewsky, 1963 Allegrettia Jeannel, 1928 Allendia Noonan, 1974 Allocinopus Broun, 1903 Allocota Motschulsky, 1860 Allolicinus Semenov-Tian-Shanskii, 1927 Allophanes Andrewes, 1939 Allophanopsis Louwerens, 1952 Allorhopalus Nagel, 2017 Allosiopelus N.Ito, 1995 Allotrechiama Ueno, 1970 Allotriopus Bates, 1882 Alpiodytes Jeannel, 1957 Altagonum Darlington, 1952 Amara Bonelli, 1810 Amara Stephens, 1827 Amarophilus Larochelle & Larivière, 2022 Amaroschema Jeannel, 1943 Amarotypus Bates, 1872 Amaroxenus Larochelle & Larivière, 2022 Ambigopenetretus Zamotajlov, 2002 Amblops Andrewes, 1931 Amblycheila Say, 1830 Amblycoleus Chaudoir, 1872 Amblygnathus Dejean, 1829 Amblypennis Amblystogenium Enderlein, 1905 Amblystomus Erichson, 1837 Amblytelus Erichson, 1842 Amelus Chaudoir, 1873 Ameoroduvalius Valentine, 1952 Amerinus Casey, 1884 Amerizus Chaudoir, 1868 Ameroduvalius Valentine, 1952 Ametroglossus Sloane, 1914 Amomotarus Amorphomerus Sloane, 1923 Amphasia Newman, 1838 Amphibia Blainville, 1816 Amphibia Linne, 1758 Amphimasoreus Piochard de la Brûlerie, 1875 Amphimenes Bates, 1873 Amphimenoides Kirschenhofer, 1999 Amphitasus Bates, 1871 Amphithasus Bates, 1871 Amphoxyne Bode, 1953 Amphyginus Haliday, 1841 Analoma Darlington, 1971 Anasis Laporte, 1867 Anatrichis LeConte, 1853 Anaulacus W.S.MacLeay, 1825 Anchista Nietner, 1856 Ancholeus Dejean, 1828 Anchomenidius Heyden, 1880 Anchomenus Bonelli, 1810 Anchonoderus Reiche, 1843 Anchotrechus Jeannel, 1927 Ancus Putzeys, 1867 Ancystroglossus Chaudoir, 1863 Andinocolpodes Perrault, 1991 Andinodontis Erwin & Toledano, 2010 Andinorites Mateu & Belles, 1980 Andinotrechus Mateu, 1981 Andrewesa Netolitzky, 1931 Andrewesia Csiki, 1932 Andrewesia Liebke, 1938 Andrewesinulus Straneo, 1938 Andrewesius Andrewes, 1939 Androzelma Dostal, 1993 Anentmetus Andrewes, 1924 Angionychus Klug, 1853 Anguloderus Anichtchenko & Sciaky, 2017 Angustanillus Baehr & Main, 2016 Anheterus Putzeys, 1868 Aniara Hope, 1838 Anillaspis Casey, 1918 Anillidius Jeannel, 1928 Anillinus Casey, 1918 Anillodes Jeannel, 1963 Anillopsidius Coiffait, 1969 Anillopsis Jeannel, 1937 Anillotarsus Mateu, 1980 Anillus Jacquelin du Val, 1851 Anisocnemus Chaudoir, 1843 Anisodactylus Dejean, 1829 Anisostichus Emden, 1953 Anomophaenus Fauvel, 1882 Anomostomus LaFerté-Sénectère, 1853 Anomotachys Jeannel, 1946 Anomotariella Baehr, 2012 Anomotarus Chaudoir, 1875 Anoncopencus Chaudoir, 1871 Anoncopeucus Chaudoir, 1872 Anophthalmus Sturm, 1844 Antarctiola Straneo, 1977 Antarctonomus Chaudoir, 1861 Antarctotrechus Ashworth & Erwin, 2016 Antennaria Dokhtouroff, 1883 Antephilorhizus Kirichenko-Babko, 2021 Anthia Weber, 1801 Anthracus Motschulsky, 1850 Antilliscaris Bänninger, 1949 Antilliscaris Lorenz, 1998 Antillodes Jeannel, 1963 Antimerina Alluaud, 1897 Antipionycha Liebke, 1928 Antireicheia Basilewsky, 1951 Antisphodrus Carret, 1907 Antroforceps Barr, 1967 Aparupa Andrewes, 1930 Apatrobus Habu & Baba, 1960 Apenes LeConte, 1851 Apenetretus Kurnakov, 1960 Aphaenopidius G.Müller, 1913 Aphaenops Bonvouloir, 1862 Aphaenopsis G.Müller, 1913 Aphaonus Reitter, 1887 Aplothorax G.R.Waterhouse, 1842 Apocimmerites Belousov, 1998 Apoduvalius Jeannel, 1953 Apophylon B.Gueorguiev & Sciaky, 2015 Apoplotrechus Alluaud, 1915 Aporesthus Bates, 1871 Apostolus Ball & Shpeley, 2002 Apotomus Illiger, 1807 Apristomimus Mateu, 1969 Apristus Chaudoir, 1846 Apritomimus Mateu, 1969 Apsaustodon Tschitscherine, 1901 Apsidocnemus Alluaud, 1936 Apterodromites Mateu, 1976 Apteroessa Hope, 1838 Apteromimus Wollaston, 1877 Aptinoderus Hubenthal, 1919 Aptinus Bonelli, 1810 Aputrechisibus Trezzi, 2007 Aquilex Moret, 1989 Arame Andrewes, 1919 Arantor Gistel, 1838 Archaeocindis Kavanaugh & Erwin, 1991 Archaeonebria Kavanaugh & J.Schmidt, 2019 Archagonum Basilewsky, 1953 Archicolliuris Liebke, 1931 Archicolpodes J.Schmidt, 2001 Archidela Rivalier, 1963 Archileistobrius Shilenkov & Kryzhanovskij, 1983 Archipatrobus Zamotajlov, 1992 Arctaphaenops Meixner, 1925 Ardistomis Putzeys, 1846 Ardistomopsis Straneo & Ball, 1989 Argentinatachoides Sallanave & Erwin&Roig-Juñent, 2008 Argiloborus Jeannel, 1937 Argutoridius Chaudoir, 1876 Arhytinus Bates, 1889 Aristes Dufour, 1820 Aristochroa Tschitscherine, 1898 Aristochroodes Marcilhac, 1993 Aristolebia Bates, 1892 Aristopus LaFerté-Sénectère, 1853 Arrhostus Motschoulsky, 1864 Arrowina R.T. & J.R.Bell, 1978 Arsinoe Laporte, 1834 Arthropterillus Wasmann, 1926 Arthropterites Wasmann, 1926 Arthropterus W.S.MacLeay, 1838 Asaphidion Gozis, 1886 Asioreicheia Bulirsch & Magrini, 2014 Asios Liebke, 1933 Askalaphium Liebke, 1938 Asklepia Liebke, 1938 Aspasiola Chaudoir, 1877 Aspidaphaenops Ueno, 2006 Aspidoglossa Putzeys, 1846 Assadecma Basilewsky, 1982 Astastus Péringuey, 1896 Astigis Rambur, 1838 Ataktosites Haupt, 1956 Atelidium Sharp, 1903 Athrostictus Bates, 1878 Atlantomasoreus Mateu, 1984 Atlantotrechus Lompe, 1999 Atractonotus Perroud, 1847 Atranodes Jedlicka, 1953 Atranopsis Baehr, 1982 Atranus LeConte, 1847 Atrirabus Hong & Wang, 1990 Atrotus Péringuey, 1896 Auchmerus Andrewes, 1929 Aulacolius Sloane, 1923 Aulacopodus Britton, 1940 Australicapitona Australomasoreus Baehr, 2007 Australovelinda Baehr, 2012 Austranillus Giachino, 2005 Austroglyptolemus Austroglyptolenus Roig-Juñent, 2003 Austropseudomorpha Baehr, 1997 Austrotrechus B.Moore, 1972 Awatrechus Ueno, 1955 Axinidium Sturm, 1843 Axinopalpus LeConte, 1846 Axinotoma Dejean, 1829 Aztecarpalus Ball, 1970 Badister Clairville, 1806 Baehrogenius Dostal, 2017 Baenningeria Reichardt, 1976 Baenningostichus Dostal, 1999 Baeoglossa Chaudoir, 1850 Bafutyphlus Bruneau de Miré, 1986 Baiopenetretus Zamotajlov, 2002 Balazucellus Deuve, 2001 Baldochlaenius Basilewsky, 1950 Balligratus Moret & Ortuño, 2017 Balticarthropterus Nagel, 1987 Balticeler Schmidt & Maddison, 2021 Bamaroodes Gueorguiev, 2014 Baripus Dejean, 1828 Barrymooreana Baehr, 1997 Barylaus Liebherr, 1985 Barysomus Dejean, 1829 Bascanus Péringuey, 1896 Basilewskya Straneo, 1948 Basilewskyana Kult, 1959 Basistichus Sloane, 1917 Batangocarabus Imura, 2002 Bathytrechus Ueno, 2005 Batoscelis Dejean, 1836 Batoscelis Lacordaire, 1854 Beckeria Jedlicka, 1931 Bedeliolus Semenov, 1900 Bellavalentis Hunting & Yang, 2019 Bellogenus Clarke, 1971 Bembidarenas Erwin, 1972 Bembidiomorphum Champion, 1918 Bembidion Latreille, 1802 Bembidium Latreille, 1802 Bembidium Stephens, 1829 Bembidromus Toledano, 2000 Bennigsenium W.Horn, 1897 Berberomerus Zamotajlov, 2002 Beronaphaenops B.V.Gueorguiev, 2012 Bhutanillus M.E.Schmid, 1975 Binaghites Jeannel, 1937 Birmaphaenops Deuve, 2017 Blackburnia Sharp, 1878 Blemus Dejean, 1821 Blennidus Motschulsky, 1866 Blenniventer Straneo, 1986 Blethisa Bonelli, 1810 Bleusea Bedel, 1897 Boeomimetes Péringuey, 1896 Bohemania Putzeys, 1863 Bohemaniella Bousquet, 2002 Boldoriella Jeannel, 1928 Bordoniella Mateu, 1989 Boreaphaenops Ueno, 2002 Bothrocolpodes Basilewsky, 1985 Bothynoproctus Tschitscherine, 1900 Bothynotrechus B.Moore, 1972 Bottchrus Jedlibcka, 1935 Bountya Townsend, 1971 Brachichila Chaudoir, 1870 Brachidius Chaudoir, 1852 Brachinites Fritsch, 1882 Brachinulus Basilewsky, 1958 Brachinus Weber, 1801 Brachyctis Chaudoir, 1870 Brachygnathus Perty, 1830 Brachynillus Reitter, 1904 Brachynopterus Bedel, 1898 Brachyodes Jeannel, 1949 Brachypelus Putzeys, 1867 Bracteon Bedel, 1879 Bradybaenus Dejean, 1829 Bradycellus Erichson, 1837 Bradycidus Casey, 1914 Brasiella Rivalier, 1954 Brigalowia Baehr, 2006 Brithysternum W.J.MacLeay, 1873 Bronislavia Semenov, 1891 Broscodera Lindroth, 1961 Broscodes Bolivar y Pieltain, 1914 Broscomimus Semenov, 1906 Broscosoma Rosenhauer, 1846 Broscus Panzer, 1813 Broter Andrewes, 1923 Bruskespar Morvan, 1998 Bruskmoal Morvan, 1998 Bryanites Valentine, 1987 Brzoskaicheila J.Moravec, 2012 Buderes Murray, 1857 Bulirschia Giachino, 2008 Burmapseudomorphus Beutel, Liu, Fikáček, Pang & Ślipiński, 2019 Bylibaraphanus Giachino & Eberhard&Perina, 2021 Caecidium Ueno, 1971 Caecocaelus Straneo, 1949 Caeconannus Jeannel, 1963 Caecoparvus Jeannel, 1937 Caelopenetretus Zamotajlov & Ito, 2000 Caelostomus W.S.MacLeay, 1825 Cainogenion Notman, 1925 Calathidius Putzeys, 1873 Calathocosmus Emden, 1928 Calathomimus Bates, 1886 Calathosoma Jeannel, 1938 Calathus Bonelli, 1810 Calathus Stephens, 1827 Caledonica Chaudoir, 1861 Caledonomorpha W.Horn, 1897 Caledyschirius Bulirsch, 2010 Calleida Latreille, 1824 Calleidomorpha Motschulsky, 1855 Callidadelpha Steinheil, 1875 Callidagonum Lorenz, 1998 Callidema Guérin-Méneville, 1843 Callindrothius Habu, 1984 Callisthenes Fischer von Waldheim, 1821 Callistochrous Chaudoir, 1850 Callistoides Motschulsky, 1865 Callistomimus Chaudoir, 1872 Callistus Bonelli, 1810 Callytron Gistel, 1848 Calochroa Hope, 1838 Calodromius Reitter, 1905 Calomera Motschulsky, 1862 Calopachys Haury, 1880 Calophaena Klug, 1821 Calophaenoidea Liebke, 1930 Calosoma Weber, 1801 Calybe