金龜子

金龜子是金龜子科昆蟲的總稱，全世界有超過30,000種。可以在除了南極洲以外的大陸發現。不同的種類生活於不同的環境，如沙漠、農地、森林和草地等。 種類繁多，像是常見的台灣青銅金龜、廣為人知的獨角仙和台灣最大的甲蟲台灣長臂金龜也都屬於金龜子科的家族成員。 幼蟲稱為蠐螬，多生活於土中，以土中有機物為食。金龜子的觸角呈鰓葉狀，鎚節的部份常呈多分叉狀。成蟲食性各異，有的以植物各部分（根、莖、葉、花、果实、种子）或汁液為食，有的以腐敗有機物為食，也有以糞便為食。 本科包括以下属： Ablabera Dejean, 1833 Ablaberoides Blanchard, 1850 Acanthaphodius Schmidt, 1909 Acanthaphodius bruchi Schmidt, 1909 Acanthobodilus Dellacasa, 1983 Acanthobodilus immundus (Creutzer, 1799) Acanthobodilus languidulus (Schmidt, 1917) Acanthonitis Janssens, 1937 Acanthoprotaetia Miksic, 1987 Acanthovalgus Kraatz, 1895 Accrosus Aceratocrates Ohaus, 1910 Aceratus Prell, 1936 Acheilo Britton, 1988 Achelyna Erichson, 1848 Achloa Erichson, 1840 Achranoxia Kraatz, 1888 Aclopus Erichson, 1835 Acoma Casey, 1889 Acraspedon Arrow, 1899 Acrobolbia Ohaus, 1912 Acrossidius Schmidt, 1913 Acrossoides Schmidt, 1913 Acrossus Mulsant, 1842 Acrothyrea Kraatz, 1882 Actinobolus Westwood, 1841 Acylochiloides Martinez, 1975 Acylochilus Ohaus, 1909 Adebrattia Bordat & Howden, 1995 Adelaeus Ratcliffe, 2014 Adeloparius Schmidt, 1913 Adonides Thomson, 1880 Adoretops Kraatz, 1883 Adoretosoma Blanchard, 1850 Adoretus Dejean, 1833 Adoretus Laporte de Castelnau, 1840 Adorodocia Brenske, 1893 Adoroleptus Brenske, 1893 Adorrhinyptia Arrow, 1917 Adoryphorus Blackburn, 1889 Adossa Britton, 1995 Adraria Villiers, 1956 Aegialatis Kono, 1938 Aegialia Latreille, 1807 Aegidiellus Paulian, 1984 Aegidinus Arrow, 1904 Aegidium Westwood, 1845 Aegopsis Burmeister, 1847 Aegostheta Dejean, 1833 Aequatoria Arrow, 1899 Aethiaratrogus Decelle, 1982 Aethiartrogus Decelle, 1982 Aethiessa Burmeister, 1842 Afroataenius Petrovitz, 1969 Afrocloeotus Afrodiapterna Dellacasa, 1984 Afrodiastictus Pittino & Mariani, 1986 Afrodiasticus Pittino & Mariani, 1986 Afrodrepanus Krikken, 2009 Afroemadiellus Bordat, 2019 Afrolepis Decelle, 1968 Afrotrichonotulus Bordat, 2016 Agamopus Bates, 1887 Aganocrossus Reitter, 1895 Agaocephala Le Peletier & Serville, 1828 Agaocnemis Moser, 1918 Agestrata Eschscholtz, 1829 Aglaphyra Brenske, 1896 Agnorimus Miyake & Iwase, 1991 Agoliinus Schmidt, 1913 Agolius Mulsant & Rey, 1869 Agrilinellus Dellacasa, Dellacasa & Gordon, 2008 Agrilinus Mulsant & Rey, 1869 Ahermodontus Báguena, 1930 Ahermodontus Corella, 1930 Aidophus (Balthasar, 1963) Airapus Stebnicka & Howden, 1996 Ajmeraphodius Král, Mencl & Rakovič, 2021 Alaia Petrovitz, 1980 Alcidiosoma Alepida Allsopp, 2018 Aliaclitopa Lacroix, 2003 Alissonotum Arrow, 1908 Aliuscanthoniola Deschodt & Scholtz, 2008 Allara Britton, 1955 Alleucosma Schenkling, 1921 Alleurostictus Allidiostoma Arrow, 1940 Alloblackburneus Bordat, 2009 Allobodilus Petrovitz, 1963 Allogymnopleurus Janssens, 1940 Allokotarsa Péringuey, 1904 Allomyrina Arrow, 1911 Allonitis Janssens, 1936 Allophileurinus Dupuis & Dechambre, 2001 Allophileurus Allophyllus Fauvel, 1903 Allorrhina Burmeister, 1842 Alloscelus Boucomont, 1923 Allothnonius Britton, 1978 Alocoderus Schmidt, 1913 Alogistotarsa Péringuey, 1904 Altonthophagus Kabakov, 1990 Alvarengius Frey, 1975 Alvarinus Blanchard, 1850 Amadotrogus Reitter, 1902 Amaecylius Huchet, 2000 Amalus Mulsant & Rey, 1870 Amaurina Kolbe, 1895 Amaurodes Westwood, 1844 Amauroleucocelis Bourgoin, 1913 Amazula Kraatz, 1882 Amblochilus Blanchard, 1850 Amblonoxia Reitter, 1902 Amblymelanoplia Dombrow, 2002 Amblyodus Westwood, 1878 Amblyoproctus Kolbe, 1910 Amblyterus MacLeay, 1819 Amerilochus Skelley, 2007 Amerisaprus Stebnicka & Skelley, 2003 Amidorus Mulsant, 1870 Amietina Cambefort, 1981 Amiserica Nomura, 1974 Amithao Thomson, 1878 Ammaecius Ammoecioides Bordat, 1999 Ammoecius Mulsant, 1842 Amorphochelus Fairmaire, 1898 Amphibolonarzon Gistl, 1848 Amphiceratodon Huchet, 2000 Amphimallon Berthold, 1827 Amphimallon Latreille, 1825 Amphionthophagus Martin Piera & Zunino, 1983 Amphistomus Lansberge, 1874 Amphistous Lansberge, 1871 Amphitrichia Brenske, 1894 Ampliodactylus Smith, 2008 Ampotis Dechambre & Madge, 1979 Amuarina Anacamphtorrhina Blanchard, 1842 Anacamptorrhina Blanchard, 1842 Anacanthodes Britton, 1990 Anaetius Petrovitz, 1969 Anagnathocera Arrow, 1922 Anahi Martinez, 1958 Analleucosma Antoine, 1989 Anamatochoranus Lacroix, 1995 Anartioschiza Kolbe, 1894 Anaspilus Kolbe, 1892 Anatista Brême, 1844 Anatona Burmeister, 1842 Anatonochilus Péringuey, 1907 Anatrinodia Casey, 1915 Ancistroma Curtis, 1835 Ancistrosoma Curtis, 1835 Ancognatha Erichson, 1847 Ancylonyx Britton, 1957 Anelaphinis Kolbe, 1892 Anenaria Lacroix, 1993 Anepsiovalgus Kolbe, 1904 Aneucomides Blackburn, 1898 Aneustypus Anisocanthon Martinez & Pereira, 1956 Anisocheirus Guérin-Méneville, 1844 Anisochelus Burmeister, 1844 Anisonyx Latreille, 1807 Anisopholis Moser, 1913 Anisoplia Dejean, 1821 Anisoplia Fischer Von Waldheim, 1824 Anisoplia Schönherr, 1817 Anisorrhina Westwood, 1842 Ankaratrotrox Paulian, 1954 Annegialia Howden, 1971 Anochilia Burmeister, 1842 Anocoela Moser, 1914 Anoctus Sharp, 1875 Anodontopopillia Ohaus, 1901 Anomacaulus Fairmaire, 1878 Anomala Leach, 1819 Anomala Samouelle, 1819 Anomalacra Casey, 1915 Anomalites Fric, 1885 Anomalochela Moser, 1913 Anomalomorpha Arrow, 1908 Anomalophylla Reitter, 1887 Anomalorhina Jameson, Paucar-Cabrera & Solis, 2003 Anomalurobius Paulian, 1952 Anomiopsoides Blackwelder, 1944 Anomiopus Westwood, 1842 Anomioserica Arrow, 1946 Anomius Mulsant & Rey, 1869 Anomocaulus Fairmaire, 1881 Anomolyna Fairmaire, 1897 Anomonyx Saylor, 1940 Anonetus Erichson, 1847 Anonychonitis Janssens, 1950 Anoplanomala Arrow, 1917 Anoplocheilus MacLeay, 1838 Anoplodrepanus Simonis, 1981 Anoplognatho Rivers, 1889 Anoplognathus Leach, 1815 Anoplogomphus Prell, 1912 Anoplostethus Brullé, 1837 Anoplostethus Reiche, 1839 Anopsoistes Anoronotum Arrow, 1939 Anoxia Laporte, 1832 Anoxoides Petrovitz, 1971 Antandroyanus Lacroix, 1993 Antecessorirhyparus Minkina, 2020 Antemnacrassa Gomez Pallerola, 1979 Anthoplia Medvedev, 1949 Anthotocus Britton, 1957 Anthracophora Burmeister, 1842 Anthracophorides Moser, 1918 Anthrencya Lacroix, 1993 Anticheira Eschscholtz, 1818 Anticheiroides Soula, 1998 Antitrochalus Brenske, 1900 Antitrogus Burmeister, 1855 Antodon Brême, 1844 Apanesthes Kraatz, 1880 Aparammoecius Petrovitz, 1958 Apeltastes Howden, 1968 Aphaenoserica Brenske, 1900 Aphanesia Britton, 1990 Aphanesthes Kraatz, 1880 Aphelinis Antoine, 1987 Aphelorhina Aphengium Harold, 1868 Aphengoecus Péringuey, 1901 Aphodaulacus Koshantschikov, 1911 Aphodiellus Schmidt, 1913 Aphodiopsis Paulian, 1942 Aphodius Hellwig, 1798 Aphodius Illiger, 1798 Aphodobius Péringuey, 1901 Aphodoharmogaster Endrödi, 1983 Aphodopsammobius Endrödi, 1964 Aphodus Aphonides Rivers, 1889 Aphonodelus Kolbe, 1905 Aphonoprocus Aphonus LeConte, 1856 Aphotaenius Cartwright, 1952 Apicencya Lacroix, 1993 Aplidia Hope, 1837 Aplopsis Blanchard, 1850 Apocnosis Thomson, 1878 Apocnosoides Antoine, 2001 Apogonia Kirby, 1818 Apohdius Aporecolpa Lansberge, 1886 Aposchiza Brenske, 1903 Apotolamprus Olsoufieff, 1947 Apotriodonta Baraud, 1962 Apsteiniella Schmidt, 1916 Aptenocanthon Matthews, 1974 Apterepilissus Montreuil, 2011 Apteroathlia Smith & Evans, 2018 Apterocanthon Apterodemidea Gutiérrez, 1952 Aptychonitis Janssens, 1937 Araboplia Uliana & Sabatinelli, 2017 Arachnimorpha Kirby, 1827 Arachnodes Westwood, 1842 Archedinus Morón & Krikken, 1990 Archeohomaloplia Nikolajev, 1982 Archeopsammoporus Minkina, 2020 Archocamenta Brenske, 1895 Archophanes Kolbe, 1905 Archophileurus Kolbe, 1910 Archoserica Brenske, 1897 Arctodium Burmeister, 1844 Ardella Paulsen, 2021 Areoda MacLeay, 1819 Argentophileurus Penco & Zubarán, 2013 Argeremazus Stebnicka & Dellacasa, 2003 Argoplia Dombrow, 2003 Argyripa Thomson, 1878 Argyripia Argyrophegges Kraatz, 1895 Arielina Rossi, 1958 Arraphytarsa Péringuey, 1904 Arriguttia Martinez, 1960 Arrowiellus Balthasar, 1933 Arupaia Stebnicka, 1999 Asactopholis Brenske, 1894 Asaphomorpha Brancsik, 1893 Aschnarhyparus Makhan, 2006 Asiactenius Nikolajev, 2000 Asiopertha (Medvedev, 1949) Asophrops Petrovitz, 1961 Aspidolea Bates, 1888 Aspilochilus Rojkoff, 2013 Aspilus Westwood, 1874 Asserador Maunders, 1852 Astaborus Thomson, 1859 Astaena Erichson, 1847 Astaenomochus Astaenoplia Martinez, 1957 Astenorella Astenorhella Westwood, 1873 Asthenopholis Brenske, 1898 Asthenorhella Westwood, 1874 Asthenorhina Westwood, 1843 Astibicola Britton, 1990 Astoxenus Péringuey, 1907 Astrocara Astroscara Schürhoff, 1937 Ataeniopsis Petrovitz, 1973 Ataenius Harold, 1867 Atanyproctus Petrovitz, 1954 Ateuchetus Bedel, 1892 Ateuchini Perty, 1830 Ateuchites Meunier, 1898 Ateuchus Weber, 1801 Athesphatoplia Lacroix, 1998 Athlia Erichson, 1835 Atlantemolanum González-Alvarado, Molano-Rendón & Vaz-de-Mello, 2019 Atrichelaphinis Kraatz, 1898 Atrichiana Distant, 1911 Attavicinus Philips & Bell, 2008 Atysilla Strand, 1942 Augoderia Burmeister, 1847 Augosoma Burmeister, 1846 Aulacopalpus Guérin-Méneville, 1838 Aulacopris White, 1859 Aulacoschiza Decelle, 1973 Aulacoserica Brenske, 1900 Aulonocnemis Klug, 1838 Aurelia Thomson, 1880 Aurum Hutchinson & Moeseneder, 2019 Ausmontins Deschodt & Davis, 2018 Australagolius Mencl, 2008 Australammoecius Petrovitz, 1958 Australaphodius (Balthasar, 1942) Australommoecius Australoxenella Howden & Storey, 1992 Automolius Britton, 1978 Autoprotaetia Mikšic, 1965 Autoserica Brenske, 1897 Axillonia Krikken, 2018 Bacchusia Mikšić, 1976 Badelina Thomson, 1880 Badiasis Machatschke, 1970 Badizoblax Thomson, 1877 Baesia Burmeister, 1842 Baisarabaeus Nikolajev, 2005 Balanogonia Paucar-cabrera, 2003 Balbera Fairmaire, 1898 Ballucus Gordon & Skelley, 2007 Baloghonthobium Paulian, 1986 Baloghonthoboum Paulian, 1986 Balsameda Thomson, 1880 Barratus Barutus Ratcliffe, 1981 Barybas Blanchard, 1850 Basilewskyanus Endrödi, 1956 Basilewskynia Schein, 1957 Batesiana Chalumeau, 1983 Bdelyopsis Bdelyropsis Pereira, Vulcano & Martinez, 1960 Bdelyrus Harold, 1869 Beckhoplia Dombrow, 2005 Beriqua Péringuey, 1904 Besourenga Vaz-de-Mello, 2008 Bietia Fairmaire, 1898 Bilga Fairmaire, 1893 Bilobatus Machatschke, 1970 Biphyllocera White, 1841 Biralus Mulsant & Rey, 1869 Bisallardiana Antoine, 2003 Bisencya Lacroix, 1993 Bislardiana Antoine, 2003 Bisoroclita Krikken, 1982 Bivalgus Paulian, 1961 Bizanus Péringuey, 1902 Blabephorus Fairmaire, 1898 Blackburneus Schmidt, 1913 Blaesia Burmeister, 1842 Blanchardoplia Lacroix, 1998 Blapephorus Blebea Lacroix, 1994 Blepharotoma Blanchard, 1850 Blikana Péringuey, 1902 Blitopertha Reitter, 1903 Bobiricola Endrödi, 1973 Bodiloides Dellacasa & Dellacasa, 2005 Bodilopsis Adam, 1994 Bodilus Mulsant & Rey, 1869 Bohepilissus Paulian, 1975 Bolax Fisher Von Waldheim, 1829 Bolbites Harold, 1868 Boletoscapter Matthews, 1974 Bombodes Westwood, 1848 Bonoraella Ruter, 1978 Bordatius Pittino & Mariani, 1986 Boreocanthon Halffter, 1958 Bosphorus Şenyüz, 2017 Bothrorrhina Burmeister, 1842 Bothrothorax Kraatz, 1900 Bothynus Hope, 1837 Boucomontiellus Balthasar, 1932 Bourgoinigora Paulian, 1994 Brachagenius Kraatz, 1890 Brachiaphodius Koshantschikov, 1913 Brachylepis Kolbe, 1894 Brachyllus Brenske, 1896 Brachymacroma Kraatz, 1896 Brachymis Thomson, 1858 Brachymys Thomson, 1858 Brachypholis Brenske, 1898 Brachysiderus Waterhouse, 1881 Brachysternus Guérin-Méneville, 1831 Bradypodidium Vaz-de-Mello, 2008 Brahmina Blanchard, 1850 Brancoplia Baraud, 1986 