Mendelian inheritance
It is a mark of his excellence as a scientist that he should have taken the trouble to follow the fate of subsequent generations and so be able to make these observation. It may seem surprising that the clear evidence for non-Mendelian inheritance should have been largely overlooked by the scientific community. It may also be that, as at that time the mechanisms of genetic inheritance in DNA were becoming open to investigation, this example of non-Mendelian inheritance was not of great interest to the scientific community. While non-mendelian inheritance is limited to the cortical structures and to the internal organelles with their own DNAk it does play a major role in some human disorders as well as "handiness" and the left/right orientation of the zygote. Very important factors.