Laporte, 1834 Calyptoglossa Jeannel, 1946 Calyptogonia Sloane, 1920 Cameroniola Baehr, 2000 Camptidius Putzeys, 1867 Camptodontus Dejean, 1826 Camptogenys Tschitscherine, 1899 Camptoscelis Dejean, 1828 Camptotoma Reiche, 1843 Canarobius Machado, 1987 Capabatus Csiki, 1930 Caphora Schmidt-Goebel, 1846 Carabidium Westwood, 1854 Carabidomemnus Kolbe, 1924 Carabites Heer, 1852 Carabites Ponomarenko, 1977 Carabomorphus Kolbe, 1895 Carabops Jacobson, 1900 Carabus Linnaeus, 1758 Carayonites Bruneau de Miré, 1986 Carbonellia Mateu, 1968 Cardiaderus Dejean, 1828 Cardiomera Bassi, 1834 Carenidium Chaudoir, 1868 Carenochyrus Solsky, 1874 Carenum Bonelli, 1813 Carterus Dejean, 1830 Casaleaphaenops Tian & Huang&Ma, 2021 Casaleius Sciaky & Wrase, 1998 Cascellius Curtis, 1838 Casnonia Latreille & Dejean, 1822 Cassolaia Wiesner, 1985 Castelnaudia Tschitscherine, 1891 Castlenaudia Catacolpodes Basilewsky, 1985 Catadromus W.S.MacLeay, 1825 Catalanodytes Sciaky, 1989 Catapiesis Solier, 1835 Catascopellus Straneo, 1969 Catascopus Kirby, 1825 Cathaiaphaenops Deuve, 1996 Caucasaphaenops Belousov, 1999 Caucasorites Belousov & Zamotajlov, 1997 Cavazzutiocarabus Imura, 1998 Cavicarabus Hong, 1991 Cedorum Borges & Serrano, 1993 Cedrorum Borges & A.Serrano, 1993 Celaenagonum Habu, 1978 Celaenephes Schmidt-Goebel, 1846 Celanida Laporte, 1867 Celioinkosa Straneo, 1951 Cenogmus Sloane, 1898 Cenothyla Rivalier, 1969 Cepahlostichus Straneo, 1977 Cephalophonus Ganglbauer, 1891 Cephalosdrophus Lassalle & Marcilhac, 1999 Cephalostichus Straneo, 1977 Cephalota Dokhtouroff, 1883 Cerabilia Castelnau, 1867 Cerabilia Laporte, 1867 Cerapterites Wasmann, 1926 Cerapterus Swederus, 1788 Ceratoderus Westwood, 1841 Ceroglossus Solier, 1848 Cerotalis Laporte, 1867 Chaetagonum Burgeon, 1933 Chaetapatrobus Lafer, 1996 Chaetauchenium Tschitscherine, 1900 Chaetobroscus Semenov, 1900 Chaetocrepis Chaudoir, 1857 Chaetodactyla Tschitscherine, 1897 Chaetodera Jeannel, 1946 Chaetoduvalius Jeannel, 1928 Chaetogenys van Emden, 1958 Chaetomargoreicheia Magrini & Bulirsch, 2005 Chaetopatrobus Lafer, 1996 Chaetosaurus J.Schmidt, 2001 Chaetotrechiama Ueno, 1982 Chalcochrous Chaudoir, 1838 Chaltenia Roig-Juñent & Cicchino, 2001 Chaudoirella Mateu, 1982 Chaudoirina Mateu, 1955 Cheilagona Baehr, 2006 Cheilonycha Lacordaire, 1842 Cheiloxya Guérin-Méneville, 1855 Chelonodema Laporte de Castelnau, 1834 Chenotrechus Tian, Huang & Jia, 2023 Chiapadytes Vigna Taglianti, 1977 Chilotomus Chaudoir, 1842 Chinapenetretus Kurnakov, 1963 Chinapterus Berlov, 1998 Chlaeminus Motschulsky, 1865 Chlaeniellus Reitter, 1908 Chlaenius Bonelli, 1810 Chrysocaraboides Deuve, 2012 Chu Tian & He, 2020 Chydaeus Chaudoir, 1854 Chylnus Sloane, 1920 Cicindela Linnaeus, 1758 Cicindelidia Rivalier, 1954 Cicindelina Latreille, 1802 Cicindelites Meunier, 1898 Cicindis Bruch, 1908 Cillenus Leach, 1819 Cillenus Samouelle, 1819 Cimbrodytes Piva, 2021 Cimmerites Jeannel, 1928 Cimmeritodes Deuve, 1996 Cinctagonum Baehr, 2012 Cintaroa Kasahara, 1989 Cintaromorpha Häckel & Anichtchenko, 2016 Cistelagonum Baehr, 2012 Ckacus Tian & Huang, 2021 Clarencia Sloane, 1917 Climax Putzeys, 1861 Clinidium Kirby, 1830 Clivina Julieni Lesne P.Lesne, 1896 Clivina Latreille, 1802 Clivinarchus Sloane, 1896 Clivinopsis Bedel, 1895 Clypeuspinus Balkenohl, 2021 Cnemalobus Guérin-Méneville, 1838 Cnides Motschulsky, 1862 Coarazuphium Gnaspini & Vanin&Godoy, 1998 Coarazuphium Pellegrini & Ferreira, 2017 Coccocarabus Imura, 2002 Colantarctia Colasidia Basilewsky, 1954 Coleolissus Bates, 1892 Colfax Andrewes, 1920 Collagonum Baehr, 1995 Colliuris DeGeer, 1774 Collyris Fabricius, 1801 Colpocaccus Sharp, 1903 Colpodes Swezey, 1942 Colpodes W.S.MacLeay, 1825 Colpodichius Straneo, 1952 Colpoides Jedlicka, 1931 Colpomimus Basilewsky, 1985 Colposhodrus Colposphodrus Jedlicka, 1953 Colpostoma Semenov, 1889 Columbitrechus Mateu, 1982 Conchitella Moore, 1962 Conexicoxa Linnaeus, 1986 Conjunctia Ponomarenko, 1977 Cophosomorpha Tschitscherine, 1891 Coptia Brullé, 1835 Coptocarpus Chaudoir, 1857 Coptodera Dejean, 1825 Coptodrenella Hansen, 1968 Coptoglossus Chaudoir, 1870 Coptolobus Chaudoir, 1857 Coptoptera Chaudoir, 1837 Coptopterella Basilewsky, 1961 Coptopterina Basilewsky, 1956 Cordorabus Ponomarenko, 1977 Coreoblemus Ueno, 1969 Coreocarabus Imura, 2002 Corintascaris Basilewsky, 1952 Corsyra Dejean, 1825 Coryza Putzeys, 1867 Cosmodela Rivalier, 1961 Cosmodiscus Sloane, 1907 Cosnania Dejean, 1821 Costitachys Erwin, 1974 Cothresia Jeannel, 1964 Crasodactylus Guérin-Méneville, 1847 Craspedonotus Schaum, 1863 Craspedophorus Hope, 1838 Craspophorus Crassacantha Baehr, 1995 Crassagena Baehr, 2006 Cratacanthus Dejean, 1829 Craterofonia Cratocechenodes Deuve, 2003 Cratocerus Dejean, 1829 Cratoferonia Tschitscherine, 1902 Cratogaster Blanchard, 1843 Cratognathus Dejean, 1829 Cratohaerea Chaudoir, 1850 Cratosoma Jeannel, 1948 Creagris Nietner, 1857 Crenulostigus Straneo, 1942 Crenulostrigus Straneo, 1942 Creobius Guérin-Méneville, 1838 Crepidogaster Boheman, 1848 Crepidogastrillus Basilewsky, 1959 Crepidogastrinus Basilewsky, 1957 Crepidolomus Basilewsky, 1959 Crepidonellus Basilewsky, 1959 Crepidopterus Chaudoir, 1855 Crepidozaena Deuve, 2001 Creprostus Imura, 2002 Cretomigadops Liu, Beutel, Makarov, Jarzembowski, Xiao & Luo, 2022 Cretomomophron Cretomophron Rosova, Prokop & Beutel, 2023 Cretorabus Ponomarenko, 1977 Cretotetracha Zhang, Zhang & Chen&Wang, 2019 Cribrodyschirius Bruneau de Miré, 1952 Criniventer Emden, 1953 Crisimus Habelmann, 1885 Croatotrechus Casale & Jalzic, 1999 Crossacantha Baehr, 1995 Cryptobatis Eschscholtz, 1829 Cryptocephalomorpha Ritsema, 1875 Cryptocharidius M.Etonti & Mateu, 1992 Cryptocolliuris Basilewsky, 1955 Cryptomma Putzeys, 1846 Cryptophonus Brandmayr & Zetto-Brandmayr, 1982 Cryptorites Jeannel, 1950 Cryptoscaphus Chaudoir, 1855 Ctenodactyla Dejean, 1825 Ctenognathus Fairmaire, 1843 Ctenostoma Klug, 1821 Cuneipectus Sloane, 1907 Curtonotus Stephens, 1827 Cyanotarus Reed, 1874 Cychropsis Boileau, 1901 Cychrus Fabricius, 1794 Cyclicus Jeannel, 1949 Cycloloba Chaudoir, 1850 Cyclosomus Latreille, 1829 Cyclotrachelus Chaudoir, 1838 Cylindera Westwood, 1831 Cylindrocranius Chaudoir, 1878 Cylindronotum Putzeys, 1845 Cylindropectus Lorenz, 1998 Cylindropectus Mateu, 1974 Cymatopterus Lomnicki, 1894 Cymbionotum Baudi di Selve, 1864 Cymenopterus Jeannel, 1948 Cymindis Latreille, 1805 Cymindoidea Laporte, 1833 Cymindoides Motschulsky, 1856 Cynthidia Chaudoir, 1874 Cyphocoleus Chaudoir, 1877 Cypholoba Chaudoir, 1850 Cyphotrechodes Jeannel, 1926 Cyptomicrus Vinson, 1939 Cyrtoderus Hope, 1842 Cyrtolaus Bates, 1882 Cyrtomoscelis Chaudoir, 1874 Cyrtopilus Basilewsky, 1985 Dabatrechus Ueno, 2004 Dactyleurys Tschitscherine, 1899 Dactylinius Straneo, 1941 Dactylotrechus Belousov & Kabak, 2003 Daer Semenov & Znojko, 1929 Daiconotrechus Ueno, 1971 Dailodontus Reiche, 1843 Dalatagonum Fedorenko, 2011 Dalmataphaenops Monguzzi, 1993 Dalmatoreicheia Magrini & Bulirsch, 2005 Dalyat Mateu, 2002 Damasteroides Deuve, 2012 Daptus Fischer von Waldheim, 1823 Darlingtonea Valentine, 1952 Darlingtonica Cassola, 1986 Darodilia Castelnau, 1868 Darodilia Laporte, 1867 Dasiosoma Britton, 1937 Decogmus Sloane, 1915 Deipyrodes Bousquet, 2002 Deliaesianum Morvan, 1999 Delinius Westwood, 1864 Deltocolpodes Morvan, 1992 Deltomerodes Deuve, 1992 Deltomerus Motschulsky, 1850 Demetrias Bonelli, 1810 Demetrida White, 1846 Demetriola Jeannel, 1949 Dendragonum Jeannel, 1948 Dendromigadops Baehr, 2013 Denudirabus Ren & Lu&Guo, 1995 Deqenocarabus Imura, 2002 Dercylinus Chaudoir, 1883 Dercylus Laporte, 1832 Derocrania Chaudoir, 1861 Deroplectoides Deuve, 2012 Derossiella Queinnec, 2008 Derostichus Motschulsky, 1860 Desera Hope, 1831 Deuveaphaenops Tian & Huang, 2017 Deuveopsis Quéinnec & Ollivier, 2021 Dhanya Andrewes, 1919 Dhysores Grouvelle, 1903 Diabaticus Bates, 1878 Diacanthostylus Habu, 1978 Diacheila Motschulsky, 1844 Diachipteryx Alluaud, 1925 Diachromus Erichson, 1837 Diamella Shi & Liang, 2013 Dianotrechus Tian, 2016 Diaphoromerus de Chaudoir, 1843 Diastrophella Rivalier, 1957 Dicaelindus W.S.MacLeay, 1825 Dicaelus Bonelli, 1813 Diceromerus Chaudoir, 1873 Diceromerus Motschulsky, 1872 Dichaetochilus Basilewsky, 1946 Dicheirotrichus Jacquelin du Val, 1855 Dicheirus Mannerheim, 1843 Dicranoglossus Chaudoir, 1872 Dicranoncoides Habu, 1978 Dicranoncus Chaudoir, 1850 Dicraspeda Chaudoir, 1863 Dicrochile Guérin-Méneville, 1846 Dicrodontus Chaudoir, 1872 Dicropterus Ehlers, 1883 Diglymma Sharp, 1886 Dilatotarsa Dokhtouroff, 1882 Dilonchus Andrewes, 1936 Dimorphopatrobus Casale & Sciaky, 1994 Dimorphoreicheia Magrini & Fancello&Leo, 2002 Dinocolpodes J.Schmidt, 2001 Dinodes Bonelli, 1810 Dinopelma Bates, 1889 Dinoscaris Alluaud, 1902 Diodercarus Lutshnik, 1931 Dioryche W.S.MacLeay, 1825 Diplacanthogaster Liebke, 1932 Diplochaetus Chaudoir, 1872 