Brenskeiella Berg, 1898 Brenskiella Berg, 1898 Breviclypeus Ricchiardi, 2017 Bricoptis Burmeister, 1842 Brindalus Landin, 1960 Brittonius Özdikmen & Demir, 2008 Bruchaphodius Martinez, 1952 Bruneixenus Howden & Storey, 1992 Bubas Dejan, 1833 Bubas Mulsant, 1842 Buettikeria Sabatinelli & Pontuale, 1998 Bunbunius Nomura, 1970 Burmeisteriellus Berg, 1898 Burmeistoplia Dombrow, 2003 Butozania Mikšić, 1955 Byrasba Harold, 1869 Byrrhidium Harold, 1869 Byrrhomorpha Blackburn, 1892 Byrromorpha Blackburn, 1892 Byrsalepis Brenske, 1898 Byrsopolis Burmeister, 1844 Caccobiomorphus Balthasar, 1964 Caccobius Thomson, 1859 Caccocnemus Caeconthobium Paulian, 1984 Caelius Lewis, 1895 Caelorrhina Burmeister, 1843 Caelorrhina Hope, 1841 Caesariatoplia Lacroix, 1998 Calamosternus Motschulsky, 1859 Calcitoryctes Krell, 2011 Calicnemis Laporte, 1832 Caligodorus Gordon & Skelley, 2007 Callabonica Blackburn, 1895 Calliferoplia Lacroix, 1998 Callipechis Burmeister, 1842 Callirhinus Blanchard, 1850 Callistemonus Péringuey, 1901 Callistethus Blanchard, 1850 Callistopopillia Ohaus, 1903 Calloodes White, 1845 Callophylla Moser, 1916 Calloserica Brenske, 1894 Callynomes Mohnike, 1873 Calocampsiura Miksic, 1987 Calocolobopterus Dellacasa, 1984 Calodactylus Blanchard, 1850 Caloglycyphana Miksic, 1968 Calomacraspis Bates, 1888 Calometopidius Bourgoin, 1917 Calometopus Blanchard, 1850 Calomosternus Motschulsky, 1860 Calopotosia Reitter, 1899 Caloprotaetia Mikšic, 1963 Calypsoryctes Howden, 1970 Calysoryctes Howden, 1970 Camapterus Ricchiardi, 2000 Cambefortantus Paulian, 1986 Cambefortius Branco, 1989 Camenta Erichson, 1847 Camentoserica Brenske, 1900 Camerounophylla Lacroix, 2005 Camerunopholis Lacroix, 2002 Campsiura Hope, 1831 Campulipes Campulipus Kirby, 1827 Campylophyllus Decelle, 1979 Campylotrochalus Brenske, 1900 Campytrochalus Brenske, 1900 Candezeollus Stebnicka & Howden, 1995 Canestera Saylor, 1938 Canthidium Erichson, 1847 Canthochilum Chapin, 1934 Canthodimorpha Davis, Scholtz & Harrison, 1999 Canthon Hoffmannsegg, 1817 Canthonella Chapin, 1930 Canthonidia Paulian, 1939 Canthonini van Lansberge, 1874 Canthonosoma MacLeay, 1871 Canthotrypes Paulian, 1939 Canudema Lacroix, 1994 Canuschiza Lacroix, 1999 Captomia Burmeister, 1842 Captomiopsis Pouillaude, 1919 Carinaulus Tesař, 1945 Carlschoenherria Bezdĕk, 2016 Carneiola Endrödi, 1974 Carneluttia Mikšić, 1976 Carneodon Özdikmen, 2009 Carneoryctes Özdikmen, 2009 Cartwrightia Islas, 1958 Castanochilus Ohaus, 1909 Castelnauphylla Lacroix, 1999 Catharsiocopris Balthasar, 1967 Catharsius Hope, 1837 Catoclastus Solier, 1851 Catrachia Coca-Abia & Robbins, 2006 Cavonus Sharp, 1875 Caymania Ratcliffe & Cave, 2010 Ceafornotensis Woolley, 2016 Celebesia Alexis & Delpont, 1998 Celebesiana Alexis & Delpont, 2002 Celidota Burmeister, 1842 Celidotella Reitter, 1909 Centrantyx Fairmaire, 1884 Centrocheilus Krikken, 1976 Centrochilus Krikken, 1976 Centrognathus Guérin-Méneville, 1840 Cephalocosmus Kraatz, 1895 Cephalocyclus Dellacasa, Gordon & Dellacasa, 1998 Cephalodesmius Westwood, 1842 Cephaloncheres Brenske, 1898 Cephaloschiza Moser, 1920 Ceramida Baraud, 1987 Ceraspis Le Peletier & Serville, 1828 Ceratochelus Dombrow, 2002 Ceratogonia Kolbe, 1899 Ceratolontha Arrow, 1948 Ceratophileurus Ohaus, 1911 Ceratoryctoderus Arrow, 1908 Cerhomalus Quedenfeldt, 1884 Ceroplophana Gestro, 1893 Cesamexico Koçak & Kemal, 2008 Cestradoretus Ohaus, 1912 Cetonia Fabricius, 1775 Cetonischema Reitter, 1899 Chacocopris Chaetadoretus Ohaus, 1914 Chaetocosmetes Moser, 1917 Chaetodermatina Heller, 1921 Chaetodermina Heller, 1921 Chaetonyx Schaum, 1862 Chaetopisthes Westwood, 1847 Chaetopisthides Wasmann, 1918 Chaetopteroplia Medvedev, 1949 Chaetovalgus Moser, 1914 Chalcasthenes Arrow, 1937 Chalcentis Burmeister, 1844 Chalcochlamys Ohaus, 1905 Chalcocopris Burmeister, 1846 Chalcocrates Heller, 1903 Chalconotus Dejean, 1833 Chalcopharis Heller, 1902 Chalcoplethis Burmeister, 1844 Chalcoprotaetia Mikšic, 1963 Chalcosoma Hope, 1837 Chalcothea Burmeister, 1842 Chalcotheomima Mikšić, 1970 Chalepides Casey, 1915 Charadronota Burmeister, 1842 Chariochilus Brenske, 1898 Chariodactylus Moser, 1919 Chariodema Blanchard, 1850 Charioserica Brenske, 1899 Charitovalgus Kolbe, 1904 Chasmatopterus Dejean, 1821 Chasmatopterus Latreille, 1825 Chasme Le Peletier & Serville, 1828 Chaunocolus Saylor, 1937 Cheilo Britton, 1990 Cheiragra MacLeay, 1864 Cheirodontus Britton, 1957 Cheirolasia Westwood, 1842 Cheironitis Lansberge, 1875 Cheiroplatys Hope, 1837 Cheiroplatys Kirby, 1837 Cheirora Britton, 1957 長臂金龜屬 Cheirotonus Hope, 1841 Cheleion Vardal & Forshage, 2010 Cheleion jendeki Král & Hájek, 2015 Cheleion malayanum Vardal & Forshage, 2010 Cheleion watanabei Kakizoe, Maruyama & Masumoto, 2019 Chelilabia Morón & Nogueira, 1998 Chelorrhina Burmeister, 1842 Cherbezatina Lacroix, 1993 Cheronia Croissandeau, 1893 Chewia Legrand, 2004 Chilamblys Kraatz, 1880 Chilenopilus Smith & Mondaca, 2016 Chilodiplus Sharp, 1877 Chiloloba Burmeister, 1842 Chiloschiza Moser, 1924 Chilothorax Motschulsky, 1859 Chilotrogus Reitter, 1905 Chioneosoma Kraatz, 1891 Chipita Soula, 2008 Chiriquiba Chiriquibia Bates, 1889 Chiron MacLeay, 1819 Chironitis Bedel, 1892 Chittius Tarasov, 2008 Chlaenobia Blanchard, 1850 Chloresthia Fairmaire, 1905 Chlorixanthe Bates, 1889 Chlorobapta Kraatz, 1880 Chlorocala Kirby, 1828 Chlorota Burmeister, 1844 Chnaunanthus Burmeister, 1844 Chondropyga Kraatz, 1880 Chondrorrhina Kraatz, 1880 Chromoptilia Westwood, 1842 Chromovalgus Kolbe, 1897 Chrysina Kirby, 1828 Chrysoliocola Alexis & Delpont, 1998 Chrysophora Dejean, 1821 Chrysophora Le Peletier & Serville, 1828 Chrysopotosia Miksic, 1966 Chrysoserica Brenske, 1897 Chthonobius Burmeister, 1847 Chtonobius Chuchina Soula, 2005 Cinacanthus Schmidt, 1913 Ciocotalpa Saylor, 1940 Circellium Latreille, 1829 Cjaradria Clania Schein, 1956 Clastocnemis Burmeister, 1840 Clavipalpus Laporte, 1832 Cleptocaccobius Cambefort, 1984 Clerota Burmeister, 1842 Clilopocha Lea, 1914 Clineterocera Clinteria Burmeister, 1842 Clinterocera Motschulsky, 1857 Clinteroides Schoch, 1898 Clipadoretus Ohaus, 1941 Clithria Burmeister, 1842 Clitocephala Burmeister, 1855 Clitopa Erichson, 1847 Clomecyra Lacroix, 1999 Clydonophorus Kraatz, 1899 Clypeasta Fairmaire, 1904 Clypeodrepanus Krikken, 2009 Clypeolontha Li & Yang, 1999 Clypeoserica Frey, 1970 Clyster Arrow, 1908 Cnemargulus Semenov, 1903 Cnematoplatys Schmidt, 1913 Cnemida Kirby, 1827 Cnemidophileurus Kolbe, 1910 Cnemisus Motschulsky, 1868 Cnemoschiza Decelle, 1979 Cocherellus Paulian, 1991 Cochinchidema Lacroix, 1996 Cochliotis Kolbe, 1894 Cochliotodes Burgeon, 1946 Coega Péringuey, 1902 Coelocorynus Kolbe, 1895 Coelocratus Burmeister & Schaum, 1841 Coelogenia Moser, 1913 Coelosis Hope, 1837 Coelosis Kirby & Spence, 1828 Coelotrachelus Schmidt, 1913 Coenochilus Schaum, 1841 Coenonycha Horn, 1876 Coenoryctoderus Prell, 1933 Coilodera Hope, 1831 Colacus Ohaus, 1910 Collagenus Ratcliffe & Hardy, 2005 Colobonthophagus Balthasar, 1935 Colobopteridius Clement, 1958 Colobopteroides Paulian, 1942 Colobopterus Mulsant, 1842 Colobostoma Blanchard, 1850 Colpochelyne Britton, 1987 Colpochila Erichson, 1843 Colpochilodes Blackburn, 1898 Colymbomorpha Blanchard, 1850 Comaserica Brenske, 1897 Comatapogonia Lacroix, 2008 Comencya Lacroix, 1993 Comophorina Strand, 1928 Comoramorphochelus Lacroix, 1997 Compsocephalus White, 1845 Compsodactylus Fuhrmann, 2012 Comythovalgus Kolbe, 1884 Conebius Fauvel, 1903 Coniopholis Erichson, 1847 Conioserica Brenske, 1900 Coniotrogus Brenske, 1903 Conradtia Kolbe, 1892 Copridaspidus Boucomont, 1920 Coprimorphus Mulsant, 1842 Copris Geoffroy, 1762 Copris Müller, 1764 Coprocanthon Martinez, 1950 Coproecus Reiche, 1841 Coprophanaeoides Edmonds, 1972 Coprophanaeus Olsoufieff, 1924 Coprophaneus Coprophnaeoides Coptochiroides Balthasar, 1938 Coptochirus Harold, 1859 Coptodactyla Burmeister, 1846 Coptognathus Burmeister, 1847 Coptomia Burmeister, 1842 Coptomiopsis Pouillaude, 1919 Coptorhina Hope, 1833 Coptorrhina Corynophyllus Hope, 1845 Corynoserica Moser, 1918 Corynotrichius Kolbe, 1892 Coryphaeus Gistl, 1848 Coryphocera Burmeister, 1842 Corythoderus Klug, 1845 Coscinocephalus Prell, 1936 Cosmesthes Kraatz, 1880 Cosmiomorpha Saunders, 1852 Cosmiophaena Kraatz, 1898 Cosmiophaenia Cosmovalgus Kolbe, 1897 Costelytra Given, 1952 Costinota Schürhoff, 1933 Cotalpa Burmeister, 1844 Cotinis Burmeister, 1842 Cotinorrhina Schoch, 1895 Craniorphnus Kolbe, 1895 Craterocyphus Schmidt, 1913 Crathoplus Blanchard, 1850 Cratocnemus Blanchard, 1842 Crecotoma Nikolajev, 2002 Cremastocheilus Knoch, 1801 Cremastodus Solier, 1851 Crematochila Crepischiza Brenske, 1903 Cretaclopus Nikolajev, 2004 Cretaegialia Nikolajev, 1993 Cretomelolontha Nikolajev, 1998 Cretonitis Nikolajev, 2007 Cretorabaeus Nikolajev, 1995 Cretoscarabaeus Nikolajev, 1995 Cretoserica Nikolajev, 1998 Criniflava Goodrich, 1966 Cryphaeobius Kraatz, 1882 Cryptaphodius Bordat, 2009 Cryptocanthon Balthasar, 1942 Cryptodontes Burmeister, 1847 Cryptodontus Burlini, 1969 Cryptodus MacLeay, 1819 Cryptoscatomaseter Gordon & Skelley, 2007 Cryptotrogus Kraatz, 1888 Crytocanthon Burmeister, 1846 Cubidens Britton, 1992 Cunderdinia Lea, 1916 Cyclidiellus Krikken, 1976 Cyclidinus Westwood, 1874 Cyclidius MacLeay, 1838 Cyclocephala Dejean, 1821 Cyclocephala Latreille, 1829 Cyclomera Klug, 1855 Cyclophorellus Krikken, 1982 Cylichnus Burmeister, 1844 Cylindrocrates Schein, 1958 Cymboptera Ohaus, 1923 Cymophorus Kirby, 1827 Cyobius Sharp, 1875 Cyphochilus Waterhouse, 1867 Cyphonistes Burmeister, 1847 Cyphonocephalus Westwood, 1842 Cyphonotus Fischer von Waldheim, 1823 Cyphoserica Brenske, 1900 Cyprolais Thomson, 1880 Cyptochirus Lesne, 1900 Cyriodera Burmeister, 1842 Cyriopertha Reitter, 1903 Cyrrhochroea Kraatz, 1900 Cyrtocamenta Brenske, 1897 Cyrtolophus Fairmaire, 1895 Cyrtothyrea Kolbe, 1895 Cyrtotrochalus Brenske, 1900 Dactylopopillia Arrow, 1917 Daemonoplus Thomson, 1858 Daintreeola Howden & Storey, 2000 Dalatamala Prokofiev, 2013 Dalgopus Dechambre, 2001 Damonplus Danielssonia Bordat & Howden, 1995 Dasygnathus MacLeay, 1819 Dasylepida Moser, 1913 Dasytrogus Reitter, 1902 Dasyus Le Peletier & Serville, 1828 Dasyvalgoides Endrödi, 1952 Dasyvalgus Kolbe, 1904 Debeckius Allsopp, 2022 Debutina Lacroix, 1993 Decellophylla Lacroix, 2005 Dechambrophylla Lacroix, 1999 Dedalopterus Sabatinelli & Pontuale, 1998 Degallieridium Vaz-de-Mello, 2008 Dellacasiellus Gordon & Skelley, 2007 Delopleurus Erichson, 1848 Delphinobius Fairmaire, 1900 Deltepilissus Pereira, 1949 Deltochilum Eschscholtz, 1822 Deltorhinum Harold, 1867 Deltorrhinum Gemminger & Harold, 1869 Demarziella Balthasar, 1961 Democrates Burmeister, 1847 Dendropaemon Perty, 1830 Denhezia Dechambre, 2006 Dentatoplia Lacroix, 1997 Denticnema Schein, 1959 Dentiheterochelus Dombrow, 2002 Dermolepida Arrow, 1941 Deroserica Moser, 1915 Descarpentriesia Ruter, 1964 Desertaclopus Ocampo & Mondaca, 2012 Desfontainesia Alexis & Delpont, 1999 Desicasta Thomson, 1878 Desmonyx Arrow, 1907 Diabroctis Gistel, 1857 Diaclaspus Brenske, 1896 Diacucephalus De Wailly, 1950 Dialithus Parry, 1849 Dialytellus Brown, 1929 Dialytes Harold, 1869 Dialytodius Gordon & Skelley, 2007 Diaphonia Newman, 1840 Diaphoroserica Péringuey, 1904 