パスツール研究所
He studied natural sciences at the University of Montpellier, followed by courses in microbiology at the Pasteur Institute in Paris. From 1907 to 1910, he served as préparateur at the laboratory of microbiology courses headed by Amédée Borrel (1867–1936). In 1910, he obtained his doctorate of medicine. Following an internship at the Pasteur Institute in Lille, he was appointed laboratory chief (microbial analysis) at the Pasteur Institute in Algiers. In 1918 he received his doctorate of natural sciences.