Diplocheila Brullé, 1835 Diplochila Brullé, 1834 Diploharpus Chaudoir, 1850 Diplous Motschulsky, 1850 Dirotus W.S.MacLeay, 1825 Dischissus Bates, 1873 Disciferella Kataev & Muilwijk, 2020 Discoderus LeConte, 1853 Discoptera Semenov, 1889 Disphaericus Waterhouse, 1842 Disphericus G.R.Waterhouse, 1842 Dister Morvan, 2006 Distichus Motschulsky, 1858 Distipsidera Westwood, 1837 Distrigidius Jeannel, 1948 Distrigus Dejean, 1828 Ditomus Bonelli, 1810 Dixus Billberg, 1820 Do Baehr, 2009 Dobodura Darlington, 1968 Doderotrechus Vigna Taglianti, 1968 Dolichocolpodes Basilewsky, 1985 Dolichoctis Schmidt-Goebel, 1846 Dolichus Bonelli, 1810 Doliodactyla Sciaky & Wrase, 1998 Dongoblemus Deuve & Tian, 2016 Dongodytes Deuve, 1993 Dontolobus Basilewsky, 1970 Dracotrechus Ueno, 2010 Dregus Motschulsky, 1864 Dromica Dejean, 1826 Dromicoida Werner, 1995 Dromidea Perroud & Montrouzier, 1864 Dromiops Péringuey, 1898 Dromistomus Jeannel, 1948 Dromius Bonelli, 1810 Dromoceryx Schmidt-Goebel, 1846 Dromochorus Guérin-Méneville, 1849 Drymatus Motschulsky, 1862 Drymonaxus Straneo, 1941 Drypta Latreille, 1797 Duvaliaphaenops Giordan, 1984 Duvalioblemus Deuve, 1995 Duvaliomimus Jeannel, 1928 Duvaliopsis Jeannel, 1928 Duvalius Delarouzée, 1859 Dyscaris Bänninger, 1940 Dyscherinus Jeannel, 1955 Dyscherus Chaudoir, 1855 Dyschiridium Chaudoir, 1861 Dyschiriodes Jeannel, 1941 Dyschiriomimus Iablokoff-Khnzorian, 1960 Dyschirius Bonelli, 1810 Dyschirius Panzer, 1813 Dyscolus Dejean, 1831 Dystipsidera Dystrichothorax Blackburn, 1892 Eccoptomenus Chaudoir, 1850 Eccoptoptera Chaudoir, 1878 Ecnomalus Bates, 1892 Ecnomolaus Bates, 1892 Ecotrechus Tian, Huang & Jia, 2023 Ectenognathus Murray, 1858 Ectinothorax Alluaud, 1941 Ectomomesa Chaudoir, 1871 Ecuadoritrechus Deuve, 2002 Ega Laporte, 1834 Egadyla Alluaud, 1916 Egaploa Alluaud, 1916 Elaphropus Motschulsky, 1839 Elaphrotites Haupt, 1856 Elaphrus Fabricius, 1775 Elgonotyphlus Sciaky & Zaballos, 1993 Ellipsoptera Dokhtouroff, 1883 Elliptoleus Bates, 1882 Elliptosoma Wollaston, 1854 Emphanes Motschulsky, 1850 Enantiola Rivalier, 1961 Enceladus Bonelli, 1813 Endosomatium Wollaston, 1877 Endynomena Chaudoir, 1873 Enoicus Péringuey, 1896 Entomoantyx Ball & McCleve, 1990 Eobroscus Kryzhanovskij, 1951 Eocarterus Stichel, 1923 Eoclivina Kult, 1959 Eocnides Jeannel, 1954 Eodontium Eohomopterus Wasmann, 1919 Eopaussus Wasmann, 1926 Eosphodrus Casale, 1988 Eotrechodes Ueno & Lafer&Sundukov, 1995 Epaphiopsis Ueno, 1953 Epaphiotrechus Deuve & Kavanaugh, 2016 Epaphius Leach, 1819 Epaphius Samouelle, 1819 Epaphius Stephens, 1827 Epelyx Blackburn, 1892 Epicolpodes Basilewsky, 1985 Epigraphodes Basilewsky, 1967 Epigraphoides Brasilewsky, 1967 Epigraphus Chaudoir, 1869 Epikastea Liebke, 1935 Epilectus Blackburn, 1888 Epimicodema Sloane, 1914 Epomis Bonelli, 1810 Erebotrechus Britton, 1964 Erectocolliuris Liebke, 1931 Eremolestes Maindron, 1905 Eremosphodrus Semenov, 1909 Erephognathus Alluaud, 1932 Eriophonus Tschitscherine, 1901 Eripus Dejean, 1829 Erwinanillus Giachino & Eberhard&Perina, 2021 Erwiniana Paulsen & A.Smith, 2003 Escolatrechus Mateu, 2003 Eucaerus LeConte, 1853 Eucamaragnathus Jeannel, 1938 Eucamptognathus Chaudoir, 1837 Eucarterus Reitter, 1900 Eucheila Dejean, 1829 Euchroa Brullé, 1835 Eucolliuris Liebke, 1931 Eucolpodes Jeannel, 1948 Eudalia Laporte, 1867 Eudromus Klug, 1835 Eujalmenus Bousquet, 2002 Eulampra Chaudoir, 1848 Euleptus Klug, 1833 Eumara Tschitscherine, 1901 Eunostus Laporte, 1835 Eunota Rivalier, 1954 Euphorticus G.Horn, 1881 Euplatia Chaudoir, 1873 Euplatyrhopalus Desneux, 1905 Euplynes Schmidt-Goebel, 1846 Euproctinus Leng & Mutchler, 1927 Euprosopus Dejean, 1825 Euryabax Fauvel, 1903 Euryaptus Bates, 1892 Euryarthron Guérin-Méneville, 1849 Eurycalleida Maindron, 1905 Eurycallida Maindron, 1905 Eurycaraboides Deuve, 2012 Eurycoleus Chaudoir, 1848 Eurydera Laporte, 1831 Euryderus LeConte, 1846 Eurygnathus Wollaston, 1854 Eurylychnus Bates, 1891 Eurymorpha Hope, 1838 Eurynebria Ganglbauer, 1891 Eurypercus Jeannel, 1948 Euryscaphus W.J.MacLeay, 1865 Eurystomis Chaudoir, 1878 Euschizomerus Chaudoir, 1850 Eustra Schmidt-Goebel, 1846 Euthenarus Bates, 1874 Eutogeneius Solier, 1849 Eutrechopsis B.Moore, 1972 Eutrechus B.Moore, 1972 Eutrichopus Tschitscherin, 1897 Euzona Rivalier, 1963 Evolenes LeConte, 1853 Externanillus Baehr & Main, 2016 Falsodromius Mateu, 1976 Fangshania Hong, 1981 Feostoma Straneo, 1941 Feronia Feroniascus Jeannel, 1951 Feroniola Tschitscherine, 1900 Feronista Moore, 1965 Feronites Fritsch, 1884 Filicerozaena Deuve, 2001 Fonsecacarabus Martins-Neto & Mendes, 2002 Forcipator Maindron, 1904 Formosiella Jedlicka, 1951 Fortagonum Darlington, 1952 Foveocrotaphus Anichtchenko, 2014 Foveodromica Cassola, 2003 Fulgenticarabus Deuve & Mourzine, 1998 Fuminoria Morita, 2006 Gaioxenus Broun, 1910 Galerita Fabricius, 1801 Galeritina Kirby, 1825 Galicioreicheia Felix & Bulirsch, 2015 Galiciotyphlotes Assmann, 1999 Gallerucidia Chaudoir, 1872 Gastragonum Darlington, 1952 Gastrellarius Casey, 1918 Gastrogmus Sloane, 1915 Geballusa Erwin, 1994 Gehringia Darlington, 1933 Geobaenus Dejean, 1829 Geobius Dejean, 1831 Geocharidius Jeannel, 1963 Geocharis Ehlers, 1883 Geodromus Dejean, 1829 Geoffreyella Baehr, 2012 Geopinus LeConte, 1847 Georgeballius Habu, 1984 Geoscaptus Chaudoir, 1855 Geotrechus Jeannel, 1919 Geotrechus Moore, 1972 Gestroania Liebke, 1938 Giachinoana Baehr, 2003 Gibbozaena Deuve, 2001 Gidda Andrewes, 1920 Gigadema J.Thomson, 1859 Gilesdytes Giachino & Eberhard&Perina, 2021 Ginema Ball & Shpeley, 2002 Giraffaphaenops Deuve, 2002 Glabroduvalius Vrbica, Curcic & Antic&Curcic, 2013 Glabrotrechus Suenson, 1957 Glaucagonum Habu, 1978 Glenopterus Heer, 1847 Globulipalpus Sciaky & Facchini, 2019 Glossocarabus Imura, 2002 Glycia Chaudoir, 1842 Glyphodactyla Chaudoir, 1837 Glyptogrus Bates, 1875 Glyptolenoides Perrault, 1991 Glyptolenus Bates, 1878 Glyptus Brullé, 1837 Gnaphon Andrewes, 1920 Gnathaphanus W.S.MacLeay, 1825 Gnatholimnaeum Sharp, 1901-1910 Gnatholymnaeum Sharp, 1903 Gnathostenus Shpeley & Ball, 2000 Gnathoxys Westwood, 1842 Goedetrechus B.Moore, 1972 Gomerina Bolivar y Pieltain, 1940 Goniocellus Casey, 1914 Goniotropis G.R.Gray, 1831 Gonogenia Chaudoir, 1844 Goriresina Matalin, Perkovsky & Vasilenko, 2021 Gotoblemus Ueno, 1970 Gouleta Erwin, 1994 Gourlaya Britton, 1964 Gourlayia Britton, 1964 Gracilanillus Baehr & Main, 2016 Graciliblemus Deuve & Tian, 2016 Grammognatha Motschulsky, 1850 Grandipenetretus Zamotajlov & Sciaky, 1999 Grandopronotalia W.Horn, 1936 Graniger Motschulsky, 1864 Graphipterus Latreille, 1802 Gregorydytes Giachino & Eberhard&Perina, 2021 Grouvellina R.T. & J.R.Bell, 1978 Grundmannius Basilewsky, 1965 Guatemalatrechus Trezzi, 2003 Guatemalteca Erwin, 2004 Gugheorites Basilewsky, 1951 Guiaphaenops Deuve, 2002 Guiaphaenops Ueno, 1987 Guineica Rivalier, 1963 Guiodytes Tian, 2013 Guizhaphaenops Vigna Taglianti, 1997 Guizhaphaenopsodes Tian & Huang, 2021 Guizhaphaenopsoides Tian & Huang&Li, 2021 Gulaphaenops Ueno, 1987 Gunvorita Landin, 1955 Guyanemorpha Erwin, 2013 Gymnetoreicheia Magrini & Fancello&Casale, 2019 Gynandromorphus Dejean, 1829 Gynandropus Dejean, 1831 Gyrochaetostylus Habu, 1978 Habragonum Ueno, 1964 Habrodera Motschulsky, 1862 Habroscelimorpha Dokhtouroff, 1883 Habutarus Ball & Hilchie, 1983 Hadrocephalus Handlirsch, 1906 Haixiaphaenops Tian, Cheng & Huang, 2021 Hakaharpalus Larochelle & Larivière, 2005 Halocoryza Alluaud, 1919 Hannaphota Landin, 1955 Hansus Ball & Shipeley, 1983 Haplaner Chaudoir, 1878 Haplanister B.Moore, 1996 Haplobothynus Tschitscherine, 1901 Haplochlaenius Lutshnik, 1933 Haplocolpodes Jeannel, 1951 Haploferonia Darlington, 1962 Haplogaster Chaudoir, 1879 Haplopeza Boheman, 1848 Haplotrachelus Chaudoir, 1855 Haptoderidius Straneo, 1975 Haptoderodes Straneo, 1986 Harapalus Harpaglossus Motschulsky, 1858 Harpalinus Jeannel, 1946 Harpaliscus Bates, 1892 Harpalobrachys Tschitscherine, 1899 Harpalodiodes Bousquet, 2002 Harpalomimes Handlirsch, 1906 Harpalomimetes Schauberger, 1932 Harpalomimus Facchini & Giachino, 2020 Harpalomorphus Péringuey, 1896 Harpalophonus Ganglbauer, 1891 Harpalus Latreille, 1802 Harpathaumas Basilewsky, 1947 Harponixus Basilewsky, 1950 Harponxius Basilewsky, 1950 Harpostomus Chaudoir, 1856 Hartonymus Casey, 1914 Hebeicarabus Hong, 1983 Helenaea Schatzmayr & Koch, 1934 Helluapterus Sloane, 1914 Helluarchus Sloane, 1914 Hellunidius Helluo Bonelli, 1813 Helluobrochus Reichardt, 1974 Helluocolpodes Liebherr, 2005 Helluodema Laporte, 1867 Helluodes Westwood, 1847 Helluomorpha Laporte, 1834 Helluomorphoides Ball, 1951 Helluonidius Chaudoir, 1872 Helluopapua Darlington, 1968 Helluosoma Laporte, 1867 Hemiaulax Bates, 1892 Hemichlaenius Bates, 1892 Hemiplatynus Casey, 1920 Hemitelestus Alluaud, 1895 Henrotiochoromus Busulini, 1958 Henrotius Jeannel, 1953 Henvelik Morvan, 1999 Heptacarabus