Diaphylla Erichson, 1847 Diaplochelus Burmeister, 1844 Diapterna Horn, 1887 Diastellopalpus Lansberge, 1886 Diastictus Mulsant, 1842 Diasticus Mulsant, 1842 Diathermus Kraatz, 1897 Dibolus Endrodi, 1976 Dicaulocephalus Gestro, 1888 Dicellachilus Waterhouse, 1905 Dicentrines Burmeister, 1844 Dichecephala Brenske, 1895 Dichelhoplia Blanchard, 1850 Dichelomorpha Burmeister, 1855 Dichelonyx Harris, 1826 Dicheloscema Moser, 1924 Dicheloschema Moser, 1924 Dichelus Le Peletier & Serville, 1828 Dicheros Gory & Percheron, 1833 Dichodontus Burmeister, 1847 Dichotomius Hope, 1838 Dichrosoma Kraatz, 1885 Dicrania Le Peletier & Serville, 1828 Dicranobia Reitter, 1900 Dicranocara Frolov & Scholtz, 2003 Dicranocephalus Hope, 1831 Dicranocnemus Burmeister, 1844 Dicranoplia Reitter, 1903 Dicronocephalus Hope, 1831 Dicronorhina Hope, 1837 Dicronorrhina Didactylia Orbigny, 1896 Didrepanephorus Wood-mason, 1878 Digenethle Thomson, 1877 Digitonthophagus Balthasar, 1959 Dihymenonyx Gutiérrez, 1949 Dikellites Britton, 1990 Dilatatoplia Lacroix, 1998 Diloboderus Reiche, 1859 Dilochrosis Thomson, 1878 Dilophochila Bates, 1888 Dilortomaeus Bordat, 2009 Dimyxus Fairmaire, 1893 Dinacoma Casey, 1889 Dinamoraza Lacroix, 1989 Dinarobina Lacroix, 1989 Dineterogomphus Prell, 1912 Dineterohgomphus Dinorhina Lacordaire, 1856 Dinoryctes Felsche, 1906 Dionychus Burmeister, 1847 Diorygopyx Matthews, 1974 Dipelicus Hope, 1842 Diphrontis Gerstaecker, 1882 Diphucephala Le Peletier & Serville, 1828 Diphycerus Fairmaire, 1878 Diphydactylus Thomson, 1858 Diphyllomorpha Hope, 1843 Diploa Kolbe, 1892 Diploeida Péringuey, 1907 Diplognatha Gory & Percheron, 1833 Diplognathoides Holm & Perissinotto, 2004 Diplotaxis Kirby, 1837 Diplotropis Boheman, 1857 Dirrhina Burmeister, 1842 Dischista Burmeister, 1842 Discopeltis Burmeister, 1842 Disphysema Harold, 1873 Djadjoua Lacroix, 1993 Djafouna Lacroix, 1993 Doaphius Balthasar, 1941 Doedycorhina Bates, 1888 Doleroserica Péringuey, 1904 Dolerotarsa Péringuey, 1904 Dolichiomicroscelis Dombrow, 2003 Dolichoplia Lacroix, 1998 Dolichostethus Kolbe, 1892 Dorbignyolus Branco, 1989 Dorcus Gistl, 1848 Doryscelis Burmeister, 1842 Doryscelis Dejean, 1836 Dorysthetus Blanchard, 1845 Doxocalia Brenske, 1897 Drepanacanthoides Schmidt, 1913 Drepanocanthoides Schmidt, 1913 Drepanocanthus Péringuey, 1901 Drepanocerus Kirby, 1828 Drepanoplatynus Boucomont, 1921 Drogo Deschodt, Davis & Scholtz, 2016 Dudleyellus Bordat, 2009 Dungoorus Carne, 1958 Dwesasilvasedis Deschodt & Scholtz, 2008 Dymusia Burmeister, 1842 Dynastes Kirby, 1825 Dynastes MacLeay, 1819 Dyscinetus Harold, 1869 Dysphanocheila Blackburn, 1898 Dysphanochila Blackburn, 1898 Dyspilophora Kraatz, 1880 Ebolowanius Dechambre, 1979 Eccoptomia Kraatz, 1880 Echyra Erichson, 1847 Ectinohoplia Redtenbacher, 1868 Ectinoplectron Ohaus, 1915 Eideria Neita & Ocampo, 2012 Elaphinis Burmeister, 1842 Elaphocera Gene, 1836 Elaphocerella Moser, 1919 Elassochiton Kolbe, 1895 Elcarmeniella Franz, 1955 Elpidus Péringuey, 1907 Emadiellus Schmidt, 1913 Embrithoplia Lacroix, 1998 Empecamenta Brenske, 1895 Empecta Erichson, 1847 Empectoides De Wailly, 1950 Emphania Erichson, 1847 Empycastes Gerstaecker, 1871 Enamillus Sharp, 1877 Enaria Erichson, 1847 Enarotadius Endrödi, 1971 Encya Dejean, 1833 Endebius Lansberge, 1880 Endoxazus Kolbe, 1892 Endreodyolus Endrodius Endroedia Mikšic, 1958 Endroedianibe Chalumeau, 1981 Endroedyolus Scholtz & Howden, 1987 Enema Hope, 1837 Energetia Dalle Torre, 1912 Engertia Dalle Torre, 1912 Engistanoxia Lacroix, 2002 Engyopsina Britton, 1990 Enicotarsus Laporte, 1831 Ennearabdus Lansberge, 1874 Enoplotarsus Lucas, 1859 Enracius Dechambre, 1999 Enthora Erichson, 1847 Entyopsis Kolbe, 1894 Entypophana Moser, 1913 Entyposis Kolbe, 1894 Eoalleucosma Antoine, 1989 Eocorythoderus Marumaya, 2012 Eocorythoderus incredibilis Marumaya, 2012 Eodrepanus Barbero, Palestrini & Roggero, 2009 Eoheterorrhina Miksic, 1974 晓扁犀金龟属 Eophileurus Arrow, 1908 Eosaulostomus Carne, 1956 Eotrichia Medvedev, 1951 Eotrochalus Burgeon, 1943 Epactoides Olsoufieff, 1947 Epadoretus Reitter, 1903 Epectinaspis Blanchard, 1850 Ephirinus Epicaulis Dejean, 1833 Epichalcoplethis Bates, 1904 Epichalcoplethis MacLeay, 1817 Epichrysus White, 1841 Epicometes Epicometis Burmeister, 1842 Epidrepanus Roggero, Barbero & Palestrini, 2015 Epilissus Blanchard, 1845 Epilissus Reiche, 1841 Epipholis Moser, 1917 Epirinus Dejean, 1833 Epironastes Carne, 1957 Epistalagma Fairmaire, 1880 Epitrichius Tagawa, 1941 Epixanthis Burmeister, 1842 Erbmahcedius Hutchinson & Allsopp, 2021 Ercomoana Lacroix, 1994 Eremazus Mulsant, 1851 Eremobothynus Ohaus, 1910 Eremonthophagus Zunino, 1979 Eremophygus Ohaus, 1910 Eremoplia Kolbe, 1914 Eremoryctes Semenov-Tian-Shanskii & Medvedev, 1932 Eremotrogus Kolbe, 1914 Eriesthis Burmeister, 1844 Erigavo Lacroix, 2006 Eriopeltastes Burmeister & Schaum, 1840 Erioscelis Burmeister, 1847 Eriphoserica Brenske, 1899 Eriulis Burmeister, 1842 Erlangeria Preiss, 1902 Erytodes Schmidt, 1916 Erytus Mulsant & Rey, 1870 Escarabaeus Zídek & Pokorný, 2011 Estenomenus Faldermann, 1835 Esymaphodius Esymus Mulsant & Rey, 1869 Etiserica Péringuey, 1904 Eucamenta Péringuey, 1904 Eucampsiura Miksic, 1987 Eucanthidium Martinez & Halffter, 1986 Eucetonia Schoch, 1894 Euchilia Burmeister, 1842 Euchirus Burmeister & Schaum, 1840 Euchirus Kirby & Spence, 1828 Euchloropus Arrow, 1907 Euchroa Gistl, 1848 Euchroea Burmeister, 1842 Euchromoplia Medvedev, 1952 Eucoelosis Iannuzzi & Marinoni, 1995 Eucopidocaulus Prell, 1912 Eucranium Brullé, 1834 Eucyclophylla Waterhouse, 1875 Eucymophorus Schein, 1954 Eudicella White, 1839 Eudinopus Burmeister, 1846 Eudolus Mulsant & Rey, 1869 Euetheola Bates, 1888 Euglycyphana Miksic, 1968 Euglypta Mohnike, 1873 Euhemicyclium Bordat, 2003 Euheptaulacus Dellacasa, 1983 Euhyboma Kolbe, 1893 Euknemoplia Lacroix, 1997 Eulaiades Fairmaire, 1899 Eulepida Kolbe, 1894 Eulepidopsis Burgeon, 1946 Euligyrus Casey, 1915 Eumacronota Mikšić, 1976 Eumaladera Nomura, 1967 Eunanus Ohaus, 1909 Euoniticellus Janssens, 1953 Euonthophagus Balthasar, 1959 Euorodalus Dellacasa, 1983 Euotophorus Dellacasa, Gordon & Dellacasa, 2016 Euparia Le Peletier & Serville, 1828 Euparixia Brown, 1927 Euparixoides Hinton, 1936 Euparotrix Stebnicka & Howden, 1996 Eupatorus Burmeister, 1847 Eupegylis Duvivier, 1892 Euphoresia Brenske, 1897 Euphoria Burmeister, 1842 Eupleurus Mulsant, 1842 Eupoecila Burmeister, 1842 Eupotosia Miksic, 1954 Euprigenia Miksic, 1974 Euprotaetia Mikšic, 1963 Euranoxia Semenov, 1890 Euremina Westwood, 1867 Europtron Marseul, 1867 Euryaspis Blanchard, 1850 Cheleion jendeki Král & Hájek, 2015 Cheleion malayanum Vardal & Forshage, 2010 Cheleion watanabei Kakizoe, Maruyama & Masumoto, 2019 Eurychelus Blanchard, 1850 Eurycorythoderus Wasmann, 1918 Euryderus Gemminger & Harold, 1869 Eurynomia Euryomia Burmeister, 1842 Eurypeza Lacroix, 2006 Eurypodea Klages, 1906 Euryschiza Brenske, 1898 Eurysternini Vulcano, Martinez & Pereira, 1960 Eurysternus Dalman, 1824 Euryvalgus Moser, 1908 Eusaproecius Branco, 1989 丽花金龟属 Euselates Thomson, 1880 Euserica Reitter, 1896 Eutelesmus Waterhouse, 1880 Eutheola Eutlesthes Eutrichesis Waterhouse, 1882 Eutrichillum Martinez, 1967 Eutyctus Semenov, 1890 Evanos Castelnau, 1840 Evbrittonia Szito, 1993 Exanticheira Soula, 1998 Exantichira Soula, 1998 Exaphodius Endrödi, 1960 Eximidema Keith & Reichenbach, 2018 Exochlorota Soula, 2002 Exochogenys Carne, 1958 Exolontha Reitter, 1902 Exomala Reitter, 1903 Exopholis Motschulsky, 1859 Exoptenomela Soula, 2002 Exostethus Britton, 1979 Exothyridium Soula, 2002 Exoxyomus Bordat, Mencl & Rakovič, 2012 Extenuoptyophis Smith & Mondaca, 2015 Faargia Martinez, 1982 Fairmairiana Alexis & Delpont, 1998 Fairmairoplia Lacroix, 1997 Falsignambia Paulian, 1987 Falsotoclinius Keith & Montreuil, 2004 Faula Blanchard, 1850 Feeridium Vaz-de-Mello, 2008 Ferreiria Ferrerianus Dellacasa, Dellacasa & Gordon, 2007 Finkia Mikšic, 1965 Firminus Coca-Abia, 2003 Flaviellus Gordon & Skelley, 2007 Flechtmanniella Stebnicka, 1999 Fornasinius Bertoloni, 1852 Fossocarus Howden, 1961 Foveopotosia Mikšic, 1957 Francmonrosia Pereira & Martinez, 1960 Frankenbergerius Balthasar, 1938 Frenchella Blackburn, 1892 Fruhstorferia Kolbe, 1894 Furconthophagus Zunino, 1979 Gametis Burmeister, 1842 Gametoides Antoine, 2005 Gamka Péringuey, 1904 Ganganomala Ratcliffe, Jameson & Zorn, 2018 Gariep Péringuey, 1907 Garreta Janssens, 1940 Gastromaladera Nomura, 1973 Gastroserica Brenske, 1897 Geminigora Paulian, 1994 Geniates Kirby, 1818 Geniatosoma Costa Lima, 1940 Genieridium Vaz-de-Mello, 2008 Genuchinus Kirby, 1825 Genuchinus Westwood, 1874 Genuchus Kirby, 1825 Geocanthon Pereira & Martínez, 1956 Geomyphilus Gordon & Skelley, 2007 Geopsammodius Gordon & Pittino, 1992 Geotrogus Guérin-Méneville, 1842 Ghania Endrodi, 1973 Gibboryctes Endrödi, 1974 Giesbertiolus Howden, 1988 Gillaspytes Howden, 1980 Gilletellus Janssens, 1937 Gilletianus Balthasar, 1933 Glacilitarsus Matsumoto, 2014 Glaphyocanthon Glaphyronyx Moser, 1924 Glaphyserica Brenske, 1899 Glenopopillia Linnaeus, 1980 Globencya Lacroix, 1993 Globulischiza Frey, 1974 Glossocheilifer Blackburn, 1898 Glycosia Schoch, 1896 Glycyphana Burmeister, 1842 Glycyphaniola Miksic, 1968 Glycyserica Brenske, 1899 Glyphoderus Agassiz, 1846 Glyphoderus Westwood, 1838 Glyptoglossa Brenske, 1895 Glyptoterus Glyptothea Bates, 1889 Glyptotheomima Mikšić, 1976 Gnaphalopoda Reiche, 1860 Gnaphalostetha Reiche, 1856 Gnathocera Kirby, 1825 Gnathogolofa Arrow, 1914 Gnatholabis Erichson, 1847 Gnorimella Casey, 1915 Gnorimidia Lansberge, 1887 Gnorimimelus Kraatz, 1880 Gnorimotrichius Krajčik, 2009 Gnorimus Le Peletier & Serville, 1828 Goetzia Mikšic, 1963 Goiginus Endrödi, 1982 Goliata Goliathinus Westwood, 1874 Goliathopsis Janson, 1881 Goliathus Lamarck, 1801 Golinca Thomson, 1878 Golofa Hope, 1837 Gomphas Brullé, 1834 Gonaphodiellus Schmidt, 1913 Gonaphodioides Dellacasa, Dellacasa & Gordon, 2012 Gonaphodiopsis Dellacasa, Dellacasa & Gordon, 2012 Gonaphodius Reitter, 1892 Gongrolophus Stebnicka & Howden, 1996 Goniaspidius Burmeister, 1844 Goniaspidius Périguey, 1902 Goniophileurus Kolbe, 1910 Goniorphnus Arrow, 1911 Goniorrhina Quedenfeldt, 1888 Gonoprotaetia Mikšic, 1965 Gorditus Ratcliffe, 2010 Gordonius Skelley, Dellacasa & Dellacasa, 2009 Gouna Péringuey, 1902 Gracilaclopus Ocampo & Mondaca, 2012 Grammodoplia Lacroix, 1997 Grammopyga Kolbe, 1895 Grandaustralis Hutchinson & Moeseneder, 2013 Grandinaphodius Ziani, 2002 Granulopsammodius Rakovič, 1981 Graviengertia Prokofiev, 2016 Grebennikovius Mlambo, Scholtz & Deschodt, 2019 Griveaudella Lacroix, 1993 Gromphas Brullé, 1837 Gronocarus Schaeffer, 1927 Gryphonycha Péringuey, 1904 Grypnocnemis Guanyinaphodius Masumoto & Kiuchi, 2001 Guatemalica Neervoort Van De Poll, 1886 Gymnephoria Ratcliffe, 2019 Gymnetina Casey, 1915 Gymnetina Kirby, 1827 Gymnetis MacLeay, 1819 Gymnogaster Blanchard, 1850 Gymnoloma Dejean, 1833 Gymnophana Arrow, 1910 Gymnopleurus Illiger, 1803 Gymnopyge Linell, 1896 Gymnoschiza Moser, 1914 Gynaecoserica Brenske, 1896 Gyronotus Lansberge, 1874 Gyroplia Brenske, 1893 Hadrodiplognatha Kraatz, 1898 Hadropechys Britton, 1990 Hadrops Britton, 1987 Hadrorhyparus Howden, 1995 Hadrosticta Kraatz, 1892 Haematonotus Kraatz, 1880 