Add recognized HTTP headers

Add X-Robots-Tag and Content-Language as HTTP headers which should not have x-amz-meta prefix.
Improve metadata processing (system vs. user-defined).

Let Key.base_fields (boto/s3/key.py) be a list of all system metadata. All
metadata processing shall refer to that list and treat everything included
as system metadata and everything else as user metadata (which gets
x-amz-meta- prefix.)
Fix stupid typo in my previous commit.
Do not percent-encode white space character in S3 metadata.
Encode ' ' as '%20' after all.

Revert earlier commit; encode space as %20 in headers, and modify tests
accordingly.
Merge pull request #2477 from opottone/develop

Add recognized HTTP headers for S3 metadata. Fixes #2477, #2050.