Imura, 1998 Heptodonta Hope, 1838 Herculagonum Baehr, 2002 Hesperanillus Baehr & Main, 2016 Heteracantha Brullé, 1835 Heteropaussus J.Thomson, 1860 Hexachaetus Chaudoir, 1872 Hexagonia Kirby, 1825 Hexaplatarthrus Jeannel, 1955 Hikosanoagonum Habu, 1954 Hiletus Schiödte, 1847 Himalaphaenops Ueno, 1980 Himalayozabrus Andujar & Serrano, 2000 Himalopenetretus Zamotajlov, 2002 Himalosphodrus Casale, 1988 Himalotrechodes Ueno, 1981 Himalotrechus Belousov & Kabak&J.Schmidt, 2019 Hippoloetis Laporte, 1835 Hirtoperigona Perrault, 1988 Holaxinidium Basilewsky, 1963 Holcaspis Chaudoir, 1865 Holcocoleus Chaudoir, 1883 Holcoderus Chaudoir, 1870 Holcoferonia Fedorenko, 2022 Holconotus Schmidt-Goebel, 1846 Hololeius LaFerté-Sénectère, 1851 Holoponerus Fairmaire, 1883 Holoprizus Putzeys, 1867 Holosoma Semenov, 1889 Homaloderodes Jeannel, 1962 Homalomorpha Brullé, 1837 Homalonesiota Maindron, 1908 Homethes Newman, 1842 Homodela Rivalier, 1950 Homopterus Westwood, 1841 Honduranillus Zaballos, 1997 Hoplauchenium Tschitscherin, 1900 Hoplizomenus Chaudoir, 1873 Hoplodactylella Strand, 1936 Hoplogaster Chaudoir, 1879 Hoplolenus LaFerté-Sénectère, 1851 Hoquedela Müller-Motzfeld, 1988 Hormacrus Sloane, 1898 Horniulus Jedlicka, 1932 Horologion Valentine, 1932 Howareutus Acorn, 1984 Hujia Song, Jarzembowski & Xiao, 2022 Huoyanodytes Tian & Huang, 2016 Hurites Tian, Huang & Jia, 2023 Hyboptera Chaudoir, 1873 Hybopteroides Erwin & Ball, 2012 Hybothecus Chaudoir, 1874 Hydraphaenops Jeannel, 1926 Hydrotrechus Carabajal, Garcia-Carrillo & Rodriguez-Fernandez, 2000 Hygranillus B.Moore, 1980 Hypaetha LeConte, 1857 Hyperectenus Alluaud, 1935 Hyperion Laporte, 1834 Hyphaereon W.S.MacLeay, 1825 Hypharpax W.S.MacLeay, 1825 Hypodipnites Jeannel, 1963 Hypotyphlus Jeannel, 1937 Hypsipezum Alluaud, 1917 Hystrichopus Boheman, 1848 Hystricosphodrus Casale & Giachino, 2004 Iason Giachino & Vailati, 2011 Iberanillus Español, 1971 Iberodytes Jeannel, 1949 Iberotrechodes Faille et al., 2021 Iberotrechus Jeannel, 1920 Ichnusodytes Magrini & Fancello&Onnis, 2019 Idacarabus Lea, 1910 Idiagonum Darlington, 1952 Idiastes Andrewes, 1931 Idiocolpodes Basilewsky, 1985 Idiomelas Tschitscherine, 1900 Idiomorphus Chaudoir, 1846 Ifasina Jeannel, 1946 Ifridytes Deuve & Queinnec, 1994 Iga Ueno, 1953 Ildobates Español, 1966 Illaphanus W.J.MacLeay, 1865 Imasakaia Arai & Morita, 2010 Incagonum Liebherr, 1994 Incatrechus Mateu & Belles, 1982 Indiophonus N.Ito, 1996 Indocarterus Kataev & Wrase, 2018 Indopatrobus Zamotajlov & Wrase, 2006 Infernophilus Larson, 1969 Inflatozaena Deuve, 2001 Inkosa Péringuey, 1926 Inotrechus Dolzhanski & Ljovushkin, 1989 Inpa Erwin, 1978 Insulanebria Kavanaugh, 2021 Iranopogonus Azadbakhsh, 2019 Iresia Dejean, 1829 Iridagonum Darlington, 1952 Ischnagonum Kasahara & Satô, 1997 Ishidatrechus Ueno, 1956 Ishikawatrechus Habu, 1950 Italaphaenops Ghidini, 1964 Italodytes G.Müller, 1938 Italozabrus Andujar & Serrano, 2000 Itamus Schmidt-Goebel, 1846 Iyotrechus Ueno & Naito, 2009 Jalzicaphaenops Lohaj & Lakota, 2010 Jansenia Chaudoir, 1865 Jeannelius Kurnakov, 1959 Jiangxiaphaenops Ueno & Clarke, 2007 Jiulongotrechus Tian & Huang&Wang, 2015 Jocqueius Basilewsky, 1988 Josefia Toledano, 2000 Jujiroa Ueno, 1952 Junaphaenops Ueno, 1997 Jundlandia Duran & Gough, 2022 Junnanotrechus Ueno & Yin, 1993 Kaffovatus Clarke, 1972 Kalchdigor Morvan, 1999 Kar Morvan, 1998 Kareya Andrewes, 1919 Karnes Morvan, 2010 Kashmirobroscus J.Schmidt & Wrase&Sciaky, 2013 Kaszabellus Jedlicka, 1954 Kaveinga R.T. & J.R.Bell, 1978 Kearophus Dajoz, 2004 Kenodactylus Broun, 1909 Kenyacus Alluaud, 1917 Kenyoreicheia Bulirsch & Magrini, 2007 Kenyoriecheia Bulirsch & Magrini, 2007 Kettlotrechus Townsend, 2010 Kimberleytyphlus Giachino & Eberhard&Perina, 2021 Kiwiharpalus Larochelle & Larivière, 2005 Kiwiplatynus Larochelle & Larivière, 2021 Kiwitachys Larochelle & Larivière, 2007 Kiwitrechus Larochelle & Larivière, 2007 Klapperichella Jedlicka, 1956 Klepteromimus Péringuey, 1898 Klepterominus Péringuey, 1898 Klepterus Péringuey, 1896 Kosswigia Jeannel, 1947 Kozlovites Jeannel, 1935 Kozyrevius Berlov, 1998 Kryzhanovskiana Kataev & Kirejtshuk, 2019 Kteatus Liebke, 1936 Kuceraianum Morvan, 2002 Kultianella Perrault, 1994 Kupeharpalus Larochelle & Larivière, 2005 Kupeplatynus Larochelle & Larivière, 2021 Kupetrechus Larochelle & Larivière, 2007 Kupeus R.T. & J.R.Bell, 1982 Kurasawatrechus Yoshida & Nomura, 1952 Kuschelinus Straneo, 1963 Kusumia Ueno, 1952 Labocephalus Chaudoir, 1848 Laccocenus Sloane, 1890 Laccopterum W.J.MacLeay, 1878 Lachenus Putzeys, 1846 Lachnaces Bates, 1872 Lachnocrepis LeConte, 1853 Lachnoderma W.J.MacLeay, 1873 Lachnolebia Maindron, 1905 Lachnophorus Dejean, 1831 Lachnothorax Motschulsky, 1862 Lacordairia Laporte, 1867 Laemostenus Bonelli, 1810 Laemostenus Tahami, Muilwijk, Lohaj & Sadeghi, 2017 Laevagonum Darlington, 1952 Lampetes Andrewes, 1940 Lamprias Bonelli, 1810 Langea W.Horn, 1901 Lanxangaphaenops Deuve, 2012 Laoblemus Ueno, 2006 Laoreicheia Balkenohl, 2005 Laosaphaenops Deuve, 2000 Laparhetes Jeannel, 1946 Lasiocera Dejean, 1831 Lassalleianum Morvan, 1999 Lebia Latreille, 1802 Lebia Rasool, Abdel-Dayem, Felix & Aldhafer, 2018 Lebidema Motschulsky, 1864 Lebidia A.Morawitz, 1862 Lebina Germar, 1813 Lebioderus Westwood, 1838 Lebiomorphica Lorenz, 1998 Lebistina Motschulsky, 1864 Lebistinida Péringuey, 1898 Lecanomerus Chaudoir, 1850 Ledouxius Zamotajlov, 1992 Ledouxnebria Deuve, 1998 Leieuporites Straneo, 1960 Leiolesticus Roux & Lassalle&Dubault, 2016 Leiradira Laporte, 1868 Leiramara Hieke, 1988 Leistus Frölich, 1799 Leja Leleupanillus Basilewsky, 1976 Leleupaussus Luna de Carvalho, 1962 Leleupidia Basilewsky, 1951 Leleuporella Basilewsky, 1956 Leleuporites Straneo, 1960 Lelis Chaudoir, 1870 Leoglymmius R.T. & J.R.Bell, 1978 Lepcha Andrewes, 1930 Leptacolpodes Basilewsky, 1985 Leptagonum Kolbe, 1897 Leptocaraboides Deuve, 2012 Leptocolpodes Basilewsky, 1985 Leptognatha Rivalier, 1963 Leptosarcus Péringuey, 1896 Leptotrachelon Liebke, 1928 Leptotrachelus Latreille, 1829 Lessinodytes Vigna Taglianti, 1982 Lesticus Dejean, 1828 Lestignathus Erichson, 1842 Letouzeya Bruneau de Miré, 1982 Leuropus Andrewes, 1947 Liagonum Jeannel, 1948 Liamegalonychus Basilewsky, 1963 Libotrechus Ueno, 1998 Licentius Jedlicka, 1939 Lichnasthenus J.Thomson, 1858 Licinoderus Sainte-Claire Deville, 1905 Licinopsis Bedel, 1899 Licinus Latreille, 1802 Limnastis Motschulsky, 1862 Limnastus Motschulsky, 1862 Limodromus Motschulsky, 1850 Liochirus Tschitscherine, 1897 Liocolpodes Basilewsky, 1985 Liodaptus Bates, 1889 Lioholus Tschitscherine, 1897 Lionedya Chaudoir, 1871 Lionepha Casey, 1918 Lionychus Wissmann, 1846 Liopasa Tschitscherine, 1901 Lioptera Chaudoir, 1870 Liotachys Bates, 1871 Lipostratia Chaudoir, 1873 Lirabus Hong, 1992 Lissagonum Habu, 1978 Lissopogonus Andrewes, 1923 Lissopterus G.R.Waterhouse, 1843 Litangocarabus Imura, 2002 Litarthrum Sloane, 1915 Lithagonum Darlington, 1952 Lithorabus Ponomarenko, 1977 Lobatodes Basilewsky, 1956 Lobobrachus Sharp, 1885 Lobocolpodes Basilewsky, 1985 Lobodontidius Basilewsky, 1970 Lobodontulus Basilewsky, 1970 Lobodontus Chaudoir, 1842 Lonchosternus LaFerté-Sénectère, 1851 Lophidius Dejean, 1831 Lophocoryza Alluaud, 1941 Lophoglossus LeConte, 1853 Lophyra Motschulsky, 1860 Lophyridia Jeannel, 1946 Lorestania Anichtchenko, 2011 Loricera Latreille, 1802 Lorostema Motschulsky, 1865 Lovricia Pretner, 1979 Loxandrus LeConte, 1852 Loxocrepis Eschscholtz, 1829 Loxodactylus Chaudoir, 1865 Loxogenius Sloane, 1907 Loxomerus Chaudoir, 1842 Loxonchus Schmidt-Goebel, 1846 Loxoncus Schmidt-Goebel, 1846 Lucicolpodes J.Schmidt, 2000 Luoxiaotrechus Tian & Yin, 2013 Luperca Laporte, 1840 Luraphaenops Giordan, 1984 Lusotyphlus Luyatrechus M.Etonti & Mateu, 2000 Luzonotrechus Ueno, 1979 Lymnaem Lymnaeum Stephens, 1828 Lymnastis Motschulsky, 1862 Macfarlandia Sumlin, 1982 Machozetus Chaudoir, 1850 Macracanthus Chaudoir, 1846 Macrocheilus Hope, 1838 Macromorphus Chaudoir, 1857 Macroprotus Chaudoir, 1878 Maculagonum Darlington, 1952 Madagascareicheia Magrini & Bulirsch, 2009 Madapelmus Dajoz, 1985 Madascaris Bänninger, 1938 Madecassellus Deuve, 2010 Madecassina Jeannel, 1949 Magnanillus Baehr, 2017 Magnirabus Hong & Wang, 1990 Malagasydipnus Giachino, 2008 Malagasyreicheia Magrini & Bulirsch&Fancello, 2021 Malagasytyphlus Giachino, 2008 Mamboicus Bates, 1886 Mamesdytes Trezzi, 2003 Manautea Deuve, 2006 Manodromius Reitter, 1905 Mantica Kolbe, 1896 Mantichora Gistl, 1837 Manticora Fabricius, 1781 Manumorpha Erwin & Geraci, 2008 Maoriharpalus Larochelle & Larivière, 2005 Maoripamborus Brookes, 1944 Maoriplatynus Larochelle & Larivière, 2021 Maoritrechus Brookes, 1932 Marsyas Putzeys, 1845 Martyr Semenov-Tian-Shanskii & Znojko, 1929 Mascarenhia Alluaud, 1933 Masoreus Dejean, 1821 Mastax Fischer von Waldheim, 1828 Masuzoa Ueno, 1960 Masuzonoblemus