Halffterinetis Morón & Nogueira, 2007 Hamatoplectris Frey, 1967 Hammondantus Cambefort, 1978 Hamonthophagus Roggero, Dierkens, Barbero & Palestrini, 2017 Hansreia Halffter & Martinez, 1977 Haplidia Hope, 1837 Haplophileurus Kolbe, 1910 Haploscapanes Arrow, 1908 Haplosoma Semenov, 1890 Harmodactylus Péringuey, 1901 Harmogaster Harold, 1861 Haroldaphodius Bordat, 2003 Haroldiataenius Chalumeau, 1981 Haroldiellus Gordon & Skelley, 2007 Haroldius Boucomont, 1914 Haroldostes Paulian, 1982 Harpechys Britton, 1957 Harpina Burmeister, 1844 Harposceles Burmeister, 1847 Hatamus Sharp, 1877 Hecistopsilus Kolbe, 1894 Hegemus Thomson, 1881 Helictopleurus Orbigny, 1915 Heliocopris Hope, 1837 Helionica Thomson, 1881 Hellaserica Baraud & Nicolas, 1966 Hemiaspidius Krikken, 1982 Hemicamenta Brenske, 1897 Hemicephalocosmus Miksic, 1974 Hemichaetoplia Baraud, 1986 Hemichalcothea Mikšić, 1970 Hemichnoodes Kraatz, 1880 Hemicoryphocera Miksic, 1974 Hemicorythoderus Tangelder & Krikken, 1982 Hemictenius Reitter, 1897 Hemicyclium Schmidt, 1913 Hemicyrthus Reiche, 1860 Hemiheterorrhina Mikšić, 1974 Hemilia Kraatz, 1880 Hemiopta Bedel, 1896 Hemiphaedimus Mikšić, 1972 Hemipharia Hemipharis Burmeister, 1842 Hemiphileurus Kolbe, 1910 Hemiprigenia Miksic, 1974 Hemiprotaetia Mikšić, 1963 Hemiserica Brenske, 1894 Hemitheodosia Sakai, 1994 Hemycorythoderus Heptaulacus Mulsant, 1842 Heptelia Brenske, 1898 Heptophylla Motschulsky, 1857 Heptophyllla Motschulsky, 1857 Hercites Burmeister, 1855 Herculaisia Seilliere, 1910 Heteralleucosma Antoine, 1989 Heterelaphinis Antoine, 2002 Heterochelus Burmeister, 1844 Heterochlorota Soula, 2002 Heteroclita Burmeister, 1842 Heteroclitopus Péringuey, 1901 Heterocnemis Albers, 1852 Heteroconus Kolbe, 1900 Heterocotinis Martinez, 1948 Heterocotinus Martinez, 1948 Heterocranus Bourgoin, 1919 Heterogeniates Ohaus, 1909 Heterogenius Moser, 1911 Heteroglobus Dupuis & Dechambre, 2008 Heteroglycyphana Miksic, 1968 Heterogomphus Burmeister, 1847 Heteroligus Kolbe, 1900 Heteronitis Gillet, 1911 Heteronychus Burmeister, 1847 Heteronychus Dejean, 1833 Heteronyx Guérin-Méneville, 1830 Heterophana Burmeister, 1842 Heterophthalmus Blanchard, 1850 Heteroprotaetia Mikšic, 1963 Heteropseudinca Valck Lucassen, 1933 Heterorhina Westwood, 1842 Heterorrhiniola Miksic, 1974 Heteroschiza Moser, 1917 Heteroserica Brenske, 1899 Heterosoma Schaum, 1844 Heterosternus Dupont, 1832 Heterostetha Moser, 1911 Heterostheta Heterostriatus Dupuis, 2006 Heterotephraea Antoine, 2002 Heterotrochalus Moser, 1918 Heterovalgus Krikken, 1978 Heteroychus Hexataenius Fairmaire, 1891 Hexodon Olivier, 1789 Hiberasta Fairmaire, 1901 Hiekeianus Endrödi, 1978 Hieritis Burmeister, 1855 Hilyotrogus Fairmaire, 1886 Himalhoplia Sabatinelli, 1983 Hispanioryctes Howden & Endrodi, 1978 Holcoribeus Nikritin, 1977 Holcorobeus Nikritin, 1977 Holisonycha Péringuey, 1904 Holocanthon Martinez & Pereira, 1956 Holocephalus Hope, 1838 Holochelus Reitter, 1889 Holocnathon Martinez & Pereira, 1956 Holocnemus Brenske, 1894 Hologymnetis Martinez, 1949 Holomelia Brenske, 1891 Holophylla Erichson, 1847 Holopycnia Brenske, 1896 Holorhopaea Britton, 1978 Holotrichia Hope, 1837 Homalochilus Blanchard, 1850 Homaloserica Brenske, 1897 Homalotarsus Janssens, 1932 Homalothyrea Kolbe, 1895 Homeochlorota Soula, 2006 Homioliogenys Gutierrez Alonso, 1952 Homocanthonides Luederwaldt, 1929 Homocopris Burmeister, 1842 Homoeomorphus Burmeister, 1847 Homoeopsopha Schürhoff, 1934 Homoeoschiza Kolbe, 1894 Homoiosternus Ohaus, 1901 Homolosternus Ohaus, 1934 Homolotropus MacLeay, 1871 Homonyx Guérin-Méneville, 1839 Homophileurus Kolbe, 1910 Homopliopsis Lacroix, 1998 Homosphileurus Kolbe, 1910 Homothermon Ohaus, 1898 Homothyrea Kolbe, 1895 Hoplebaea Brenske, 1899 Hopleidos Künckel D'Herculais, 1887 Hoplia Illiger, 1803 Hopliopsis Blanchard, 1850 Hoplitogomphus Prell, 1912 Hoplitovalgus Kolbe, 1904 Hoplochelus Blanchard, 1850 Hoplocnemis Harold, 1869 Hoplocnemus Blanchard, 1851 Hoplolontha Fairmaire, 1904 Hoplomaladera Nomura, 1974 Hoplopelidnota Bates, 1904 Hoplopus Laporte, 1832 Hoplopyga Thomson, 1880 Hoplopygothrix Schürhoff, 1933 Hoplorida Moser, 1918 Hoploryctoderus Prell, 1933 Hoplosternodes Burgeon, 1946 Hornietus Stebnicka, 2000 Hornosus Dellacasa, Dellacasa & Gordon, 2015 Horpnus Paulian, 1951 Horridocalia Endrödi, 1974 Hovachelus Fairmaire, 1897 Hovophileurus Arrow, 1911 Howdenocarus Hardy, 1978 Howdenypa Arnaud, 1993 Humblotania Lacroix, 1993 Hyalonthophagus Palestrini & Giacone, 1988 Hybaloides Quedenfeldt, 1884 Hybalus Dejean, 1833 Hybocamenta Brenske, 1898 Hyboschema Péringuey, 1901 Hyboserica Péringuey, 1904 Hybosorites Nikolajev, 1996 Hybovalgus Kolbe, 1904 Hylamorpha Arrow, 1899 Hylobothynus Ohaus, 1910 Hymenochelus Reitter, 1890 Hymenoplia Eschscholz, 1830 Hypaspidius Arrow, 1899 Hyperius Fairmaire, 1878 Hyphelithia Kraatz, 1880 Hyphoryctes Blackburn, 1895 Hypocanthidium Balthasar, 1938 Hypolepida Britton, 1978 Hyposerica Brenske, 1897 Hypothyce Howden, 1968 Hypotrichia LeConte, 1861 Hypselogenia Burmeister, 1846 Iapurea Martinez, 1953 Iarupea Martinez, 1953 Iberoaphodius Rössner, 2018 Ichnestoma Gory & Percheron, 1833 Ictigaster Britton, 1986 Idaecamenta Péringuey, 1904 Idaeserica Péringuey, 1904 Idanastes Britton, 1987 Idiapogonia Arrow, 1916 Idiochelyna Arrow, 1916 Idionycha Arrow, 1932 Idiovalgus Arrow, 1910 Idutywa Péringuey, 1902 Ignambia Heller, 1916 Iguazua Stebnicka, 1997 Ihosyus Montreuil & Lacroix, 2020 Inanda Péringuey, 1902 Inca Le Peletier & Serville, 1828 Incala Thomson, 1857 Incalidia Janson, 1907 Indieraligus Dechambre, 1979 Indoblackburneus Mencl, Král & Rakovič, 2019 Indocetonia Mikšic, 1965 Indognorimus Krikken, 2009 Indoliocola Alexis & Delpont, 1998 Indoprotaetia Indotrichius Krikken, 2009 Indovalgus Ricchiardi, 2012 Ingogius Endrödi, 1976 Ingrisma Fairmaire, 1893 Inhambane Péringuey, 1907 Insimuloissacaris Smith & Mondaca, 2015 Insuloryctes Dechambre, 2000 Iranotrogus Nikolajev, 2006 Irazua Ratcliffe, 2003 Iridisoma Delgado & Morón, 1991 Irrasinus Gordon & Skelley, 2007 Isacanthon Pacheco & Vaz-de-Mello, 2019 Ischiopsopha Gestro, 1874 Ischnestoma Ischnocelis Ischnopopillia Kraatz, 1892 Ischnoscelis Burmeister, 1842 Ischnostomiella Krikken, 1978 Ischnotarsia Kraatz, 1880 Ischnovalgus Kolbe, 1897 Ischyrocera Hauser, 1904 Ishiopsopha Isocopris Pereira & Martinez, 1960 Isonychus Mannerheim, 1828 Isoplia Burmeister, 1855 Issacaris Fairmaire, 1889 Ixodina Roth, 1851 Ixorida Thomson, 1880 Jalalabadia Balthasar, 1967 Jalisco Dellacasa, Gordon & Dellacasa, 2003 Jansongora Paulian, 1994 Jansonia Schürhoff, 1937 Janssensantus Paulian, 1976 Janssensellus Cambefort, 1975 Jossonthophagus Dierkens, Moretto & Génier, 2017 Joziratia Lacroix, 1993 Jumnos Saunders, 1839 Junkia Dalla Torre, 1913 Juraclopus Nikolajev, 2005 Kabindeknomiosoma Lacroix, 2005 Kadlecia Frey, 1965 Kareiga Péringuey, 1902 Karolinella Minkina, 2017 Karooida Perissinotto, 2020 Kenyalepis Lacroix, 2011 Kenyatopadema Lacroix, 2009 Kerowagia Delpont, 1996 Kheper Janssens, 1940 Khoina Péringuey, 1902 Kibakoganea Nagai, 1984 Kirprellius Allsopp, 2022 Knysna Péringuey, 1902 Kolbeellus Jacobson, 1906 Komrina Lacroix, 1993 Koreoxyomus Kim, 1996 Korisaba Péringuey, 1902 Koshantschikovius Schmidt, 1913 Kraseophylla Péringuey, 1904 Krikkenius Branco, 1991 Kryzhanovskia Nikolajev & Kabakov, 1977 Kubousa Péringuey, 1902 Kuenckeliana Lacroix, 1989 Kurtops Roggero, Barbero & Palestrini, 2017 Kuytenia Mikšic, 1963 Labarrus Mulsant & Rey, 1869 Labroma Sharp, 1873 Laceratoplia Lacroix, 1997 Lachnodera Erichson, 1847 Lachnosterna Hope, 1837 Lachnota Reitter, 1889 Lacroixidema Keith, 2000 Lacroixilepis Keith, 2007 Lagochile Hoffmannsegg, 1817 Lamellothyrea Krikken, 1980 Lamproserica Brenske, 1897 Lansbergia Ritsema, 1888 Laonthophagus Frey, 1971 Laotrichia Keith, 2007 Larhodius Balthasar, 1963 Lasiexis Semenov-tian-shanskii & Medvedev, 1936 Lasiocala Blanchard, 1850 Lasiopsis Erichson, 1847 Lasioserica Brenske, 1897 Lasiotrichius Reitter, 1899 Lasiotropus Reitter, 1913 Latemnis Thomson, 1880 Latescutella Ruter, 1972 Latipalpus Moser, 1921 Latodrepanus Krikken, 2009 Laurentiana Ruter, 1952 Lawangia Schenkling, 1921 Lebbea Lacroix, 1993 Lecanoderus Kolbe, 1907 Lecanotrogus Kolbe, 1894 Lecantotrogus Kolbe, 1894 Lechorodius Gordon & Skelley, 2007 Leiopsammodius Rakovič, 1981 Lenosoma MacLeay, 1863 Leonotus Britton, 1986 Leoserica Fabrizi, Eberle & Ahrens, 2019 Leotrichillum Vaz-de-Mello, 2008 Lepadoretus Reitter, 1903 Lepanus Balthasar, 1966 Lepesmonyx Paulian, 1991 Lepidiota Kirby, 1828 Lepidomela Kolbe, 1894 Lepidoserica Nikolajev, 1979 Lepidotrogus Kolbe, 1894 Lepischiza Moser, 1914 Lepisia Le Peletier & Serville, 1828 Lepithrix Le Peletier & Serville, 1828 Lepitrichula Schein, 1959 Lepivalgus Moser, 1914 Leptochristina Baraud & Branco, 1991 Leptocnemis Dombrow, 2001 Leptohoplia Saylor, 1935 Leptorhyparus Howden, 2003 Leptothyrea Kraatz, 1882 Leribe Péringuey, 1904 Leucocelis Burmeister, 1842 Leucopholis Dejean, 1833 Leucophorus Brenske, 1892 Leucoserica Reitter, 1896 Leucothyreus MacLeay, 1819 Leuretra Erichson, 1847 Liatongus Reitter, 1892 Liberocera Deloya & Ratcliffe, 1988 Lichnia Erichson, 1835 Lichniops Gutiérrez, 1946 Lichniopsoides Martinez, 1953 Licnostrategus Prell, 1933 Lignarius Gistl, 1848 Ligyrus Burmeister, 1847 Limarus Mulsant & Rey, 1869 Limepomera Lacroix, 1999 Linargius Rakovič & Mencl, 2012 Lindbergianus Petrovitz, 1961 Liogenys Guérin-Méneville, 1830 Lioglyptoxenus Pittino, 2006 Liomenochilus Hutchinson & Allsopp, 2022 Liostraca Burmeister, 1842 Liothorax Motschulsky, 1859 Liotrichius Kolbe, 1892 Liparetrus Guérin-Méneville, 1830 Lipothyrea Péringuey, 1907 Liptothyrea Lissadoretus Arrow, 1917 Lissogenius Schaum, 1844 Lissomelas Bates, 1889 Listrochelus Blanchard, 1850 Lithanomala Nikolajev, 1992 Lithoscarabaeus Nikolajev, 1992 Litocopris Waterhouse, 1891 Lobateuchus Montreuil, Génier & Nel, 2010 Lobidion Genier, 2010 Lobogeniates Ohaus, 1917 Loboparius Schmidt, 1913 Loboserica Kolbe, 1914 Lobovalgus Kolbe, 1897 Lomanoxia Martinez, 1951 Lomanoxoides Stebnicka, 1999 Lomaptera Gory & Percheron, 1833 Lonchothyrea Kolbe, 1895 Lonchotus Burmeister, 1847 Longaphodius Endrödi, 1980 Longicrura Frey, 1974 Lophodonitis Janssens, 1938 Lophorrhina Westwood, 1842 Lophorrhinides Perissinotto, Clennell & Beinhundner, 2019 Loraphodius Reitter, 1892 Loraspis Mulsant & Rey, 1869 Lorditomaeus Péringuey, 1901 Loricatarsa Frey, 1969 Lorkovitschia Mikšić, 1968 Lucassenia Olsoufieff, 1940 Luispenaia Martinez, 1972 Lunaphodius Balthasar, 1964 Lutera Westwood, 1875 Lutfius Özdikmen, 2009 Luxolinus Gordon & Skelley, 2007 Lycomedes Brême, 1844 Lyraphora Kraatz, 1880 Machala Lacroix, 1993 Macleayella Britton, 1990 Macraspis MacLeay, 1819 Macrocanius Schürhoff, 1935 Macrocranius Schürhoff, 1935 Macrocyphonistes Ohaus, 1910 Macrodactylus Berthold, 1827 Macrodactylus Dejean, 1821 Macrodactylus Latreille, 1825 Macroderes Westwood, 1842 Macroglycyphana Miksic, 1968 Macroliocola Alexis & Delpont, 1998 Macromina Westwood, 1874 Macrominus McLeay, 1838 Macronota Hoffmannsegg, 1817 Macronotops Krikken, 1977 Macronthophagus Ochi, 2003 Macropanelus Ochi, Kon & Araya, 1998 Macroplia Brenske, 1898 Macropliopsis Lacroix, 1997 Macropodina Casey, 1915 Macropoidelimus Morón, 1983 Macropoides