중동호흡기증후군
Rousettus bat coronavirus HKU9 (HKU9-1) is an enveloped, positive-sense, single-stranded RNA mammalian Group 2 "Betacoronavirus" discovered in "Rousettus" bats in China in 2011. This strain of coronavirus is closely related to the EMC/2012 strain found in London which is related to the "Middle East respiratory syndrome-related coronavirus" ("MERS-CoV"). The "MERS-CoV" species is responsible for the 2012 Middle East respiratory syndrome coronavirus outbreak in Saudi Arabia, Jordan, United Arab Emirates, the United Kingdom, France, and Italy.