Ueno, 1989 Matabele Péringuey, 1896 Mateuellus Deuve, 1990 Maungazolus Larochelle & Larivière, 2017 Mayaphaenops Vigna Taglianti, 1977 Mecodema Blanchard, 1843 Mecostomus Sharp, 1903 Mecyclothorax Sharp, 1903 Mecynognathus W.J.MacLeay, 1873 Mecynoscaris Alluaud, 1930 Medisores R.T. & J.R.Bell, 1987 Megacarabus Hong, 1983 Megacephala Latreille, 1802 Megadromus Motschulsky, 1866 Megadrypta Sciaky & Anichtchenko, 2020 Megalebia Mateu, 1972 Megalomma Westwood, 1841 Megaloodes Lesne, 1896 Megalopaussus Lea, 1906 Megamorio Chaudoir, 1880 Meganophthalmus Kurnakov, 1959 Megastylulus Giachino & Sciaky, 2003 Megodonthus Meileyingia Hong, 1987 Meladroma Motschulsky, 1855 Melaenus Dejean, 1831 Melanchiton Andrewes, 1940 Melanchrous Andrewes, 1940 Melanospilus Westwood, 1847 Meleagros Kirschenhofer, 1999 Melisodera Westwood, 1835 Menarus Andrewes, 1939 Menigius Chaudoir, 1879 Meonis Laporte, 1867 Meonochilus Liebherr & Marris, 2009 Meotachys Erwin, 1974 Merizodus Solier, 1849 Merizomena Chaudoir, 1873 Meroctenus Gemminger & Harold, 1868 Meropalpus Tschitscherine, 1900 Mesacanthina Rivalier, 1969 Mesarthropterus Wasmann, 1926 Mesocaraboides Deuve, 2012 Mesochila Rivalier, 1969 Mesocolpodes Basilewsky, 1985 Mesolestes Schatzmayr, 1943 Mesorabus Ponomarenko, 1977 Mesus Chevrolat, 1858 Metabacetus Bates, 1892 Metablus Jedlicka, 1958 Metacaraboides Deuve, 2012 Metacolpodes Jeannel, 1948 Metadromius Bedel, 1907 Metagrilium Handlirsch, 1906 Metalebia Basilewsky, 1948 Metallanchista Shi & Liang, 2013 Metallica Chaudoir, 1873 Metallina Motschulsky, 1850 Metascopus Basilewsky, 1970 Metaxellus Straneo, 1960 Metaxidius Chaudoir, 1852 Metaxinidium Basilewsky, 1963 Metaxymorphus Chaudoir, 1850 Metaxys Chaudoir, 1857 Metazuphium Mateu, 1992 Metius Curtis, 1839 Metoncidus Bates, 1871 Metriocheila J.Thomson, 1857 Metrius Eschscholtz, 1829 Metrocidium Sharp, 1903 Mexanillus Vigna Taglianti, 1973 Mexaphaenops Bolivar y Pieltain, 1942 Mexisphodrus Barr, 1965 Mexitrechus Barr, 1982 Mexyclothorax Mianningocarabus Imura, 2002 Miaotrechus Tian & Chen&Ma, 2020 Micratopus Casey, 1914 Micrixys LeConte, 1854 Microblemus Ueno, 2007 Microcephalus Berthold, 1827 Microcephalus Dejean, 1828 Microcheila Brullé, 1835 Microcosmodes Strand, 1936 Microdaccus Schaum, 1864 Microderes Faldermann, 1836 Microdipnites Jeannel, 1957 Microdipnodes Basilewsky, 1960 Microdipnus Jeannel, 1937 Microferonia Blackburn, 1890 Microlestes Schmidt-Goebel, 1846 Microlestodes Baehr, 1987 Micromentignatha Sumlin, 1982 Microodes Jeannel, 1949 Microschemus Andrewes, 1940 Microserrullula Netolitzky, 1921 Microthylax Rivalier, 1954 Microtyphlus Linder, 1863 Microus Chaudoir, 1876 Microzaena Fairmaire, 1901 Microzargus Sciaky & Facchini, 1997 Migadopidius Jeannel, 1938 Migadopiella Baehr, 2009 Migadops G.R.Waterhouse, 1842 Miguihuana Barr, 1982 Miltodes Andrewes, 1922 Mimanillus B.Moore, 1972 Mimocolliuris Liebke, 1933 Mimodromius Chaudoir, 1873 Mimophilorhizus Mateu, 1993 Mimotrechus B.Moore, 1972 Mimovelindopsis Mateu, 1963 Mimozaena Deuve, 2001 Minideutera Moravec, 2002 Minimaphaenops Deuve, 2000 Minipenetretus Zamotajlov, 2002 Minosaphaenops Queinnec, 2008 Minuphloeus Darlington, 1968 Minuthodes Andrewes, 1941 Minutotrechus Deuve & Kavanaugh, 2016 Minypatrobus Ueno, 1955 Miocarabus Hong, 1983 Miolebidia Zhang & Liu&Shangguan, 1989 Mioptachys Bates, 1882 Miquihuana Barr, 1982 Mirachlaenius Facchini, 2011 Miscelus Klug, 1834 Mischocephalus Chaudoir, 1863 Miscodera Eschscholtz, 1830 Mizotrechus Bates, 1872 Mnuphorus Chaudoir, 1873 Mochtherus Schmidt-Goebel, 1846 Moirainpa Erwin, 1984 Molopidius Jeannel, 1942 Molopinus Jeannel, 1948 Molops Bonelli, 1810 Molopsida White, 1846 Monnea Mateu, 1970 Monocentrum Chaudoir, 1868 Monodryxus Straneo, 1942 Monolobus Solier, 1849 Montagonum Darlington, 1952 Monteremita Seldon & Holwell, 2019 Mooreagonum Baehr, 2016 Morimotoides Morimotoidius Habu, 1954 Moriodema Laporte, 1867 Moriomorpha Laporte, 1867 Moriomorphina Morion Latreille, 1810 Morionidius Chaudoir, 1880 Moriosomus Motschulsky, 1855 Mormolyce Hagenbach, 1825 Mormolycina Jeannel, 1949 Morphodactyla Semenov, 1889 Motianocarabus Imura, 2007 Mouhotia Laporte, 1862 Myanmadera Wiesner & Phyu, 2015 Myas Sturm, 1826 Myriochila Motschulsky, 1858 Myriochile Motschoulsky, 1862 Myrmecodemus Sloane, 1923 Mystroceridius Reichardt, 1972 Mystropomus Chaudoir, 1848 Nannoryctes Baehr, 2000 Nannotrechus Winkler, 1926 Nanodiodes Bousquet, 1996 Nanotrechiama Belousov & Kabak, 2018 Naviauxella Cassola, 1988 Naxipenetretus Zamotajlov, 1999 Neanops Britton, 1962 Neaphaenops Jeannel, 1920 Nearctic Pachyotominae Nebria Latreille, 1802 Nebriagonum Darlington, 1952 Nebrioferonia Straneo, 1939 Nebriosoma Laporte, 1867 Nebrorabus Ponomarenko, 1989 Negrea Mateu, 1968 Negreum Habu, 1958 Nelidus Chaudoir, 1878 Nelsonites Valentine, 1952 Nemaglossa Solier, 1849 Nemotarsus LeConte, 1853 Neoaulacoryssus Noonan, 1985 Neobatenus Jeannel, 1948 Neoblemus Jeannel, 1923 Neocarenum Laporte, 1867 Neochila Basilewsky, 1953 Neochryopus Bänninger, 1932 Neocicindela Rivalier, 1963 Neocollyris W.Horn, 1901 Neocolpodes Jeannel, 1948 Neocoptodera Jeannel, 1949 Neodendragonum Basilewsky, 1953 Neodhysores R.T. & J.R.Bell, 1978 Neodiachipteryx Noonan, 1985 Neodipnus Jeannel, 1957 Neodrypta Basilewsky, 1960 Neodyschirius Kult, 1954 Neoeudalia Baehr, 2005 Neoferonia Britton, 1940 Neoglyptus Basilewsky, 1953 Neohelluo Sloane, 1914 Neohyparpalus Clarke, 1981 Neoillaphanus Giachino & Eberhard&Perina, 2021 Neolaphyra Bedel, 1895 Neoleleupidia Basilewsky, 1953 Neolovricia Lakota & Jalzic&J.Moravec, 2009 Neomauna Turgut & Özdikmen, 2006 Neomegalonychus Jeannel, 1948 Neonomius B.Moore, 1963 Neoodes Basilewsky, 1953 Neoorthogonius Tian & Deuve, 2006 Neophygas Noonan, 1976 Neoplectes Gottwald, 1982 Neoreicheia Kult, 1950 Neoscaphus Sloane, 1888 Neotalus Will, 2002 Neothanes Scudder, 1890 Neotrechus G.Müller, 1913 Nepalorthogonius Habu, 1979 Nepalorthorgonius Habu, 1979 Nesacinopus Tschitscherine, 1900 Nesaeoduvalius Casale & Giachino, 2016 Nesamblyops Jeannel, 1937 Nesarpalus Bedel, 1897 Nesarpax Alluaud, 1936 Nesiocolpodes Jeannel, 1948 Nesiodrypta Jeannel, 1949 Nesiotrechus Ueno, 1995 Nesites Andrewes, 1939 Nesocidium Sharp, 1903 Nesolimnaeum Sharp, 1901-1910 Nesomicrops Sharp, 1903 Nesorthomus Bedel, 1899 Nesosteropus Apfelbeck, 1904 Netrodera Chaudoir, 1850 Nickerlea W.Horn, 1899 Nilotrechus Quéinnec & Ollivier, 2021 Nipponagonum Habu, 1978 Nipponaphaenops Ueno, 1971 Nipponoharpalus Habu, 1973 Nippononebria Ueno, 1955 Nipponosynuchus Morita, 1998 Nirmala Andrewes, 1930 Nomius Laporte, 1835 Nornalupia Kataev, 2002 Notabax B.Moore, 1976 Notagonum Darlington, 1952 Notaphus Dejean, 1821 Nothanillus Jeannel, 1962 Nothobroscus Roig-Juñent & Ball, 1995 Nothocascellius Roig-Juñent, 1995 Nothodaptus Maindron, 1906 Nothoderis Boyd & Erwin, 2016 Nothoodes Gueorguiev & Liang, 2020 Notiobia Perty, 1830 Notiokasis Kavanaugh & Nègre, 1983 Notiophilus Duméril, 1805 Notocolpodes Basilewsky, 1985 Notokasis Kavanaugh & Nègre, 1983 Notokistus Handlirsch, 1906 Notolestus Sloane, 1894 Notonomus Chaudoir, 1865 Notoplatynus B.Moore, 1985 Notopseudomorpha Baehr, 1997 Notospeophonus B.Moore, 1962 Notospira Rivalier, 1961 Nototachys Alluaud, 1930 Nototarsus Nototarus de Chaudoir, 1875 Nototrechus B.Moore, 1972 Nototylus Gemminger & Harold, 1868 Novillidius Straneo, 1941 Nunbergites Pawlowski & Stachowiak, 1991 Nurus Motschulsky, 1866 Nycteis Laporte, 1835 Nyctosyles Putzeys, 1867 Nymphagonum Habu, 1978 Oarotrechus Townsend, 2010 Obadius Burmeister, 1875 Oberthueria Vuillet, 1911 Ochropisus Bates, 1883 Ochtozetus Chaudoir, 1872 Ochyropus Schiödte, 1847 Ocydromus Clairville, 1806 Ocys Stephens, 1828 Odacantha Paykull, 1798 Odontium LeConte, 1848 Odontocarus Solier, 1835 Odontocheila Laporte, 1834 Odontomasoreus Darlington, 1968 Odotoncarus Solier, 1835 Oechalius Liebke, 1935 Oecornis Britton, 1940 Oedesis Motschulsky, 1850 Oesyperus Andrewes, 1923 Ogmophora Tschitscherine, 1896 Ogygium Liebke, 1935 Oilea Liebke, 1931 Olegius Komarov, 1996 Olisthopus Dejean, 1828 Omalodera Blanchard, 1842 Omaseus Mannerheim, 1824 Omestes Andrewes, 1933 Omobrus Andrewes, 1930 Omoglymmius Ganglbauer, 1891 Omophagus Andrewes, 1937 Omophron Latreille, 1802 Omostropus Péringuey, 1896 Omphra Dejean, 1825 Omphra Reiche, 1842 Omphreoides Fairmaire, 1896 Omphreus Dejean, 1828 Omus Eschscholtz, 1829 Onawea Johns, 2007 Oncostylus Habu, 1978 Onota Chaudoir, 1873 Onotokiba Alluaud, 1927 Onycholabis Bates, 1873 Onypterygia Dejean, 1831 Oodes Bonelli, 1810 Oodinkosa Straneo, 1939 Oodinotrechus Ueno, 1998 Oodinus Motschulsky, 1865 Ooidius Chaudoir, 1847 Oopterus Guérin-Méneville, 1841 Oosoma Nietner, 1857 Ophionea Klug, 1821 Ophonichius Straneo, 1942 Ophoniscus Bates, 1892 Ophonus Dejean, 1821 Ophonus Stephens, 1828 Opilidia Rivalier, 1954 Opisthencentrus W.Horn, 1893 Opisthius Kirby, 