Guérin-Méneville, 1844 Macropopillia Ohaus, 1905 Macroprotaetia Mikšic, 1965 Macroretroides Bordat, Dellacasa & Dellacasa, 2000 Macroretrus Péringuey, 1908 Macroryctes Semenov-Tian-Shanskii & Medvedev, 1932 Macrotetrus Peringuey, 1908 Macrotina Strand, 1934 Macrotrigonophorus Miksic, 1974 Maculaphodius Gordon & Skelley, 2007 Madahoplia Lacroix, 1998 Madecorphnus Paulian, 1992 Madiana Ratcliffe & Romé, 2019 Madiniella Chalumeau & Gruner, 1976 Maechidinus Lea, 1919 Maechidius MacLeay, 1819 Maghrebotrogus Montreuil & Keith, 2022 Magniscarabaeus Hong, 1988 Mahafalyanus Lacroix, 1993 Makalaka Péringuey, 1908 Makoanus Lacroix, 1993 Maladera Mulsant, 1871 Malagoniella Martinez, 1961 Malaia Heller, 1891 Malaisius Arrow, 1941 Malawinophylla Lacroix, 2013 Mallotarsus Blanchard, 1850 Maneschetocia Delpont, 1996 Manjarivolo Paulian, 1974 Manodactylus Moser, 1919 Manodema Moser, 1907 Manonychus Moser, 1919 Manopus Castelnau, 1840 Manopus Laporte, 1840 Manoserica Moser, 1924 Marmarina Kirby, 1827 Marmylida Thomson, 1880 Marronus Coquerel, 1866 Martineziana Chalumeau & Özdikmen, 2006 Martinezidium Vaz-de-Mello, 2008 Mascarena Arrow, 1919 Masumotokoganea Hirasawa, 1992 Matashia Matsumura, 1938 Matthewsius Gunter & Weir, 2017 Mauromecistoplia Dombrow, 2002 Mausoleopsis Lansberge, 1882 Mawenzhena Alexis & Delpont, 2001 Mayataia Lacroix, 1993 Mazahuapertha Morón & Nogueira, 1998 Mazoe Péringuey, 1896 Mecaspidiellus Antoine, 1997 Mecaspidus Bourgoin, 1921 Mecinonota Kraatz, 1892 Mecopelidinota Mecopelidnota Bates, 1904 Mecynodes Mulsant, 1870 Mecynorhina Hope, 1837 Mecynorrhinella Marais & Holm, 1992 Medeotrogus Keith, 2001 Megaceras Hope, 1837 Megaceropsis Dechambre, 1976 Megacoryne Britton, 1987 Megalonitis Janssens, 1937 Megalorhina Westwood, 1847 Megalorrhina Meganthracophora Megapertha (Reitter, 1903) Megaphanaeus d'Olsoufieff, 1924 Megaphonia Schürhoff, 1933 Megarhopaea Britton, 1978 Megascarabaeus Megasoma Kirby, 1825 Megasybacodes Kakizoe, Maruyama & Masumoto, 2019 Megateloides Landin, 1974 Megatelus Reitter, 1892 Megatharsis Waterhouse, 1891 Megathopa Eschscholtz, 1822 Megathoposoma Balthasar, 1939 Megistophylla Burmeister, 1855 Megistophyllas Megistoplia Lacroix, 1997 Melanchroea Kraatz, 1900 Melanhyphus Fairmaire, 1881 Melanocamenta Brenske, 1899 Melanocanthon Halffter, 1958 Melanopopillia Linnaeus, 1980 Melanoptera Melanosa Mulsant & Rey, 1871 Melichus Péringuey, 1901 Melinesthes Kraatz, 1880 Melinopterus Mulsant, 1842 Mellissius Wollaston, 1869 Melolontha Fabricius, 1775 Melolontha Müller, 1764 Melolonthini Leach, 1819 Melolonthites Heer, 1847 Melolonthoides De Wailly, 1950 Meloroba Mendidaphodius Reitter, 1901 Mendidius Harold, 1868 Menigionyx Arrow, 1941 Menthophilus Mentophilus Castelnau, 1840 Mericserica Brenske, 1897 Meridioclita Krikken, 1982 Merinanus Lacroix, 1993 Merogyrus Gordon & Skelley, 2007 Meroloba Thomson, 1880 Mesaclopus Nikolajev, 1992 Mesomerodon Ohaus, 1905 Mesontoplatys Motschulsky, 1863 Mesorrhopa Mesoserica Brenske, 1897 Mesosternus Morón, 1987 Mesotrochalus Kolbe, 1914 Messyrhus Rakovič & Král, 1997 Mesydra Ponomarenko, 1977 Mesystoechus Waterhouse, 1878 Metacatharsius Paulian, 1939 Metadorodocia Machatschke, 1957 Metallesthes Kraatz, 1880 Metallophanaeus d'Olsoufieff, 1924 Metallopseudinca Valck Lucassen, 1933 Metanastes Arrow, 1911 Metapachylus Bates, 1890 Metaphileurus Kolbe, 1910 Metapogonia Lacroix, 2008 Metascelis Westwood, 1842 Metatrogus Britton, 1978 Metoporaphodius Frolov & Akhmetova, 2017 Mettalophanaeus Meurguesia Ruter, 1969 Michaeloplia Lacroix, 1997 Micraegialia Brown, 1931 Micranomala Arrow, 1911 Micrelaphinis Schoch, 1896 Micreuchilia Pouillaude, 1917 Microchalcothea Moser, 1910 Microchilus Blanchard, 1850 Microcoenus Britton, 1957 Microcopris Balthasar, 1958 Microcosmiomorpha Miksic, 1974 Microdilochrosis Jákl, 2009 Microdoris Burmeister, 1844 Microglycyphana Miksic, 1968 Microlomaptera Kraatz, 1885 Microlontha Petrovitz, 1967 Microogenius Gutiérrez, 1951 Micropeltus Blanchard, 1842 Micropeocila Micropertha Baraud, 1991 Microphileurus Kolbe, 1910 Microphylla Kraatz, 1890 Microplidus Péringuey, 1902 Microplus Burmeister, 1844 Micropoecila Kraatz, 1880 Microrhopaea Lea, 1920 Microrutela Bates, 1904 Microryctes Arrow, 1908 Microserica Brenske, 1894 Microsericaria Nikolajev, 1979 Microtermitodius Pittino, 2006 Microtertnitodius Pittino, 2006 Microteuchestes Landin, 1974 Microthops Microthopus Burmeister, 1855 Microthyrea Kraatz, 1882 Microtrochalus Brenske, 1900 Microvalgus Kraatz, 1883 Mictophileurus Ohaus, 1911 Miksicus Özdikmen & Turgut, 2009 Milatrogus Montreuil & Keith, 2022 Milichus Péringuey, 1901 Millotsis Bourgin, 1945 Miltotrogus Reitter, 1902 Mimadoretus Arrow, 1901 Mimaegialia Nikolajev, 2007 Mimela Kirby, 1823 Mimeoma Casey, 1915 Mimogeniates Martinez, 1964 Mimognorimus Kirkken Mimonthophagus Balthasar, 1963 Mimotribostethes Gutiérrez, 1949 Mimovalgus Arrow, 1944 Minidorysthetus Soula, 1998 Minisiderus Arrow, 1902 Minisiderus Endrodi, 1970 Minthophilus Minutoplia Lacroix, 1998 Miochaetopisthes Tangelder & Krikken, 1982 Miolachnosterna Wickham, 1914 Miotemna Lacordaire, 1856 Mireia Ruter, 1953 Miridiba Reitter, 1902 Mitracamenta Brenske, 1903 Mitracephala Thomson, 1859 Mitrophorus Burmeister, 1844 Mixigenus Thomson, 1859 Mnematidium Ritsema, 1888 Mnematium MacLeay, 1821 Modialis Fairmaire & Germain, 1860 Molynoptera Kraatz, 1897 Molynopteroides Antoine, 1989 Mongolrobeus Nikolajev, 2004 Monochelus Le Peletier & Serville, 1828 Monoplistes Lansberge, 1874 Monotropus Erichson, 1847 Monteitheolus Howden & Storey, 2000 Monteithocanthon Gunter & Weir, 2017 Morettius Roggero, Dierkens, Barbero & Palestrini, 2017 Moriaphila Kraatz, 1880 Morokia Janson, 1905 Moroniella Ramírez-Ponce, 2015 Moronius Grossi & Vaz-de-Mello, 2015 Moseriana Ruter, 1965 Mothon Semenov & Medvedev, 1927 Motuotrichius Huang & Chen, 2016 Mozartius Nomura & Nakane, 1951 Mucama Soula, 2002 Mucrencya Lacroix, 1993 Mucterophallus Poll, 1886 Musurgus Vauloger, 1898 Mycernus Broun, 1904 Mycterophallus Kraatz, 1886 Mycterophallus Neervoort Van De Poll, 1886 Mycterophalus Myloxena Berg, 1881 Myloxena Ohaus, 1909 Myloxenoides Martinez, 1975 Myoderma Dejean, 1833 Myodermides Ruter, 1964 Myodermidius Bourgoin, 1920 Myodermidius Moser, 1919 Myodermum Burmeister, 1840 Myrhessus Balthasar, 1955 Myrmecaphodius Martinez, 1952 Myrmecochilus Wasmann, 1900 Myrmecotonus Mann, 1914 Mysarus Petrovitz, 1962 Mystroceros Burmeister, 1842 Mzansica Perissinotto, 2020 Nagainokoganea Hirasawa, 1992 Namakwanus Scholtz & Howden, 1987 Namaphilus Deschodt & Davis, 2017 Nanaga Péringuey, 1902 Nanniscus Burmeister, 1844 Nannopopillia Kolbe, 1894 Nannovalgus Kolbe, 1904 Nanorhopaea Britton, 1978 Nanos Westwood, 1842 Nanotermitodius Howden, 2003 Napoa Stebnicka, 1999 Narycius Dupont, 1835 Navigator Moeseneder & Hutchinson, 2016 Nayarita Morón & Nogueira, 1998 Neagolius Koshantschikov, 1912 Nebulasilvius Deschodt & Scholtz, 2008 Nedymoserica Brenske, 1900 Nefoncerus Saylor, 1938 Nematophylla Arrow, 1906 Nematoserica Arrow, 1917 Neoblitopertha Baraud, 1991 Neocalaphodius Bordat, 1990 Neocamenta Burgeon, 1945 Neochaetopisthes Wasmann, 1918 Neoclita Perissinotto, 2017 Neoclithria Neervoort Van De Poll, 1886 Neoclitopa Lacroix, 1997 Neocnecus Sharp, 1878 Neocolobopterus Landin, 1974 Neocorvicoana Ratcliffe & Mico, 2001 Neocorynophyllus Carne, 1985 Neocymophorus Schein, 1954 Neodasygnathus Carne, 1957 Neodiapterina Dellacasa, 1986 Neodiapterna Dellacasa, 1984 Neoemadiellus Clement, 1986 Neogutierrezia Martinez, 1953 Neoheptaulacus Paulian & Villiers, 1941 Neoheteronyx Blackburn, 1890 Neohyphus Heller, 1896 Neomaladera Baraud, 1965 Neometanastes Yamaya, 2010 Neomystroceros Alexis & Delpont, 1998 Neonastes Carne, 1957 Neonitis Péringuey, 1901 Neopegylis Lacroix, 2015 Neophaedimus Lucas, 1870 Neophaedimus Schoch, 1894 Neophaenognatha Allsopp, 1983 Neopharis Neophoedimus Lucas, 1870 Neophyllotocus Blackburn, 1898 Neopsammodius Rakovič, 1986 Neorhyssemus Gordon & Pittino, 1992 Neorrhina Thomson, 1878 Neoryctes Arrow, 1908 Neosaproecius Branco, 1990 Neoscelis Schoch, 1897 Neoserica Brenske, 1894 Neosericania Miyake & Yamaya, 1994 Neosisyphus Müller, 1942 Neotemnorrhynchus Janssens, 1949 Neotrichaphodioides Dellacasa, Dellacasa & Skelley, 2010 Neotrichiorhyssemus Rakovič & Král, 1997 Neotrichonotulus Dellacasa, Gordon & Dellacasa, 2004 Neovalgus Miyake, 1985 Nepaloserica Frey, 1965 Nephrodophus Sharp, 1873 Nephrodopus Sharp, 1873 Neprodopus Neptunides Thomson, 1879 Nepytis Erichson, 1842 Nesadmiralitia Prokofiev, 2016 Neso Blackburn, 1898 Nesocanthon Pereira & Martínez, 1956 Nesohoplia Scott, 1912 Nesosisyphus Vinson, 1946 Nesovinsonia Martinez & Pereira, 1958 Netocia Costa, 1852 Netociomima Mikšic, 1963 Nettelislasia Martinez, 1952 Neuquenodactylus Smith & Mondaca, 2015 Neuroserica Brenske, 1900 Nialaphodius Kolbe, 1908 Nialosternus Hollande & Therond, 1998 Nialus Mulsant & Rey, 1869 Nigroplia Dombrow, 2002 Nigrotrichia Matsumoto, 2016 Nimbacola Paulian, 1970 Nimbus Mulsant & Rey, 1869 Niphetophora Kraatz, 1883 Niphobleta Kraatz, 1880 Niponoprotaetia Miksic, 1966 Nipponaphodius Nakane, 1959 Nipponoagoliinus Ochi & Kawahara, 2001 Nipponoserica Nomura, 1973 Nipponovalgus Sawada, 1941 Nitiocellus Branco, 2010 Nitorellus Britton, 1957 Niuailan Allsopp & Hutchinson, 2020 Nobiellus Dellacasa & Dellacasa, 2005 Nobius Mulsant & Rey, 1869 Nolicus Petrovitz, 1962 Nosphisthis Blackburn, 1898 Notheterogomphus Prell, 1912 Nothophanus Heller, 1896 Notocaulus Quedenfeldt, 1884 Novapus Sharp, 1875 Nselenius (Holm & Perissinotto, 2004) Nudipleurus Lebis, 1960 Nuiba Prokofiev, 2016 Nunoidium Vaz-de-Mello, 2008 Nyassinus Westwood, 1879 Obaphodius Bordat, 1988 Oberthuroplia Lacroix, 1998 Ochicanthon Vaz-de-mello, 2003 Ochodaeus Serville, 1825 Octocollis Moeseneder & Hutchinson, 2012 Octoplasia Brenske, 1892 Odochilus Harold, 1877 Odontacrossus Dellacasa, Král, Dellacasa & Bordat, 2014 Odontolochus Schmidt, 1916 Odontoloma Boheman, 1857 Odontolytes Koshantschikov, 1916 Odontoplia Fairmaire, 1897 Odontoprotaetia Mikšic, 1963 Odontopsammodius Gordon & Pittino, 1992 Odontorrhina Burmeister, 1842 Odontothyrea Schoch, 1897 Odontotonyx MacLeay, 1871 Odontoxyomus Král, Rakovič & Mencl, 2015 Odontria White, 1846 Oedanomerus Sehnal, 2018 Oedanomerus Waterhouse, 1875 Oedichira Burmeister, 1855 Oedipovalgus Kolbe, 1897 Oficanthon Paulian, 1985 Okeurophorus Oligophylla Kraatz, 1894 Omaloplia Schönherr, 1817 Omocnemus Schein, 1958 Omocrates Burmeister, 1844 Oncerus LeConte, 1856 Oncochirus Kolbe, 1891 Oncosterna Thomson, 1880 Oniticellus Dejean, 1821 Oniticellus LePeletier & Serville, 1828 Onitis Fabricius, 1798 Onochaeta Erichson, 1847 Onochirus Kolbe, 1891 Onoreidium Vaz-de-Mello, 2008 Onorius Frolov & Vaz-de-Mello, 2015 Ontherus Erichson, 1847 Onthobium Reiche, 1860 Onthohpagus Onthophagini Burmeister, 1846 Onthophagus Latreille, 1802 Onthophahus Onychionyx Arrow, 1914 Onychoserica Moser, 1916 Onychosophrops Frey, 1972 Onychotarsus Schoch, 1897 Onychothecus Boucomont, 1912 Oocamenta Péringuey, 1904 Oogenius Solier, 1851 Ophropyx Britton, 1987 Opistocus Britton, 1957 Oplognathus MacLeay, 1819 Oplostomus MacLeay, 1838 Opsitocus Britton, 1957 Orammoecius Král, Rakovič & Mencl, 2016 Oreoderus Burmeister, 1842 Oreogenoplia