血縁選択説
Kin selection based on Hamiliton’s Rule provides an explanation to the population genetics framework and specifies the patterns of local and family genetic makeup that results in the most favorable outcome. The key to this is that the genetic structure spawns nonrandom genotypes which coupled with differential reproduction makes kin selection possible. Inter-nest relatedness was high resulting in few independent haploid genomes which makes possible the development of a social hierarchy.