Fixed #23600 - Made default_storage take account of settings changes.

Updated file_storage_changed signal handler to check for changes to

MEDIA_URL,
FILE_UPLOAD_PERMISSIONS, and
FILE_UPLOAD_DIRECTORY_PERMISSIONS

as well as MEDIA_ROOT which was handled in #17744. This means that
overriding any of these settings will now be reflected in
django.files.storage.default_storage.