Lacroix, 1998 Oreopotosia Medvedev, 1964 Oreotrogus Kolbe, 1897 Oreovalgus Kolbe, 1904 Orizabus Fairmaire, 1878 Orocanthus Endrödi, 1983 Oroclita Krikken, 1982 Orodaliscoides Schmidt, 1913 Orodaliscus Reitter, 1900 Oromus Mulsant & Rey, 1869 Orphnus MacLeay, 1819 Orrodoretus Ohaus, 1912 Orsilochus Burmeister, 1847 Ortheterogomphus Prell, 1912 Orthocavonus Carne, 1957 Orthoserica Brenske, 1900 Orubesa Reitter, 1895 Oruscatuen Bates, 1870 Oruscatus Bates, 1870 Oryctes Hellwig, 1798 Oryctes Illiger, 1798 Oryctoantiquus Ratcliffe, Smith & Erwin, 2005 Oryctoderinus Endrödi, 1978 Oryctoderus Boisduval, 1835 Oryctomorphus Guérin-Méneville, 1830 Oryctophileurus Kolbe, 1910 Oryctus Oscarinus Gordon & Skelley, 2007 Osmanius Branco & Baraud, 1988 Osmoderma Le Peletier & Serville, 1828 Othnonius Olliff, 1890 Otoclinius Brenske, 1896 Otoclinus Brenske, 1896 Otophorus Mulsant, 1842 Ottokelleria D'Andretta & Martinez, 1957 Outenikwanus Scholtz & Howden, 1987 Outeniqua Péringuey, 1902 Ovomanonychus Costa, Cherman & Iannuzzi, 2020 Ovoserica Frey, 1968 Oxyadoretus Arrow, 1917 Oxyataenius Dellacasa & Stebnicka, 2001 Oxychirus Quedenfeldt, 1888 Oxycorythus Solsky, 1876 Oxygrylius Casey, 1915 Oxyligyrus Arrow, 1908 Oxyomoides Dellacasa, Dellacasa & Gordon, 2016 Oxyomus Dejean, 1833 Oxyomus Kirby & Spence, 1828 Oxypelta Pouillaude, 1920 Oxyperatex Krikken, 1982 Oxyrraphia Oxyserica Brenske, 1899 Oxysternon Castelnau, 1840 Oxysternon Laporte, 1840 Oxythyrea Mulsant, 1842 Ozodius Skelley, 2007 Pachacama Soula, 2006 Pachnessa Brenske, 1894 Pachnoda Burmeister, 1842 Pachnodoides Alexis & Delpont, 2002 Pachnotosia Reitter, 1891 Pachrodema Blanchard, 1850 Pachycamenta Brenske, 1895 Pachychilecamenta Brenske, 1903 Pachycnema Le Peletier & Serville, 1828 Pachycolus Erichson, 1847 Pachydema Laporte, 1832 Pachydemocera Reitter, 1902 Pachyderoserica Moser, 1920 Pachygastra Germar, 1848 Pachylomera Griffith & Pidgeon, 1831 Pachylomera Kirby, 1832 Pachyorphnus Paulian, 1951 Pachyoryctes Arrow, 1908 Pachypoides Fairmaire, 1884 Pachyprotaetia Mikšic, 1965 Pachypus Dejean, 1821 Pachyrhinadoretus Ohaus, 1912 Pachyserica Brenske, 1897 Pachysoma MacLeay, 1821 Pachystethus Blanchard, 1850 Pachytephraea De Palma & Malec, 2020 Pachytricha Hope, 1841 Pacuvia Curtis, 1845 Paediovalgus Kolbe, 1904 Pagonotarsus Burmeister, 1842 Paidiovalgus Palacephala Lacroix, 1999 Palaeophileurus Kolbe, 1910 Paleira Reiche, 1871 Paleopragma Thomson, 1880 Paleotrichius Poinar, 2011 Palnia Stebnicka, 1985 Panelus Lewis, 1895 Panglaphyra Kraatz, 1880 Panotrogus Reitter, 1902 Pantodinus Burmeister, 1847 Pantolia Burmeister, 1842 Papuana Arrow, 1911 Parabodilus Hollande & Thérond, 1998 Parabyrsopolis Ohaus, 1915 Paracamenta Péringuey, 1904 Paracanthon Balthasar, 1938 Paraceratochelus Dombrow, 2002 Parachilia Burmeister, 1842 Parachlorota Soula, 2002 Parachorius Harold, 1873 Parachrysina Bates, 1888 Paraclitopa Waterhouse, 1875 Paracoptochirus Balthasar, 1963 Paracorythoderus Wasmann, 1918 Paracotalpa Ohaus, 1915 Paracrossidius Balthasar, 1932 Paracryptocanthon Howden & Cook, 2002 Paracyclidius Howden, 1971 Paradeloparius Landin, 1959 Paradendropaemon Edmonds, 1972 Paradidactylia Balthasar, 1937 Paradorodocia Machatschke, 1957 Paradorysthetus Soula, 1998 Paraegidium Vulcano, Pereira & Martinez, 1966 Paragnorimus Becker, 1910 Paragymnopleurus Shipp, 1897 Paraheteronyx Moser, 1924 Paraheterosternus Morón, 1983 Paraholotrichia Lacroix & Coache, 2017 Parahyboma Paulian, 1938 Paraixodina Roggero, Barbero & Palestrini, 2015 Paraliatongus Balthasar, 1963 Paralleucosma Antoine, 1989 Parallidiostoma Ocampo & Colby, 2009 Paramacraspis Ohaus, 1915 Paramaladera Nomura, 1974 Paramicroplus Lacroix, 1998 Parammoecius Seidlitz, 1891 Paramorphochelus Lacroix, 1997 Paranelaphinis Antoine, 1988 Paranimbus Schmidt, 1913 Paranodon Cockerell, 1905 Paranomala Casey, 1915 Parapeltastes Ricchiardi, Perissinotto & Clennell, 2004 Parapetiia Martinez, 1961 Paraphacosomoides Balthasar, 1969 Paraphanaemorphus Paraphileurus Endrödi, 1978 Paraphytus Harold, 1877 Parapilinurgus Arrow, 1910 Paraplectrone Mikšić, 1985 Paraplesiataenius Chalumeau, 1992 Parapoecilophila Hauser, 1904 Paraprotaetia Moser, 1907 Parapsammodius Gordon & Pittino, 1992 Parapsammodius Verdu, Stebnicka & Galante, 2006 Paraptenomela Soula, 2002 Parapucaya Prell, 1934 Pararhabdotis Kraatz, 1899 Pararhopaea Blackburn, 1911 Pararhynchocephala Paulian, 1991 Pararhyssemus Balthasar, 1955 Parascatonomus Paulian, 1932 Paraschizognathus Ohaus, 1904 Parasciton Britton, 1990 Paraserica Reitter, 1896 Parastasia Westwood, 1842 Parastraella Antoine, 2005 Parasymmela Pacheco, Monné, Vaz-de-Mello & Ahrens, 2022 Parataenius Balthasar, 1961 Paratelaugis Ohaus, 1915 Parateuchus Shipp, 1895 Parathyridium Ohaus, 1915 Paratrichaphodius Bordat, 2013 Paratrichius Janson, 1881 Paratriodonta Baraud, 1962 Parautoserica Lacroix, Coache & Filippi, 2022 Paraxeloma Holm, 1988 Parbomonyx Ohaus, 1915 Pardalosus Gordon & Skelley, 2007 Parelaphinis Marais & Holm, 1989 Parentius Zunino, 1979 Parepixanthis Kraatz, 1893 Pareriesthis Moser, 1918 Pareteronychus Prell, 1934 Pareuchilia Kraatz, 1880 Parhomonyx Ohaus, 1915 Parhoplognathus Ohaus, 1915 Parisolea Bates, 1888 Parisomorphus Schaufuss, 1890 Parodochilus Rakovic, 1997 Paroniticellus Balthasar, 1963 Paronthobium Paulian, 1984 Paronthophagus Balthasar, 1959 Paronyx Britton, 1990 Paroplia Fairmaire, 1903 Parorphnus Paulian, 1951 Paroryctoderus Dechambre, 1994 Parthenoserica Brenske, 1899 Parvuhowdenius Deschodt & Scholtz, 2008 Parvulomaladera Ahrens, 1995 Pasaphylla Péringuey, 1904 Passaliolla Balthasar, 1945 Patatra Soula, 2008 Paulianellus Balthasar, 1938 Paulosawaya Martinez & D'andretta, 1956 Peckolus Scholtz & Howden, 1987 Pectinichelus Ballion, 1870 Pectinosoma Arrow, 1913 Pedaria Laporte, 1832 Pedaridium Harold, 1868 Pedinorrhina Kraatz, 1880 Pedinotrichia Matsumoto, 2016 Pegylis Erichson, 1847 Pelidnota MacLeay, 1819 Pelidnotites Cockerell, 1920 Peltonotus Burmeister, 1847 Peltotrichus Howden, 1968 Penalus Paulian, 1985 Penrithochilus Krikken, 1990 Pentacoryna Moser, 1926 Pentanomala Ohaus, 1919 Pentaphylla De Wailly, 1950 Pentecamenta Brenske, 1896 Pentelia Brenske, 1891 Penthima Kraatz, 1892 Pentodina Endrödi, 1968 Pentodon Hope, 1837 Pentodontoschema Péringuey, 1901 Peotoxus Krikken, 1983 Peperonota Westwood, 1847 Pereiraidium Vaz-de-Mello, 2008 Pericamenta Péringuey, 1904 Periclitopa Brenske, 1896 Pericoptus Burmeister, 1847 Periphanesthes Kraatz, 1880 Periproctoides Lacroix, 2006 Periproctus Kolbe, 1910 Periserica Brenske, 1898 Perissosoma Waterhouse, 1875 Peritrichia Burmeister, 1844 Perrindema Lacroix, 1997 Peruquime Mondaca & Valencia, 2016 Petinopus Blackburn, 1898 Petrovitzia Mikšić, 1965 Petrovitzius Rakovič, 1979 Petroyitzia Miksic, 1965 Phaeadoretus Reitter, 1903 Phaeaphodius Reitter, 1892 Phaedimus Waterhouse, 1841 Phaedotrodus Phaedotrogus Paulian, 1985 Phaenognatha Hope, 1841 Phaenomeris Hope, 1833 Phalacronothus Motschulsky, 1859 Phalangogonia Burmeister, 1844 Phalangonyx Reitter, 1889 Phalangosoma Quedenfeldt, 1884 Phalops Erichson, 1848 Phanaeomorphus Balthasar, 1935 Phanaeus MacLeay, 1819 Phanaresthes Phaneresthes Kraatz, 1894 Pharaonus Blanchard, 1850 Pharaphodius Reitter, 1892 Phiara Brenske, 1897 Philacelota Heller, 1900 Phileucourtus Dechambre, 2008 Phileurus Latreille, 1807 Philgertia Zídek, 2021 Philistina MacLeay, 1838 Philochlaenea Hope, 1839 Philochloenia Dejean, 1833 Philoscaptus Brèthes, 1919 Philoserica Brenske, 1897 Phobetus LeConte, 1856 Pholeoaphodius Bordat, 2009 Pholidochris Kolbe, 1894 Phonopleurus Moser, 1919 Phonotaenia Kraatz, 1880 Phorine Britton, 1987 Phornus Paulian, 1951 Photyna Brenske, 1897 Phoxomela Schaum, 1844 Phoxomeloides Schoch, 1898 Phycochus Broun, 1886 Phycocus Broun, 1883 Phylliocephala Blackburn, 1890 Phyllocamenta Moser, 1918 Phyllochlaenia Blanchard, 1846 Phyllococerus Waterhouse, 1876 Phyllognathus Eschscholtz, 1830 Phyllopertha Stephens, 1830 Phyllophaga Harris, 1826 Phylloputha Kirby & Spence, 1828 Phylloserica Brenske, 1899 Phyllotocidium Blackburn, 1898 Phyllotocus Fischer von Waldheim, 1823 Phyllotrochalus Brenske, 1900 Phymatopterix Phytholaema Blanchard, 1850 Pichica Soula, 2002 Pileotrichius Bourgoin, 1921 Pilinopyga Kraatz, 1888 Pilinurgus Burmeister, 1842 Pilleriana Baraud, 1991 Pimelomera Moser, 1924 Pimelopus Erichson, 1842 Pinacopodius Branco, 1991 Pinacotarsus Harold, 1875 Piscoperus Ratcliffe & Giraldo, 2014 Pittinius Rakovič & Král, 1997 Placodidus Péringuey, 1900 Plaesiorrhina Westwood, 1842 Plaesiorrhinella Plaesioserica Brenske, 1899 Plagiogonus Mulsant, 1842 Planolinellus Dellacasa & Dellacasa, 2005 Planolinoides Dellacasa & Dellacasa, 2005 Planolinus Mulsant & Rey, 1869 Planontherus Genier, 1996 Planophileurus Chapin, 1932 Platychelus Burmeister, 1844 Platycoelia Dejean, 1833 Platyderides Schmidt, 1916 Platygenia MacLeay, 1819 Platygeniops Krikken, 1978 Platynocephalus Westwood, 1854 Platyonitis Janssens, 1942 Platyphileurus Ohaus, 1910 Platyrutela Bates, 1888 Platysodes Westwood, 1874 Platytomus Mulsant, 1842 Plectris Le Peletier & Serville, 1828 Plectrodes Horn, 1867 Plectrone Wallace, 1867 Pleistophylla Péringuey, 1904 Pleophylla Erichson, 1847 Plesiocamenta Lacroix & Montreuil, 2019 Plesiopalacephala Lacroix, 2006 Plesiorutela Jameson, 1997 Plesioserica Brenske, 1900 Plesiosternus Morón, 1983 Pleuraphodius Schmidt, 1913 Pleuronitis Lansberge, 1875 Pleuronota Kraatz, 1892 Pleurophorus Mulsant, 1842 Plinurgus Plochilia Fairmaire, 1896 Plochiliana Ruter, 1978 Plocovalgus Kolbe, 1904 Plotopuserica Brenske, 1899 Plusioserica Brenske, 1899 Plusiotus Podalgus Burmeister, 1847 Podischnus Burmeister, 1847 Podolasia Harold, 1869 Podopholis Moser, 1915 Podopogonus Moser, 1917 Podoserica Breit, 1912 Podostena Howden, 1997 Podotenoides Paulian, 1942 Podotenus Schmidt, 1913 Podovalgus Arrow, 1910 Poecilophana Kraatz, 1895 Poecilopharis Kraatz, 1880 Poecilophilides Kraatz, 1898 Pogoniotarsus Kraatz, 1880 Pogonopus Arrow, 1910 Pogonotarsus Burmeister, 1842 Pollaplonyx Waterhouse, 1875 Polybathes Kirby, 1827 Polyenaria De Wailly, 1950 Polylamina Hardy, 1974 Polyphylla Harris, 1841 Polyphyllum Blanchard, 1850 Polyplastus Janson, 1888 Polystalactica Kraatz, 1882 Popillia Dejean, 1821 Popillia Serville, 1825 Porphyronota Burmeister, 1842 Potosiomima Praogolofa Bates, 1892 Prigenia Mohnike, 1871 Prionoryctes Arrow, 1911 Priscorrhina Krikken, 1984 Priscyparus Skelley, 2021 Priska Jákl, 2018 Pristerophora Harold, 1869 Proagoderus Lansberge, 1883 Proagopertha Reitter, 1903 Proagosternus Blanchard, 1850 Problerhinus Deyrolle, 1864 Proborhinus Britton, 1988 Prochelyna Erichson, 1847 Proctophanes Harold, 1861 Prodontria Broun, 1904 Prodoretus Brenske, 1893 Progastor Thomson, 1880 Promacropoides Sigwalt, 1987 Prophaenognatha Bai, Ren & Yang, 2011 Propomacrus Newman, 1836 Propomacus Prosphileurus Kolbe, 1905 Protaetia Burmeister, 1842 Protaetiola Mikšic, 1963 Protaetiomorpha Mikšić, 1968 Protaetiophana Heller, 1896 Protelura Britton, 1987 Proteroscarabeus Grabau, 1923 Proteroschiza Brenske, 1896 Protochilus Krikken, 1976 Protoclita Krikken, 1978 Protoclitopa Decelle, 1979 Protoleura Britton, 1987 Protopalnia Nikolajev, 2007 Prototrox Nikolajev, 2000 Psammaegialia Nikolajev, Wang & Zhang, 2014 