1837 Opsis Handlirsch, 1906 Oregus Putzeys, 1868 Oreodicastes Maindron, 1905 Oreonebria K.Daniel, 1903 Oribazus Chaudoir, 1874 Orictites Andrewes, 1931 Orientoreicheia Bulirsch & Hurka, 1994 Orionella Jedlicka, 1964 Oroblemites Ueno & Pawlowski, 1981 Oroblemus Ueno & Yoshida, 1966 Orophicus Alluaud, 1925 Orotrechus G.Müller, 1913 Orthobasis Chaudoir, 1871 Orthocerodus Basilewsky, 1946 Orthocindela Rivalier, 1972 Orthogenium Chaudoir, 1835 Orthoglymma Liebherr, Marris, Emberson & Syrett&Roig-Juñent, 2011 Orthogonius W.S.MacLeay, 1825 Orthomus Chaudoir, 1838 Orthotrichus Peyron, 1856 Orthotyphlus Mateu & Zaballos, 1998 Orzolina Machado, 1987 Oscadytes Lagar Mascaro, 1975 Osimus Motschulsky, 1850 Otoglossa Chaudoir, 1873 Ovrabites Ponomarenko, 1977 Oxephloena Shpeley & Ball, 2000 Oxoides Solier, 1849 Oxycentrus Chaudoir, 1854 Oxycheila Dejean, 1825 Oxycheilopsis Cassola & Werner, 2004 Oxycrepis Reiche, 1843 Oxydrepanus Putzeys, 1867 Oxyglychus Straneo, 1938 Oxygnathopsis Louwerens, 1953 Oxygnathus Dejean, 1826 Oxygonia Mannerheim, 1837 Oxygoniola W.Horn, 1892 Oxygonium Basilewsky, 1951 Oxylobus Chaudoir, 1855 Oxyodontus Chaudoir, 1870 Oxypselaphus Chaudoir, 1843 Oxytrechus Jeannel, 1927 Ozaena Olivier, 1812 Ozaenapheaenops Deuve, 1986 Ozaeniella Basilewsky, 1976 Pachybatenus Basilewsky, 1973 Pachycaecus Straneo, 1971 Pachycallida Jeannel, 1949 Pachycarus Solier, 1835 Pachydesus Motschulsky, 1864 Pachyferonia Jeannel, 1951 Pachylebia Jeannel, 1949 Pachylebiodes Mateu, 1972 Pachymorphus Chaudoir, 1838 Pachyodontus Chaudoir, 1879 Pachyroxochus Straneo, 1942 Pachystoides Deuve, 2012 Pachyteles Perty, 1830 Pachythecus Chaudoir, 1874 Pachytrachelus Chaudoir, 1852 Pachytrechodes Jeannel, 1960 Palaeoaxinidium McKay, 1991 Palaeoiresina Wiesner & Will&J.Schmidt, 2017 Palaeopronyssiformia Wiesner & Will&J.Schmidt, 2017 Pamborus Latreille, 1812 Panagaeus Bonelli, 1810 Panagaeus Latreille, 1802 Panagrius Andrewes, 1933 Panaphaenops Tian & Huang&Ma, 2021 Pangaphaenops Tian, Huang & Jia, 2023 Pangus Dejean, 1821 Paniestichus Will, 2011 Papuadolichoctis Baehr, 1998 Parabaris Broun, 1881 Parabradycellus N.Ito, 2003 Parabroscus Lindroth, 1956 Parachlaenius Kolbe, 1894 Paracimmerites Belousov, 1998 Paracimmerities Belousov, 1998 Paraclivina Kult, 1947 Paracollagonum Baehr, 2001 Paracolliuris Kirschenhofer, 1994 Paracolpodes Basilewsky, 1985 Paracoryza Basilewsky, 1952 Paradolichoctis Baehr, 2006 Paradolichus Semenov, 1889 Paradromius Fowler, 1887 Paradyscherus Basilewsky, 1971 Paraeutrichopus Mateu, 1954 Paraglycia Bedel, 1904 Paraleleupidia Basilewsky, 1951 Paraliagonum Basilewsky, 1957 Parallelomorphus Motschulsky, 1850 Paralovricia Giachino & B.Gueorguiev&Vailati, 2011 Paramecus Dejean, 1829 Paramegalonychus Basilewsky, 1953 Paranchodemus Habu, 1978 Paranchus Lindroth, 1974 Paranillopsis Cicchino & Roig-Juñent, 2001 Paranillus Jeannel, 1949 Paranurus Tschitscherine, 1901 Parapatrobus Zamotajlov, 1992 Parapenetretus Kurnakov, 1960 Parapenthes Kataev & Wrase, 2018 Parapenthus Kataev & Wrase, 2018 Paraphaea Bates, 1873 Paraphaenops Jeannel, 1916 Paraphysodeutera J.Moravec, 2002 Parapionycha Liebke, 1929 Paraplatynus Baehr, 2016 Parapterostichus Desbrochers, 1906 Parareicheia Jeannel, 1957 Parascopodes Darlington, 1968 Parasiopelus Basilewsky, 1946 Parasyleter Balkenohl, 2021 Paratachys Casey, 1918 Parathlibops Basilewsky, 1958 Paratrechiotes Ueno, 1995 Paratrechodes Jeannel, 1926 Paratrechus Jeannel, 1920 Paratrichothorax Baehr, 2004 Paraulacoryssus Shpeley & Hunting&Ball, 2017 Paraxinidium Basilewsky, 1963 Parazuphium Jeannel, 1942 Paregaploa Müller, 1948 Paregraphus Basilewsky, 1967 Parena Motschulsky, 1860 Parepinebriola Kavanaugh, 2021 Pareuryaptus Dubault & Lassalle&Roux, 2008 Pareurydera Jeannel, 1949 Pareutrichopus Mateu, 1954 Parhypates Motschulsky, 1866 Paridiagonum Baehr, 2009 Parillaphanus Baehr, 2018 Parophonus Ganglbauer, 1891 Paropisthius Casey, 1920 Parorthomus B.Gueorguiev & Wrase&Farkac, 2014 Parvindela Duran & Gough, 2019 Parvocaecus Coiffait, 1956 Pasimachus Bonelli, 1813 Passalidius Chaudoir, 1863 Patanitretus Zamotajlov, 2002 Patrobus Dejean, 1821 Paulianites Jeannel, 1949 Paulianolebia Mateu, 1972 Paulianoscirtus Basilewsky, 1976 Paussoides Motschoulsky, 1857 Paussomorphus Raffray, 1885 Paussopsis Cockerell, 1911 Paussotropus C.O.Waterhouse, 1877 Paussus Linnaeus, 1775 Pawgammm Morvan, 1999 Pectinitarsus Fairmaire, 1881 Pediomorphus Chaudoir, 1878 Pedius Motschulsky, 1850 Pelecium Kirby, 1819 Peliocypas Schmidt-Goebel, 1846 Pelmatellus Bates, 1882 Pelocharis Jeannel, 1960 Pelodiaetodes B.Moore, 1980 Pelodiaetus Jeannel, 1937 Pelonomites Jeannel, 1963 Pelophila Dejean, 1821 Pelorus Fischer-Waldheim, 1821 Penecupes Ren, 1995 Penetretus Motschulsky, 1865 Pentacomia Bates, 1872 Pentagonica Schmidt-Goebel, 1846 Pentaplatarthrus Jeannel, 1946 Pentaplatarthrus Westwood, 1833 Penthophonus Reitter, 1900 Penthus Chaudoir, 1843 Pephrica Alluaud, 1936 Percodermus Sloane, 1920 Percolaus Bates, 1882 Percolestus Sloane, 1892 Percosoma Schaum, 1858 Percus Bonelli, 1810 Pericalina Hope, 1838 Pericalus W.S.MacLeay, 1825 Pericompsus LeConte, 1852 Peridexia Chaudoir, 1861 Perigona Castelnau, 1835 Perigona Laporte, 1835 Perigona Serrano, Capela & Santos, 2017 Perileptus Schaum, 1860 Periphobus Péringuey, 1896 Perissostomus Alluaud, 1930 Permunda Ponomarenko & Volkov, 2013 Perochnoristhus Basilewsky, 1973 Peronomerus Schaum, 1854 Perucharidius Mateu & M.Etonti, 2002 Peruphorticus Erwin & Zamorano, 2014 Petraphaenops Delic & Kapla&Colla, 2019 Petrimagnia Kryzhanovskij & Mikhailov, 1971 Petrimagonia Kryzhanovskihj & Michailov, 1971 Petrodromeus Ponomarenko & Volkov, 2013 Peyrieraselus Deuve, 1981 Phacocerus Chaudoir, 1873 Phacothorax Jeannel, 1944 Phaenaulax Tschitscherine, 1898 Phaeoxantha Chaudoir, 1850 Phalantha Gistl, 1839 Phanagnathus Basilewsky, 1950 Pharetis Liebherr, 2020 Pheggomisetes Globiceps Ciniglavcensis Pheggomisetes Knirsch, 1923 Pheropsophus Solier, 1833 Pherosophus Pherosopsophus Philipis Erwin, 1994 Philochthus Stephens, 1828 Philodes LeConte, 1861 Philophlaeus Chaudoir, 1844 Philophuga Motschulsky, 1859 Philorhizus Hope, 1838 Philoscaphus W.J.MacLeay, 1871 Phloeocarabus W.J.MacLeay, 1871 Phloeoxena Chaudoir, 1870 Phloeozetus Peyron, 1856 Pholeodytes Britton, 1962 Phorticosomus Schaum, 1863 Phrator Semenov, 1922 Phrypeus Casey, 1924 Phyla Motschulsky, 1844 Phyllodroma Lacordaire, 1843 Physea Brullé, 1835 Physeomorpha Ogueta, 1963 Physoclivina Kult, 1959 Physocrotaphus Parry, 1849 Physodera Eschscholtz, 1829 Physodeutera Lacordaire, 1843 Physoglossus Akhil & Sabu, 2020 Physolaesthus Chaudoir, 1850 Physomerus Chaudoir, 1883 Picnochile Motschulsky, 1856 Piesmus LeConte, 1846 Piezia Brullé, 1834 Pilades Heyne, 1895 Pilbaradytes Giachino & Eberhard&Perina, 2021 Pilbaranillus Baehr & Main, 2016 Pilbaraphanus Giachino & Eberhard&Perina, 2021 Pilosaphaenops Deuve & Tian, 2008 Pinacodera Schaum, 1857 Pionycha Chaudoir, 1848 Pioprosopus Tschitscherine, 1899 Piosoma LeConte, 1847 Plagiopyga Boheman, 1848 Plagiotelum Solier, 1849 Planetes W.S.MacLeay, 1825 Plataphus Motschulsky, 1864 Platidiolus Chaudoir, 1878 Platyagonum Habu, 1978 Platycerozaena Bänninger, 1927 Platychile W.S.MacLeay, 1825 Platycoelus Blanchard, 1843 Platyderus Stephens, 1827 Platyhelluo Baehr, 2005 Platylytron W.J.MacLeay, 1873 Platymetopsis Ball & Maddison, 1987 Platymetopus Dejean, 1829 Platynodes Westwood, 1847 Platynus Bonelli, 1810 Platypatrobus Darlington, 1938 Platyrhopalides Wasmann, 1918 Platyrhopalopsis Desneux, 1905 Platyrhopalus Westwood, 1833 Platysmodes Fauvel, 1903 Platysphyrus Sloane, 1905 Platytarus Fairmaire, 1850 Platyxythrius Lorenz, 1998 Platyxythrius Straneo, 1942 Plaumannium Liebke, 1939 Playtnus Plesioaphaenops Deuve & Tian, 2011 Plesioglymmius R.T. & J.R.Bell, 1978 Pleuarthropterus Wasmann, 1929 Pleuracanthus Gray, 1832 Plicagonum Darlington, 1952 Plocamostehus Britton, 1940 Plocamostethus Britton, 1940 Plochionus Dejean, 1821 Plochionus Wiedemann, 1823 Poecilinus Jeannel, 1948 Poecillus Samouelle, 1819 Poecilus Bonelli, 1810 Pogonistes Chaudoir, 1872 Pogonochila Pogonodaptus G.Horn, 1881 Pogonoglosssus Pogonoglossus Chaudoir, 1863 Pogonophorus Latreille, 1802 Pogonopsis Bedel, 1898 Pogonoschema Jeannel, 1927 Pogonostoma Klug, 1835 Pogonus Dejean, 1821 Polistichus Bonelli, 1810 Pollicobius Vinson, 1939 Polpochila Solier, 1849 Polyaulacus Chaudoir, 1878 Polychaetus Chaudoir, 1882 Polydamasium Liebke, 1938 Polyderidius Jeannel, 1962 Polyderis Motschulsky, 1862 Polyrhanis Rivalier, 1963 Pometon Fleutiaux, 1899 Ponapagonum Darlington, 1970 Pontodytes Casale & Giachino, 1989 Pontonoa Liebke, 1935 Porocara Sloane, 1917 Porocimmerites Belousov, 1998 Porotachys Netolitzky, 1914 Potamagonum Darlington, 1952 Praeanchodemus J.Schmidt & Göpel&Will, 2017 Praedodromeus Strada, Montagna & Tintori, 2014 Praepristus Kirschenhofer, 