Psammodaphodius Endrödi, 1976 Psammodius Fallen, 1807 Psammoporus Thomson, 1859 Psammorpha Stebnicka, 1994 Psednoserica Brenske, 1899 Pseudachloa Péringuey, 1904 Pseudacrossus Reitter, 1892 Pseudadoretus Semenov, 1890 Pseudagenius Heller, 1923 Pseudagoliinus Pseudagolius Schmidt, 1913 Pseudalleucosma Antoine, 1989 Pseudanatona Kraatz, 1895 Pseudanthracophora Kraatz, 1898 Pseudastoxenus Bourgoin, 1921 Pseudataenius Brown, 1927 Pseudenaria Fairmaire, 1901 Pseudencya Lacroix, 1991 Pseudepilissus Pseudepixanthis Kraatz, 1880 Pseuderytus Hollande & Thérond, 1998 Pseudesymus Orbigny, 1896 Pseudeuryomia Kraatz, 1894 Pseudheterochelus Schein, 1959 Pseudholophylla Blackburn, 1911 Pseudignambia Paulian, 1984 Pseudinca Kraatz, 1880 Pseudisonychus Frey, 1971 Pseudoagoliinus Bordat, 1994 Pseudoahermodontus Dellacasa & Dellacasa, 1997 Pseudoanticheiroides Soula, 1998 Pseudoblackburneus Bordat, 2016 Pseudoblitopertha Keith, 2000 Pseudocalopotosia Miksic, 1966 Pseudocanthon Bates, 1887 Pseudocavonus Blackburn, 1890 Pseudocetonischema Mikšic, 1965 Pseudochalcothea Ritsema, 1882 Pseudochalcotheomima Mikšić, 1985 Pseudochironitis Ferreira, 1977 Pseudochlorota Ohaus, 1905 Pseudoclinteria Kraatz, 1882 Pseudoclithria Neervoort Van De Poll, 1886 Pseudocoelotrachelus Dellacasa, Dellacasa & Gordon, 2013 Pseudocongella Moser, 1924 Pseudocopris Ferreira, 1960 Pseudocotalpa Hardy, 1971 Pseudocymophorus Schein, 1954 Pseudocyrtolophus Frolov & Montreuil, 2019 Pseudodiceros Mikšić, 1974 Pseudodicrania Gutiérrez, 1950 Pseudodiplotaxis Nonfried, 1894 Pseudodorysthetus Soula, 1998 Pseudodrepanocanthus (Bordat, 1993) Pseudoeuselates Miksic, 1974 Pseudogeniates Ohaus, 1910 Pseudogonaphodiellus Dellacasa, Gordon & Dellacasa, 2007 Pseudoheliocopris Ferreira, 1970 Pseudoheptaulacus Dellacasa & Dellacasa, 2000 Pseudoheteronyx Blackburn, 1892 Pseudoheterophana Allard, 1990 Pseudoholophylla Blackburn, 1911 Pseudohomonyx Arrow, 1908 Pseudohomonyx apoensis Miyake & Yamaya, 1997 Pseudohomonyx belitungensis Miyake & Yamaya, 1997 Pseudohomonyx borneensis Arrow, 1908 Pseudohomonyx continentalis Yang & Pathomwattananurak, 2023 Pseudohomonyx crassus Arrow, 1911 Pseudohomonyx extradentatus Miyake & Yamaya, 1997 Pseudohomonyx javanus (Burmeister, 1847) Pseudohomonyx loksadoensis Jákl & Zídek, 2017 Pseudohomonyx luzonicus Miyake & Yamaya, 2000 Pseudohomonyx mindroensis Miyake & Yamaya, 2000 Pseudohomonyx minor Jákl & Zídek, 2017 Pseudohomonyx negros Miyake & Yamaya, 1997 Pseudohomonyx palavanus Prell, 1912 Pseudohomonyx samarensis Miyake & Yamaya, 1997 Pseudohomonyx similis Miyake & Yamaya, 1999 Pseudohypaspidius Soula, 1998 Pseudoisonychus Frey, 1971 Pseudoliogenys Moser, 1919 Pseudolontha Fairmaire, 1897 Pseudomacraspis Ohaus, 1903 Pseudomaladera Nomura, 1974 Pseudomecinonota Mikšic, 1972 Pseudomesystoechus Ohaus, 1912 Pseudomicroplus Lacroix, 1998 Pseudomothon Pittino, 1984 Pseudoneptunides Csiki, 1909 Pseudonetocia Medvedev, 1964 Pseudonthobium Paulian, 1984 Pseudonthophagus Balthasar, 1959 Pseudooxythyrea Baraud, 1985 Pseudopachydema Balthasar, 1930 Pseudopanotrogus Petrovitz, 1969 Pseudopantrogus Petrovitz, 1969 Pseudoparoplia Lacroix, 1997 Pseudopectinosoma Katovich, 2011 Pseudopedaria Felsche, 1904 Pseudoperiproctus Keith & Reichenbach, 2018 Pseudopharaphodius Bordat, 1988 Pseudopholis Duvivier, 1892 Pseudopilinurgus Moser, 1918 Pseudopodotenus Dellacasa, 1990 Pseudoptenomela Soula, 2002 Pseudopygora Kraatz, 1897 Pseudorhomborrhina Miksic, 1967 Pseudorphnus Benderitter, 1913 Pseudoryctes Sharp, 1873 Pseudorycytes Pseudosaproecius Balthasar, 1941 Pseudosaprosites Balthasar, 1972 Pseudoscaptobius Krikken, 1976 Pseudoschizognathus Ohaus, 1904 Pseudoschizonycha Keith, 2006 Pseudoserica Guérin-Méneville, 1830 Pseudosericania Kobayashi, 1980 Pseudostegopterus Ricchiardi, 2015 Pseudostereomera Bordat & Howden, 1995 Pseudosymmachia Dalle Torre, 1912 Pseudosyrichthus Péringuey, 1901 Pseudotephraea Kraatz, 1882 Pseudotetraclipeoides Dellacasa, Dellacasa & Gordon, 2015 Pseudoteuchestes Dellacasa, 1986 Pseudothyridium Soula, 2002 Pseudotocama Keith, 2008 Pseudotorynorrhina Mikšić, 1967 Pseudotrematodes Jacquelin Du Val, 1860 Pseudotrigonocnemis Keith, 2007 Pseudotrochalus Quedenfeldt, 1884 Pseudourbania Mikšic, 1965 Pseudouteniqua Schein, 1959 Pseudoxyomus Petrovitz, 1962 Pseustophylla Péringuey, 1904 Psileterogomphus Prell, 1912 Psilocnemis Burmeister, 1842 Psilodontria Broun, 1895 Psilonychus Burmeister, 1855 Psilopholis Brenske, 1892 Psuedotephraea Kraatz, 1882 Ptenomela Bates, 1888 Pterobius Murayama, 2010 Pterobius itiokai Murayama, 2010 Pterothochaetes Ptychodestes Ptychodesthes Kraatz, 1883 Ptychophorus Burmeister & Schaum, 1841 Ptyophis Redtenbacher, 1868 Puapana Heller, 1913 Pubinus Mulsant & Rey, 1869 Pucaya Ohaus, 1910 Puelchesia Ocampo & Smith, 2006 Pukupuku Muramoto, 2006 Pulgilotrichius Krikken Pusiodactylus Smith, 2008 Pycnopanelus Arrow, 1931 Pycnoschema Thomson, 1859 Pygora Burmeister, 1842 Pygovalgus Kolbe, 1884 Pyronota Boisduval, 1835 Pyropotosia Reitter, 1898 Pyrrhopoda Kraatz, 1880 Qingaphodius Král, 1997 Rabula Péringuey, 1902 Raceloma Thomson, 1877 Radama Fairmaire, 1895 Rakovicius Pittino, 2006 Ramilia Kolbe, 1894 Ravautiana Lacroix, 1991 Raysymmela Saylor, 1947 Rectoscutaria Schein, 1958 Redotus Fairmaire, 1860 Reineria Mikšić, 1968 Relictonthophagus Kabakov, 1979 Renaudiana Lacroix, 1993 Renaudius Balthasar, 1951 Reneria Renorphnus Frolov & Montreuil, 2009 Repsimus MacLeay, 1819 Rhabdopholis Burmeister, 1855 Rhabdotis Burmeister, 1842 Rhabdotops Krikken, 1981 Rhachistoplia Lacroix, 1997 Rhadinolontha Arrow, 1930 Rhadinotaenia Kraatz, 1900 Rhagopteryx Burmeister, 1842 Rhamphadoretus Ohaus, 1912 Rhamphorrhina Klug, 1855 Rhinacosmus Kraatz, 1895 Rhinarion Ruter, 1965 Rhinaspis Perty, 1833 Rhamphadoretus Ohaus, 1912 Rhamphorrhina Klug, 1855 Rhinacosmus Kraatz, 1895 Rhinarion Ruter, 1965 Rhinaspis Perty, 1833 Rhinocerotopsis Maruyama, 2009 Rhinocoeta Burmeister, 1842 Rhinyptia Burmeister, 1844 Rhizogeniates Ohaus, 1909 Rhizoplatodes Péringuey, 1901 Rhizoplatys Westwood, 1841 Rhizoproctus Kolbe, 1894 Rhizotrogus Latreille, 1825 Rhodaphodius Adam, 1994 Rhomborhina Hope, 1837 Rhomborrhina Rhomborrhine Rhopaea Erichson, 1847 Rhynchadoretus Ohaus, 1913 Rhynchapogonia Lacroix, 2008 Rhynchocephala Fairmaire, 1883 Rhynchoserica Burgeon, 1942 Rhyncoserica Burgeon, 1942 Rhyncosymmela Frey, 1974 Rhyparus Agassiz, 1846 Rhyparus Westwood, 1845 Rhysothorax Bedel, 1911 Rhyssemodes Reitter, 1892 Rhyssemorphus Clouët, 1900 Rhyssemus Mulsant, 1842 Rhyxicephalus Moser, 1921 Rhyxiderma Moser, 1924 Rhyxiphloea Burmeister, 1842 Rigoutorum Hutchinson & Moeseneder, 2022 Rimuloplia Lacroix, 1997 Robinsonelliana Lacroix, 1996 Ronphus Paulian, 1951 Rubilepis De Wailly, 1950 Rubrohyboma Paulian, 1939 Rufotrichia Matsumoto, 2016 Rugaphodius Gordon & Skelley, 2007 Rugopertha Machatschke, 1957 Rutela Latreille, 1802 Rutelarcha Waterhouse, 1874 Rutelisca Bates, 1888 Rutelorcytes Arrow, 1908 Ruteloryctes Arrow, 1908 Ruteraetia Krikken, 1980 Rykanes Minck, 1919 Rykanoryctes Minck, 1919 Ryteloryctes Sabatinellius Montreuil & Lacroix, 2020 Sabatinelloplia Lacroix, 1998 Saccharoscaptus Morón & Grossi, 2015 Sahlbergianus Dellacasa & Dellacasa, 2000 Saphobiamorpha Brookes, 1944 Saphobius Sharp, 1873 Saprolochus Stebnicka & Galante, 2007 Saprositellus Balthasar, 1967 Saprosites Redtenbacher, 1857 Saprosotellus Balthasar, 1967 Saprosties Redtenbacher, 1858 Saprovisca Stebnicka, 1993 Saprus Blackburn, 1904 Sariangus Rakovič & Mencl, 2011 Sarophorus Erichson, 1847 Sarothermus Blackburn, 1907 Sarothromerus Blackburn, 1907 Saulostomus Waterhouse, 1878 Sauvagesinella Paulian, 1934 Sayloria Frey, 1975 Scabrostomus Gordon & Skelley, 2007 Scapanes Burmeister, 1847 Scapanoclypeus Evans, 1987 Scaphorhina Quedenfeldt, 1884 Scaphorhinadoretus Ohaus, 1912 Scaptobius Schaum, 1841 Scaptocnemis Péringuey, 1901 Scaptophilus Burmeister, 1847 Scarabaeolus Balthasar, 1965 Scarabaes Scarabaeus Linnaeus, 1758 Scatimus Erichson, 1847 Scatonomus Erichson, 1835 Scatrichus Génier & Kohlmann, 2003 Sceliages Westwood, 1837 Scelidothrix Dombrow, 2001 Scelophysa Burmeister, 1844 Schaefferellus Gordon & Skelley, 2007 Scheinia Ruter, 1957 Schismatocera Gautier Des Cottes, 1872 Schizadoretus Arrow, 1917 Schizochelus Blanchard, 1850 Schizognathus Fischer Von Waldheim, 1823 Schizonycha Dejean, 1833 Schizorhina Kirby, 1825 Schochidia Berg, 1898 Scitala Erichson, 1842 Scitaloides Britton, 1990 Sciton Blackburn, 1892 Scybalocanthon Martinez, 1948 Scybalophagus Martinez, 1953 Scythrodes Broun, 1886 Scythropesthes Kraatz, 1880 Sebakwe Péringuey, 1904 Sebaris Castelnau, 1840 Sebaris Laporte, 1840 Selaserica Brenske, 1897 Selomothus Fairmaire, 1891 Selviria Stebnicka, 1999 Semanopterus Hope, 1847 Semanopterus angustatus Blackburn, 1888 Semanopterus calabyi Carne, 1957 Semanopterus kingstoni Reid & Tees, 2023 Semanopterus leai (Blackburn, 1897) Semanopterus rectangulus Blackburn, 1895 Semanopterus solidus (Burmeister, 1847) Semanopterus subcostatus (Castelnau, 1840) Semanopterus tricostatus Blackburn, 1896 Semienaria De Wailly, 1950 Serica MacLeay, 1819 Sericania Motschulsky, 1860 Sericaria Motschoulsky, 1860 Sericesthis Boisduval, 1835 Sericini Hope, 1837 Sericoides Guérin-Méneville, 1839 Sericospilus Sharp, 1882 Serraphodius Kabakov, 1996 Serrophorus Balthasar, 1935 Setiserica Miyake, 2002 Setodius Gordon & Skelley, 2007 Setylaides Stebnicka, 1994 Seudinca Shangaia Lucas, 1920 Siamaphodius Masumoto, 1991 Siamophylla Keith, 2005 Sicardia Reitter, 1896 Sigorus Mulsant & Rey, 1869 Silluvia Landin, 1949 Silvaphilus Roets & Oberlander, 2010 Silvinha Vaz-de-Mello, 2008 Simogonious Harold, 1871 Simogonius Harold, 1871 Simorrhina Kraatz, 1886 Sinaphodius Červenka, 1994 Sinapisoma Boucomont, 1928 Singhala Blanchard, 1851 Singhalopotosia Miksic, 1987 Sinodiapterna Dellacasa, 1986 Sinodrepanus Simonis, 1985 Sinoliocola Alexis & Delpont, 1998 Sinoplia Medvedev, 1952 Siralus Dechambre, 1998 Sissantobius Ritsema, 1888 Sisyphe Latreille, 1806 Sisyphus Latreille, 1807 Sisyphus Wiedemann, 1823 Sitiphus Fairmaire, 1894 Skelleyanus Dellacasa, Dellacasa, Gordon & Stebnicka, 2011 Smicorhina Westwood, 1847 Socotraproctus Král, Sehnal & Bezdĕk, 2012 Somalibia Lansberge, 1882 Somatoserica Brenske, 1899 Sophrops Fairmaire, 1887 Sorocha Soula, 2006 Spaeromorphus Spaniolepis Kolbe, 1894 Sparrmannia Castelnau, 1840 Spathoschiza Arrow, 1902 Spelaiorrhina Lansberge, 1886 Sphaeraphodius Kakizoe, Jiang & Wang, 2021 Sphaeromrophus Sphaerorutela Jameson, 1997 Sphaeroscelis Burmeister, 1855 Sphaerotrochalus Brenske, 1900 Sphecoserica Brenske, 1899 Spherocanthon Olsoufieff, 1947 Sphinctovalgus Kolbe, 1904 Sphodroxia Kraatz, 1890 Sphyrocallus Sharp, 1877 Spilophorus Schaum, 1848 Spilophorus Westwood, 1848 Spilopopillia Kraatz, 1892 Spinanomala Ohaus, 1910 Spinochlamys Machatschke, 1970 Spodistes Burmeister, 1847 Spodochlamyini Burmeister, 1855 Spodochlamys Burmeister, 1855 Stalagmosoma Burmeister, 1842 Stebnickiella Skelley, 2007 Stegopterus Burmeister & Schaum, 1840 Stenochelyne Britton, 1990 Stenocorythoderus Wasmann, 1918 Stenocrates Burmeister, 1847 Stenolomaptera Kraatz, 1898 Stenonota Fairmaire, 1889 Stenopegylis Arrow, 1943 Stenopisthes Moser, 1913 Stenoserica Brenske, 1900 Stenosophrops Nomura, 1977 Stenosternus Karsch, 1881 Stenotarsia Burmeister, 1842 Stenotothorax Schmidt, 1914 Stenovalgus Kolbe, 1892 Stephanopholis Brenske, 1896 Stephanorrhina Burmeister, 1842 Stephopseudinca Lucassen, 1933 Stereomera Arrow, 1905 Stethaspis Hope, 1837 Stethodesma Bainbridge, 1840 Stethodesma Hope, 1841 Stethopseudinca Valck Lucassen, 1933 Stichothyrea Kraatz, 1882 Stigmatoplia Dombrow, 2001 Stilbotrochalus Kolbe, 1914 Stiptocnemis Branco, 1989 Stiptopodius Harold, 1871 Stiptotarsus Branco, 1989 Stizopygora Kraatz, 1881 Stomanomala Kolbe, 1897 Storeyus Hasenpusch & Moeseneder, 2010 Straliga Fairmaire, 1901 Strategus Hope, 1837 Strategus Kirby, 1828 Streblopus Lansberge, 1874 Streblopus-Van Lansberge, 1874 Strigoderma Burmeister, 1844 Strigodius Gordon & Skelley, 2007 Stripsipher Gory & Percheron, 1833 Strumadoretus Ohaus, 1912 Stypotrupes Burmeister, 1847 Subrinus Mulsant & Rey, 1869 Sugrames Reitter, 1894 Sukelus Branco, 1992 Sulcophanaeus Olsoufieff, 1924 Sunenaga Ochi, 2003 Suntemnonycha Péringuey, 1904 Supertermitoderus Mencl, 2011 Surutu Martinez, 1955 Sussorca Paulian, 1954 Svobia Mikšic, 1965 Sybacodes Fairmaire, 1896 Sybax Boheman, 1857 Sylvicanthon Halffter & Martinez, 1977 Symmela Erichson, 1835 Symphodon Schmidt, 1914 Synacta Fairmaire, 1899 Synapsis Bates, 1868 Synchilus Britton, 1956 Synclitopa Kolbe, 1897 Synenaria Lacroix, 1993 Syneterogomphu Syngeneschiza Brenske, 1898 Synistovalgus Kolbe, 1897 Syntomopteryx Kraatz, 1900 Syrichthodontus Péringuey, 1901 Syrichthomorphus Péringuey, 1901 Syrichthoschema Janssens, 1942 Syrichtomorphus Systellopus Sharp, 1877 Systellorhina Kraatz, 1895 Systellorrhina Kraatz, 1895 Taeniesthes Kraatz, 1880 Taeniodera Burmeister, 1842 Tafaia Valck Lucassens, 1939 Taiwanoserica Taiwanotrichia Kobayashi, 1990 Talautoclyster Yamaya, 2001 Tamnoserica Brenske, 1899 Tanyana Stebnicka, 2006 Tanyproctoides Petrovitz, 1971 Tanyproctus Faldermann, 1832 Tanyproctus Ménétriés, 1832 Tanzanilepis Lacroix, 2008 Tanzanipholis Lacroix, 2002 Tanzanolus Scholtz & Howden, 1987 Taphraeoserica Brenske, 1900 Taphrocephala Quedenfeldt, 1888 Tapinoschema Thomson, 1880 Tarsocamenta Moser, 1924 Tarsovalgus Miyake, 1993 Taurhina Burmeister, 1842 Taurrhina Taxipertha Baraud, 1991 Tehuacania Endrödi, 1975 Teinogenys Sharp, 1873 Telaugis Burmeister, 1844 Telochilus Krikken, 1975 Telura Erichson, 1842 Teluroides Britton, 1990 Temnoplectron Westwood, 1841 Temnorhynchus Hope, 1837 Temnorrhynchodes Prell, 1934 Temnorrhynchus Hope, 1837 Tephraea Burmeister, 1842 Tephraeoserica Brenske, 1900 Teraserica Brenske, 1897 Terebrogaster Lacroix, 1989 Termitaxis Krikken, 1970 Termitoderus Mateu, 1966 Termitodiellus Nakane, 1961 Termitodius Wasmann, 1894 Termitophilus Britton, 1957 Termitopisthes Wasmann, 1918 Termitotrox Reichensperger, 1915 Termitotrox Wasmann, 1918 Territonia Krikken, 2018 Tesarius Rakovič, 1981 Tesserodon Hope, 1837 Tesserodoniella Vaz-de-Mello & Halffter, 2006 Testaceitrichia Matsumoto, 2021 Tetraaechma Blanchard, 1846 Tetraclipeoides Schmidt, 1914 Tetraechma Agassiz, 1845 Tetraechma Blanchard, 1845 Tetragonorhina Kraatz, 1896 Tetragonorrhina Kraatz, 1896 Tetragus Duarte & Grossi, 2022 Tetraodorhina Blanchard, 1842 Tetrarhabdotis Kraatz, 1883 Tetraserica Ahrens, 2004 Tetrodontus Zhang, 1991 Teuchestes Mulsant, 1842 Thabina Péringuey, 1902 Thamnastopeos Kraatz, 1883 Theodosia Thomson, 1880 Theotimius Huchet, 2000 Theuremaripa Soula, 2006 Thinorycter Semenov & Reichardt, 1925 Thoracoplia Prokofiev, 2015 Thoracotrichia Decelle, 1968 Thronistes Burmeister, 1847 Thrymoserica Brenske, 1897 Thyce LeConte, 1856 Thyregis Blackburn, 1904 Thyreogonia Reitter, 1898 Thyridium Burmeister, 1844 Thyriochlorota Ohaus, 1915 Tiamidema Prokofiev, 2014 Tiarocera Burmeister, 1842 Tiaronthophagus Roggero, Moretto, Barbero & Palestrini, 2019 Tibiodrepanus Krikken, 2009 Tibiotrichius Miyake, 1994 Tibiovalgus Kolbe, 1904 Tibiovalgus Krikken, 1978 Tichocelis Moser, 1908 Tiniocellus Péringuey, 1901 Tipicha Soula, 2002 Tlaocera Péringuey, 1904 Tmesorrhina Westwood, 1841 Tocama Reitter, 1902 Tomarus Erichson, 1847 Tomentoprotaetia Miksic, 1987 Tomogonus Orbigny, 1904 Tonkinilla Prokofiev, 2021 Torynorrhina Arrow, 1907 Tosevskiana Pavicevic, 1985 Toxocnemis Dombrow, 2002 Toxospathius Fairmaire, 1878 Tracheterogomphus Prell, 1912 Trachyserica Brenske, 1899 Tragiscus Klug, 1855 Trapezorrhina Hauser, 1904 Trematodes Faldermann, 1835 Triacmoserica Kolbe, 1914 Triaphodius Tribialus Dupuis & Perrin, 2022 Tribopertha Reitter, 1903 Trichanomala Arrow, 1917 Trichaphodiellus Schmidt, 1914 Trichaphodioides Paulian, 1942 Trichaphodius Schmidt, 1914 Trichaulax Kraatz, 1880 Trichesthes Erichson, 1847 Trichillidium Vaz-de-Mello, 2008 Trichillum Harold, 1868 Trichinopus Waterhouse, 1875 Trichioaparammoecius Minkina, 2018 Trichiodera Burmeister, 1855 Trichiopsammobius Petrovitz, 1963 Trichiorhyssemus Clouët, 1901 Trichiotinus Casey, 1915 Trichius Fabricius, 1775 Trichocanthon Pereira & Martinez, 1960 Trichocelis Moser, 1908 Trichocephala Moser, 1916 Trichogomphus Burmeister, 1847 Tricholepis Blanchard, 1851 Tricholontha Nomura, 1952 Trichomaladera Nomura, 1974 Trichomia Trichoneptunides Legrand, 2001 Trichonotuloides Balthasar, 1945 Trichonotulus Bedel, 1911 Trichonthophagus Zunino, 1979 Trichoparadidactylia Bordat, 2022 Trichoplus Burmeister, 1842 Trichopopillia Ohaus, 1897 Trichoschiza Moser, 1917 Trichoserica Reitter, 1896 Trichostetha Burmeister, 1842 Trichothyrea Kolbe, 1895 Triclirea Schoch, 1896 Trigonochilus Brenske, 1896 Trigonocnemis Kraatz, 1894 Trigonocnemus Kraatz, 1894 Trigonopeltastes Burmeister & Schaum, 1840 Trigonophorinus Pouillaude, 1913 Trigonophorus Hope, 1831 Trigonoscelus (Petrovitz, 1963) Trigonostomus Brenske, 1893 Trinodia Casey, 1915 Trinoxia Brenske, 1894 Triodontella Reitter, 1919 Triodontonyx Moser, 1921 Triodontus Westwood, 1845 Triodontus alticola Paulian, 1976 Triodontus ankarafantsikae Frolov, Montreuil & Akhmetova, 2016 Triodontus bicavatus (Fairmaire, 1905) Triodontus copridoides Paulian, 1976 Triodontus fairmairei Frolov, Montreuil & Akhmetova, 2016 Triodontus hanskii Frolov, 2010 Triodontus hildebrandti (Fairmaire, 1883) Triodontus hova (Fairmaire, 1868) Triodontus inexpectatus Frolov, Montreuil & Akhmetova, 2016 Triodontus itremoi Paulian, 1976 Triodontus lemoulti Frolov, Montreuil & Akhmetova, 2016 Triodontus modestus (Benderitter, 1914) Triodontus nitidulus (Guérin-Méneville, 1829) Triodontus owas Westwood, 1852 Triodontus viettei Frolov, Montreuil & Akhmetova, 2016 Triodonyx Saylor, 1942 Trioplognathus Ohaus, 1904 Trioplus Burmeister, 1847 Trioserica Moser, 1922 Triplognatha Krikken, 1987 Trissodon Burmeister, 1847 Trissodon Prell, 1936 Tristaphodius Balthasar, 1932 Trizogeniates Ohaus, 1917 Trochaloschema Reitter, 1896 Trochaloserica Brenske, 1900 Trochalus Laporte, 1832 Trogodes Boheman, 1857 Tropidonitis Janssens, 1937 Tropinota Mulsant, 1842 Tropiorhynchus Blanchard, 1851 Trymodera Gerstaecker, 1867 Trypoxylus Minck, 1920 Tryssus Erichson, 1847 Tsaratanoplia Lacroix, 1997 Tulbaghia Péringuey, 1904 Turanella Semenov, 1905 Uenoeus Ochi & Kon, 2017 Ulata Saylor, 1945 Uloptera Burmeister, 1842 Ulopterus Burmeister, 1842 Unguiculenaris Dewailly, 1950 Unidentis Josso & Prévost, 2008 Upsa Deschodt, Sole & Scholtz, 2020 Urbania Mikšic, 1963 Uroxys Westwood, 1842 Vadonaria De Wailly, 1950 Vadonidella Ruter, 1973 Valgoides Fairmaire, 1899 Valgus Scriba, 1790 Vansonia Schein, 1956 Vansoniola Schein, 1958 Vanstaronia Kammerer, 2006 Varencya Lacroix, 1993 Vayana Ohaus, 1915 Versicorpus Deschodt, Davis & Scholtz, 2011 Vezoanus Lacroix, 1993 Vietnamoprotaetia Mikšic, 1971 Vieuella Ruter, 1964 Viridimicus Jameson, 1990 Vladimirellus Dellacasa, Dellacasa & Bordat, 2002 Volinus Mulsant & Rey, 1869 Vulcanocanthon Pereira & Martinez, 1960 Vulcanoia Martinez, 1954 Wadaia Itoh, 1993 Walsternoplus Allsopp, Jákl & Rey, 2023 Walterantus Cambefort, 1977 Wambo Allsopp, 1988 Warwickia Smith & Evans, 2005 Watkinsia Britton, 1995 Webbella Britton, 1988 Wernoryctes Dechambre, 1999 Xalpirta Skelley & Cekalovic, 2001 Xanthotrogus Reitter, 1902 Xeloma Kraatz, 1881 Xenaclopus Arrow, 1915 Xenocanthon Martinez, 1952 Xenoceraspis Arrow, 1920 Xenochlorota Soula, 2002 Xenodorus Brême, 1844 Xenogeniates Villatoro & Jameson, 2001 Xenoheptaulacus Hinton, 1934 Xenoloba Bates, 1889 Xenopelidnota Bates, 1904 Xenoproctis Kolbe, 1896 Xenoreoderus Arrow, 1910 Xenoserica Ahrens, 2005 Xenotrochalus Moser, 1917 Xenotrogus Britton, 1978 Xeropsamobeus Saylor, 1937 Xinidium Harold, 1869 Xiphoscelis Burmeister, 1842 Xiphosceloides Holm, 1992 Xylonichus Boisduval, 1835 Xylorectes Xyloryctes Hope, 1837 Xyloscaptes Prell, 1934 Xylotrupes Hope, 1837 Xyridea Britton, 1990 Xyrine Britton, 1987 Xyroa Britton, 1987 Xyrodes Britton, 1987 Yaaxkumukia Morón & Nogueira, 1998 Yanovalgus Nomura, 1952 Yixianscarabaeus Nikolajev, 2015 Youngaphodius Bordat, 1988 Yvescambefortius Ochi & Kon, 1996 Zaburina Saylor, 1945 Zagurya Fuhrmann, 2021 Zebinus Fairmaire, 1894 Zietzia Blackburn, 1894 Zimbabuephylla Lacroix, 1997 Zomba Lacroix, 2006 Zonocopris Arrow, 1932 古埃及人將糞金龜（推糞蟲、屎壳郎）視為生命永恆的象徵，因為其推動糞球有如太陽東昇西落運轉一般，象徵埃及太陽神拉。 許多種類會對經濟植物造成危害。如在北美危害花卉、果樹、農作物的日本豆金龜（Popillia japonica）。 糞金龜能分解動物的糞便，因此在非洲和澳洲的生態上扮演重要的角色。 一種豆金龜 Phyllopertha horticola 犀角金龜 Oryctes nasicornis 金花金龜 Cetonia aurata 糞金龜的一種（南非） 欧洲鳃金龟 Melolontha melolontha 欧洲鳃金龟 Melolontha melolontha 欧洲鳃金龟 Melolontha melolontha 欧洲鳃金龟 Melolontha melolontha 犀角金龜 Oryctes nasicornis 美東白兜蟲 Dynastes tityus 粪金龟 Oxysternon conspicillatum [在维基数据编辑] 《欽定古今圖書集成·博物彙編·禽蟲典·金龜部》，出自陈梦雷《古今圖書集成》 小林準治; 手塚治虫. 朱耀沂（翻譯） , 编. 《手塚治虫 昆蟲圖鑑》. 玉山社. 2004-01-03 [2004]. ISBN 986-781-944-6 （中文）. （繁體中文） 维基共享资源上的相关多媒体资源：金龜子 Vårdal, H. & Forshage, M. 2010. A new genus and species and a revised phylogeny of Stereomerini (Coleoptera, Scarabaeidae, Aphodiinae), with notes on assumedly termitophilic aphodiines. ZooKeys, 34, 55–76. DOI: 10.3897/zookeys.34.264 Maruyama, Munetoshi. 2012. A new genus and species of flightless, microphthalmic Corythoderini (Coleoptera: Scarabaeidae: Aphodiinae) from Cambodia, associated with Macrotermes termites. Zootaxa. (3555): 83-88. Qiao-Zhi Yang (Georgi Geshev) and Wuttipon Pathomwattananurak. 2023. A New Species of the Genus Pseudohomonyx Arrow, 1908 (Coleoptera: Scarabaeidae: Dynastinae) from Malaysia and Thailand, with First Description of P. apoensis Miyake & Yamaya, 1997 male and A Catalogue of the Genus. Zootaxa. 5361(1); 114-124. DOI: 10.11646/zootaxa.5361.1.6 Maruyama M (2010) A new genus and species of myrmecophilous aphodiine beetle (Coleoptera, Scarabaeidae) inhabiting the myrmecophytic epiphyte Platycerium sp. (Polypodiaceae) in the Bornean rainforest canopy. In: Ratcliff e B, Krell F-T (Eds) Current advances in Scarabaeoidea research. ZooKeys. 34: 49–54. doi: 10.3897/zookeys.34.281 Chris A.M. Reid, Natalie A. Tees. 2023. A New, but possibly extinct, Species of Semanopterus Hope, 1847 from Lord Howe Island, in the southwestern Pacific Ocean (Coleoptera: Scarabaeidae: Dynastinae). Zootaxa. 5306(5); 563-570. DOI: 10.11646/zootaxa.5306.5.4 Andrey V. Frolov, Olivier Montreuil and Lilia A. Akhmetova. 2016. Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with A Revision of the Genus Triodontus Westwood. Zootaxa. 4207(1); 1–93. DOI: 10.11646/zootaxa.4207.1.1