1999 Prakasha Andrewes, 1919 Princidium Motschulsky, 1864 Prioniomus Jeannel, 1937 Prionodrypta Jeannel, 1949 Prionognathus LaFerté-Sénectère, 1851 Pristacrus Chaudoir, 1870 Pristonychus Dejean, 1828 Pristoscelis Chaudoir, 1878 Pristosia Motschulsky, 1865 Probstia Cassola, 2002 Procarabus Oppenheim, 1888 Procasnoidea Baehr, 1996 Procechenochilus Imura & Belousov, 2002 Procerus Sturm, 1826 Procletodema Péringuey, 1898 Procletus Péringuey, 1896 Procollagonum Baehr, 2001 Procrustes Bonelli, 1810 Procrustoides Deuve, 2012 Prodiplous Zamotajlov & Sciaky, 2006 Proditomus Schauberger, 1934 Prodyscherodes Jeannel, 1955 Prodyscherus Jeannel, 1946 Progonochaetus G.Müller, 1938 Promecoderus Dejean, 1829 Promecognathus Chaudoir, 1846 Promecoptera Dejean, 1831 Promegalonychus Basilewsky, 1953 Pronyssa Bates, 1874 Pronyssiformia W.Horn, 1929 Proozaena Deuve, 2001 Prophenorites Basilewsky, 1985 Propionycha Liebke, 1928 Prosopogmus Chaudoir, 1865 Prosphodrus Britton, 1959 Prostalomus Basilewsky, 1950 Prostenognathus Shpeley & Ball, 2000 Prosynactus Bode, 1953 Prothyma Hope, 1838 Prothymidia Rivalier, 1957 Protocaccus Liebherr, 2000 Protocerapterus Wasmann, 1926 Protocolliuris Liebke, 1931 Protocollyris Mandl, 1975 Protocolpodes Basilewsky, 1985 Protomegodontus Imura, 2002 Protopaussus Gestro, 1892 Protopidius Basilewsky, 1949 Protorabus Ponomarenko, 1977 Protoscalidion L.Schaufuss, 1888 Protrechiama Belousov & Kabak, 2003 Psammocoryza Hogan, 2006 Psammodromius Peyerimhoff, 1927 Psecadius Alluaud, 1911 Pseudabacetus Burgeon, 1935 Pseudabarys Chaudoir, 1874 Pseudagonica B.Moore, 1960 Pseudamara Lindroth, 1968 Pseudamblytelus Baehr, 2004 Pseudanchomenus Tarnier, 1860 Pseudanillus Bedel, 1896 Pseudanisotarsus Noonan, 1973 Pseudanophthalmus Jeannel, 1920 Pseudaphaenops Winkler, 1912 Pseudaptinus Laporte, 1834 Pseudaristus Reitter, 1900 Pseudillaphanus Giachino, 2005 Pseudobatenus Basilewsky, 1951 Pseudoceneus Tschitscherine, 1891 Pseudoclivina Kult, 1947 Pseudocnides Jeannel, 1927 Pseudocollyris Bi & Wiesner, 2021 Pseudodiachipteryx Burgeon, 1936 Pseudodromica Cassola, 2003 Pseudognathaphanus Schauberger, 1932 Pseudohyparpalus Basilewsky, 1946 Pseudomasoreus Desbrochers des Loges, 1904 Pseudomegalonychus Basilewsky, 1950 Pseudometabletus Liebke, 1930 Pseudomigadops Jeannel, 1938 Pseudomonnea Mateu, 1983 Pseudomorpha Kirby, 1823 Pseudomyas Uyttenboogaart, 1929 Pseudonebria Ledoux & Roux, 1988 Pseudoodes Gueorguiev & Liang, 2020 Pseudoophonus Motschulsky, 1844 Pseudoopterus Csiki, 1928 Pseudopachylebia Mateu, 1972 Pseudophilochthus Wollaston, 1877 Pseudophorticus Erwin, 2004 Pseudoplatiella Baehr, 2012 Pseudoplatyderus Bolivar y Pieltain, 1940 Pseudorhysopus Kataev & Wrase, 2001 Pseudoselenophorus Péringuey, 1896 Pseudosphaerodes Jeannel, 1949 Pseudotaphoxenus L.Schaufuss, 1865 Pseudotetracha Fleutiaux, 1894 Pseudotoglossa Mateu, 1961 Pseudotrechisibus Mateu & Belles, 1982 Pseudotrechus Rosenhauer, 1856 Pseudoxycheila Guérin-Méneville, 1838 Pseudozaena Laporte, 1834 Psilidius Jeannel, 1957 Psychristus Andrewes, 1930 Psydrus LeConte, 1846 Pterocyrtus Sloane, 1920 Pterogmus Sloane, 1920 Pterorhopalus Maruyama, 2011 Pterostichus Bonelli, 1810 Pterostillichus Straneo, 1949 Puertrechus Belousov & Kabak, 2014 Punctoduvalius Curic & Curic, 2012 Putzeysius Jeannel, 1962 Pylartesius Liebke, 1939 Pyramoides Bousquet, 2002 Qianaphaenops Ueno, 2000 Qiangopatrobus Zamotajlov, 2002 Qianlongius Tian & Jia, 2021 Qianotrechus Ueno, 2000 Qinlingocarabus Imura, 1998 Quammenis Erwin, 2000 Quasicalathus Schmidt & Will, 2022 Quasipenetretus Zamotajlov, 2002 Queinnectrechus Deuve, 1992 Rakantrechus Ueno, 1951 Rambousekiella Knirsch, 1925 Raphetis B.Moore, 1963 Raptor Lucnik, 1927 Rascioduvalius Curcic, Brajkovic, Mitic & Curcic, 2003 Rathymus Dejean, 1831 Rawlinsius Davidson & Ball, 1998 Reflexisphodrus Casale, 1988 Reichea Saulcy, 1862 Reicheadella Reitter, 1913 Reicheia Saulcy, 1862 Reicheidius Jeannel, 1957 Reicheiodes Ganglbauer, 1891 Reicheodes Ganglbauer, 1892 Renneria Baehr, 1999 Rhabdotus Chaudoir, 1865 Rhabidius Basilewsky, 1948 Rhadine LeConte, 1846 Rhaebolestes Sloane, 1903 Rhagadillius Straneo, 1951 Rhanagnathus Basilewsky, 1950 Rhegmatobius Jeannel, 1937 Rhipocarabus Reitter, 1896 Rhopalochlaenius Zhang & Sun&Zhang, 1994 Rhopalomelus Boheman, 1848 Rhopalostyla Chaudoir, 1850 Rhysocara Sloane, 1916 Rhysodes Germar, 1822 Rhysopleura Sloane, 1906 Rhysopus Andrewes, 1929 Rhysotrachelus Boheman, 1848 Rhytidocarabus Imura, 2002 Rhytidognathus Chaudoir, 1861 Rhytidophaena Bates, 1891 Rhytiferonia Darlington, 1962 Rhytisternus Chaudoir, 1865 Rhyzodiastes Fairmaire, 1895 Ripogenites Basilewsky, 1954 Rivacindela Brouerius van Nidek, 1973 Robustopenetretus Zamotajlov & Sciaky, 1999 Ronhuberia J.Moravec & Kudrna, 2002 Ropaloteres Guerin-Meneville, 1849 Rossjoycea Liebherr, 2011 Rubidiclivina Baehr, 2015 Rugiluclivina Balkenohl, 1996 Rugitarus Baehr, 2009 Rupa Jedlicka, 1935 Ryugadous Habu, 1950 Salcedia Fairmaire, 1899 Salpingophora Rivalier, 1950 Samiriamorpha Erwin & Geraci, 2008 Sangocarabus Imura, 2002 Sardaphaenops Cerruti & Henrot, 1956 Sarothrocrepis Chaudoir, 1850 Sarticus Motschulsky, 1866 Satotrechus Ueno, 2006 Sbordoniella Vigna Taglianti, 1980 Scalidion Schmidt-Goebel, 1846 Scaphinotus Dejean, 1826 Scaphinotus Latreille Scapterus Dejean, 1826 Scaraphites Westwood, 1842 Scarites Fabricius, 1775 Scaurotrechodes Geginat, 2006 Schidonychus Klug, 1834 Schizogenius Putzeys, 1846 Schuelea Baehr, 2004 Scolyptus Putzeys, 1861 Scopodes Erichson, 1842 Scotagonum Habu, 1978 Scotodipnus Schaum, 1860 Scotoplanetes Absolon, 1913 Scototrechus Britton, 1962 Scutellarius Morvan, 2010 Scybalicus Schaum, 1862 Secatophus Laporte, 1867 Selenochilus Chaudoir, 1878 Selenodipnus Jeannel, 1963 Selenophori Selenophorus Dejean, 1829 Selenotichnus Kataev, 1999 Selina Motschulsky, 1858 Semiaphaenops Deuve, 2000 Semiardistomis Kult, 1950 Semiclivina Kult, 1947 Semiophonus Schauberger, 1933 Sericoda Kirby, 1837 Serranillus Barr, 1995 Serratotyphlus Giachino, 2008 Serratozaena Deuve, 2001 Serrimargo Chaudoir, 1870 Setalidius Chaudoir, 1878 Setalimorphus Sloane, 1895 Setalis Laporte, 1867 Setitachys Baehr, 2016 Setodyschirius Fedorenko, 1996 Setolebia Jedlicka, 1941 Sfitakantha Andrewes, 1919 Shanwangicarabus Hong, 1985 Shenaphaenops Ueno, 1999 Shennongotrechus Ueno, 2004 Shenoblemus Tian & Fang, 2020 Shenocoptolabrus Imura, Zhou & Su, 1999 Shibataia Habu, 1978 Shilinotrechus Ueno, 2003 Shiqianaphaenops Tian, 2016 Shuangheaphaenops Tian, 2017 Shuaphaenops Ueno, 1999 Shuocychropsis Imura, 2002 Shyrodes Grouvelle, 1903 Siagona Latreille, 1804 Siagonyx W.J.MacLeay, 1871 Sichuanotrechus Deuve, 2005 Sidublemus Tian & Yin, 2013 Sigenacarabus Kleinfeld, 2000 Sikelioreicheia Magrini & Fancello, 2019 Simodontus Chaudoir, 1843 Simous Chaudoir, 1882 Sinaphaenopoides Tian, Huang & Jia, 2023 Sinaphaenops Ueno & Wang, 1991 Sinechostictus Motschulsky, 1864 Sinesetosa Balkenohl, 1996 Singiliomimus Péringuey, 1896 Singilis Rambur, 1837 Singillis Rambur, 1837-1838 Sinis Heer, 1862 Sinocalosoma Hong & Wang, 1986 Sinocarabus Hong, 1982 Sinocaralosoma Hong, 1984 Sinocimmerites Deuve, 2007 Sinocolpodes J.Schmidt, 2001 Sinocychropsis Imura, 2001 Sinoleptocarabus Imura, 2002 Sinometrius Wrase & J.Schmidt, 2006 Sinotrechiama Ueno, 2000 Sinotroglodytes Deuve, 1996 Sinozolus Deuve, 1997 Sinurus Chaudoir, 1870 Siopelophonus Kataev, 2015 Siopelus Murray, 1859 Sirdenus Dejean, 1828 Sitaphe B.Moore, 1963 Skoeda Morvan, 1995 Skorlagad Morvan, 1999 Skouedhirraad Morvan, 1999 Sloaneana Csiki, 1933 Sloanella Jeannel, 1927 Sloanoglymmius R.T. & J.R.Bell, 1991 Smeringocera Chaudoir, 1863 Smirnowia Lutshnik, 1922 Socotrana Cassola & Wranik, 1998 Soesilascarites Makhan, 2010 Sofota Jedlicka, 1951 Solenogenys Westwood, 1859 Somalodromius Mateu, 1967 Somoplatodes Basilewsky, 1986 Somoplatus Dejean, 1829 Somotrichus Seidlitz, 1887 Sophroferonia Alluaud, 1933 Spallomorpha Sparostes Putzeys, 1867 Speagonum B.Moore, 1977 Spelaeodytes L.Miller, 1863 Spelaeovulcania Machado, 1987 Speleodesmoides Mateu, 1978 Speleopidius Vives & Comas, 2021 Speluncarius Reitter, 1886 Speocolpodes Barr, 1974 Speokokosia Alluaud, 1932 Speomolops Patrizi, 1955 Speosuphium Moore, 1995 Speotarus B.Moore, 1964 Speothalpius B.Moore, 1995 Speotrechus Jeannel, 1922 Speozuphium B.Moore, 1995 Sperkanhir Morvan, 2010 Sphaeroderus Dejean, 1826 Sphaerodinus Jeannel, 1949 Sphaerostylus Chaudoir, 1848 Sphaerotachys G.Müller, 1926 Sphallomorpha Westwood, 1840 Spherita Liebherr, 2020 Sphodropsis Reitter, 1894 Sphodropsis Seidlitz, 1887 Sphodroschema Alluaud, 1930 Sphodrosomus Perroud, 1864 Sphodrus Clairville, 1806 Sphodrus Schellenberg, 1806 Sphoerodes Chaudoir, 1883 Sphrodus Spinidolichoctis Baehr, 1998 Sporades Fauvel, 1882 Srimara R.T. & J.R.Bell, 1978 Steganomma W.J.MacLeay, 1887 Stegazopteryx Will, 2004 Stenidia Brullé, 1834 Stenocheila Laporte, 1832 Stenochoromus L.Miller, 1866 Stenocnemus Mannerheim, 1837 Stenocosmia Rivalier, 1965 Stenocrepis Chaudoir, 1857 Stenognathus Chaudoir, 1843 Stenolepta Semenov, 1889 Stenolophidius Jeannel, 1948 Stenolophus Dejean, 1821 Stenomorphus Dejean, 1831 Stenoodes Basilewsky, 1953 Stenotelus Bouchard, 1903 Stereocerus Kirby, 1837 Stereostoma Murray, 1857 Steropus Mannerheim, 1824 Stevensius Jeannel, 1923 Sthenocranion Tschitscherine, 1899 Stichonotus Sloane, 1910 Stigmatachys Boyd & Erwin, 2016 Stigomerodes Straneo, 1939 Stilboma Andrewes, 1933 Stolonis Motschulsky, 1866 Stomis Clairville, 1806 Stomis Schellenberg, 1806 Stomonaxellus Tschitscherine, 1901 Storthodontus Chaudoir, 1855 Straneoa Basilewsky, 1953 Straneoites Basilewsky, 1947 Straneomelus Sciaky & Facchini, 2019 Straneostichus Sciaky, 1994 Straneotia Mateu, 1961 Stratiotes Putzeys, 1846 Stricklandiana Bousquet, 2002 Stricteripus Straneo & Ball, 1989 Striganoviella Fedorenko, 2012 Strigia Brullé, 1835 Strigomerodes Straneo, 1939 Strigomerus Chaudoir, 1873 Stuhlmannium Kolbe, 1894 Stygiotrechus Ueno, 1958 Stylulus L.Schaufuss, 1882 Styphlodromus Basilewsky, 1959 Styphlomerus Chaudoir, 1875 Styracoderus Chaudoir, 1874 Subilsia Español, 1967 Succinarthropterus Kolbe, 1926 Sugimotoa Habu, 1975 Sulciclivina Balkenohl, 2022 Sumatrechus Deuve, 2005 Sumlinia Cassola & Werner, 2001 Sundaoodes Gueorguiev & Liang, 2020 Sunocarabus Hong, 1987 Superbotrechus Deuve & Tian, 2009 Suzuka Ueno, 1956 Syleter Andrewes, 1941 Syletor Tschitscherine, 1899 Syllectus Bates, 1878 Synteratus Broun, 1909 Syntomus Hope, 1838 Synuchidius Apfelbeck, 1908 Synuchus Gyllenhal, 1810 Syrdenoidius Baehr & Hudson, 2001 Syrdenus Dejean, 1828 Systolocranius Chaudoir, 1857 Tachycellus Ganglbauer, 1891 Tachyina Motschulsky, 1862 Tachypeles Deuve, 2001 Tachyphanes Jeannel, 1946 Tachypus Stephens, 1828 Tachys Dejean, 1821 Tachysbembix Erwin, 2004 Tachyta Kirby, 1837 Tachyura Motschulsky, 1862 Tachyuropsis Shilenkov, 2002 Tachyxysta Boyd & Erwin, 2016 Taenarthrus Broun, 1914 Taiwanotrechus Ueno, 1987 Takasagoagonum Habu, 1977 Tangarona R.T. & J.R.Bell, 1982 Taniatrechus Belousov & Dolzhanski, 1994 Tantillus Chaudoir, 1870 Tanystoma Motschulsky, 1845 Tanythrix Schaum, 1858 Taphoxenus Motschulsky, 1850 Taphoxomimus Straneo, 1975 Tapinopterus Schaum, 1858 Tapinoscaris Jeannel, 1946 Tarastethus Sharp, 1883 Taridius Chaudoir, 1875 Tarsagonum Darlington, 1952 Tarsitachys Erwin, 1971 Tasmanillus Giachino, 2005 Tasmanitachoides Erwin, 1972 Tasmanorites Jeannel, 1927 Tasmanotrechus B.Moore, 1972 Tauredon Handlirsch, 1910 Taurocimmerites Belousov, 1998 Tecnophilus Chaudoir, 1877 Tefflus Latreille, 1822 Tefflus Leach, 1819 Teiresia Liebke, 1935 Temnostega Enderlein, 1905 Tenuistilus Habu, 1978 Teraphis Laporte, 1867 Teratexis Semenov-Tian-Shanskii & Znoiko, 1933 Teratotarsa Tschitscherine, 1893 Termophilum Basilewsky Teropha Laporte, 1867 Tetracha Hope, 1838 Tetragonoderus Dejean, 1829 Tetraleucus Casey, 1920 Tetungonebria Shilenkov, 1982 Teukrus Liebke, 1931 Thaioodes Gueorguiev, 2014 Thalassobius Solier, 1849 Thalassoduvalius Ueno, 1956 Thalassophilus Wollaston, 1854 Thalassotrechus Van Dyke, 1918 Thaumastaphaenops Magrini, Vanni & Zanon, 1997 Thayerella Baehr, 2016 Theprisa B.Moore, 1963 Therates Latreille, 1816 Thermoscelis Putzeys, 1873 Thliboclivina Kult, 1959 Thlibops Putzeys, 1867 Thoasia Liebke, 1939 Thopeutica Schaum, 1861 Thryptocerus Chaudoir, 1878 Thyreochaetus Basilewsky, 1959 Thyreopterus Dejean, 1831 Thysanotus Chaudoir, 1848 Tianeotrechus Tian & Tang, 2016 Tianzhuaphaenops Zhao & Tian, 2016 Tibetopenetretus Zamotajlov & Sciaky, 1999 Tibetoplataphus Schmidt, 2003 Tibetorinocarabus Imura, 2002 Tibioscarites Bänninger, 1929 Tienmutrechus Suenson, 1957 Tiferonia Darlington, 1962 Tilius Chaudoir, 1876 Tinautius Mateu, 1997 Tinoderus Chaudoir, 1879 Tinognathus Chaudoir, 1879 Titaresius Liebke, 1935 Tonkinaphaenops Deuve, 2013 Tonkinoscaris Bänninger, 1956 Torretassoa Schatzmayr & Koch, 1933 Toshiaphaenops Ueno, 1999 Tostkar Morvan, 1998 Trachelocyphoides Alluaud, 1934 Trachelocyphoides Straneo, 1942 Trachelocyphus Tschitscherine, 1900 Trachysarus Reed, 1874 Trachyta Trechepaphiama Deuve & Kavanaugh, 2016 Trechepaphiopsis Deuve & Kavanaugh, 2016 Trechiama Jeannel, 1927 Trechiamiotes Deuve, 1998 Trechicomimus Mateu & Nègre, 1972 Trechicus LeConte, 1853 Trechiella Jeannel, 1927 Trechimorphus Jeannel, 1927 Trechinites Heer, 1862 Trechinotus Jeannel, 1962 Trechiotes Jeannel, 1954 Trechisibus Motschulsky, 1862 Trechistus B.Moore, 1972 Trechobembix Jeannel, 1926 Trechoblemus Ganglbauer, 1891 Trechodes Blackburn, 1901 Trechoides Motschulsky, 1856 Trechosia Jeannel, 1926 Trechosiella Jeannel, 1960 Trechus Clairville, 1806 Tremembecarabus Martins-Neto, 1998 Trepanes Motschulsky, 1864 Trephionus Bates, 1883 Trephisa B.Moore, 1963 Triaenogenius Chaudoir, 1877 Tribacoides Deuve, 2012 Trichamblytelus Baehr, 2004 Trichaphaenops Jeannel, 1916 Tricharnhemia Baehr, 2009 Trichidema Basilewsky, 1956 Trichillinus Straneo, 1938 Trichis Klug, 1832 Trichisia Motschulsky, 1865 Trichocarenum Blackburn, 1892 Trichocoptodera Louwerens, 1958 Trichognatha Latreille, 1829 Trichognathus Latrielle, 1829 Tricholicinus Poppius, 1912 Trichopalpoodes B.Gueorguiev & J.Schmidt, 2016 Trichopedius Bedel, 1899 Trichopiezia Nègre, 1955 Trichopselaphus Chaudoir, 1843 Trichopsida Larochelle & Larivière, 2013 Trichoputzeysius Sciaky & Grottolo, 1996 Trichosternus Chaudoir, 1865 Trichotichnus A.Morawitz, 1863 Trichoxycentrus N.Ito, 2000 Tricondyla Latreille, 1822 Trigonaptus Fedorenko, 2020 Trigonothops W.J.MacLeay, 1864 Trigonotoma Dejean, 1828 Trilophidius Jeannel, 1957 Trilophus Andrewes, 1927 Triplosarus Bates, 1874 Trirammatus Chaudoir, 1835 Trogloagonum Casale, 1982 Troglocimmerites Ljovuschkin, 1970 Trogloclivina Deuve, 2003 Troglorites Jeannel, 1919 Tropidocerus Chaudoir, 1878 Tropidotrechus Jeannel, 1927 Tropopsis Solier, 1849 Tropopterus Solier, 1849 Trymosternus Chaudoir, 1873 Tschitscherinellus Csiki, 1906 Tshitsherinella Semenov, 1905 Tuiharpalus Larochelle & Larivière, 2005 Tuiplatynus Larochelle & Larivière, 2021 Tuxtlamorpha Erwin & Geraci, 2008 Typhlocharis Dieck, 1869 Typhlochoromus Moczarski, 1913 Typhlonesiotes Jeannel, 1937 Typhloreicheia Holdhaus, 1924 Typhloreichia Holdhaus, 1924 Typhloscaris Kuntzen, 1914 Typhlotrechus G.Müller, 1913 Typhlozuphium Baehr, 2014 Typsiharpalus Tschitscherine, 1901 Uenanthracus Kasahara, 1994 Uenoaphaenops Tian & He, 2020 Uenoites Belousov & Kabak, 2016 Uenotrechus Deuve & Tian, 1999 Ushijimaella Ueno, 1980 Vachinius Casale, 1984 Valeriaaschero Erwin, 2004 Vata Fauvel, 1882 Velebitaphaenops Casale & Jalzic, 2012 Velesaphaenops S.Curcic et al., 2018 Velinda Andrewes, 1921 Velindastus Schüle & Lorenz, 2008 Velindomimus Jeannel, 1955 Velindopsis Burgeon, 1937 Vianasia Mateu, 1955 Vietotrechus Ueno, 1995 Viridagonum Lassalle, 2015 Vitagonum B.Moore, 1999 Vulcanophilus Heller, 1898 Wahlbergiana Bousquet, 2002 Waiputrechus Townsend, 2010 Walterhornia Olsoufieff, 1934 Waltherhornia Olsoufieff, 1934 万华盲步甲属 Wanhuaphaenops Tian & Wang, 2016 Wanoblemus Tian & Fang, 2016 Watanabeocarabus Imura, 2002 Wate Liebke, 1928 Whiteheadiana Perrault, 1994 Winklerites Jeannel, 1937 Wolltinerfia Machado, 1985 Wuchangicarabus Hong, 1991 Wulongius Tian & Huang, 2021 Wulongoblemus Ueno, 2007 Wumengius Tian, Huang & Jia, 2023 Xanthophoea Xanthos Kirschenhofer, 2003 Xenaroswelliana Erwin, 2007 Xenion Tschitscherine, 1902 Xenitenopsis Basilewsky, 1956 Xenitenus Péringuey, 1896 Xenodochus Andrewes, 1941 Xenodromius Bates, 1891 Xenophonus Müller, 1942 Xenotrechus Barr & Krekeler, 1967 Xestagonum Habu, 1978 Xestonotus LeConte, 1853 Xestopus Andrewes, 1937 Xhosores R.T. & J.R.Bell, 1978 Xiangxius Tian & Huang&Li, 2021 Xishanocarabus Hong, 1984 Xoloncus Kataev, 2002 Xuedytes Tian & Huang, 2017 Xystosomus Schaum, 1863 Xystosormis Yalongaphaenops Belousov & Kabak, 2021 Yamatosa R.T. & J.R.Bell, 1979 Yamautidius Ueno, 1957 Yanzaphaenops Ueno, 2010 Yasunimorpha Erwin & Geraci, 2008 Yezacoptolabrus Imura, 2002 Youtrechus Tian, Huang & Jia, 2023 Yunnanocarabus Linnaeus, 1977 Yunotrechus Tian & Huang, 2014 Zabronothus Broun, 1893 Zabrus Clairville, 1806 Zabrus Schellenberg, 1806 Zacotus LeConte, 1869 Zangoharpalus Huang, 1998 Zapotecanillus Sokolov, 2013 Zargus Wollaston, 1854 Zariquieya Jeannel, 1924 Zeanillus Jeannel, 1937 Zecicindela Larochelle & Larivière, 2013 Zecillenus Lindroth, 1980 Zemetallina Lindroth, 1976 Zeodera Laporte, 1867 Zeopoecilus Sharp, 1886 Zheduocarabus Imura, 2002 Zhijinaphaenops Ueno & Ran, 2002 Zhongdianocarabus Imura, 2002 Zoianillus Sciaky, 1994 Zolanillus Sciaky, 1994 Zolotarevskyella Mateu, 1953 Zolus Sharp, 1886 Zunotrechus Tian, Huang & Jia, 2023 Zuphioides Ball & Shpeley, 2013 Zuphium Latreille, 1805 Carabidae Taxa. 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