Gairdner Foundation International Award
In 1998, he won the Albert Lasker Award for Basic Medical Research with Lee Hartwell and Paul Nurse for their pioneering work on cell division. He was elected a Fellow of the Royal Society (FRS) in 1998 and an officer of the Order of Canada in 2003 in recognition of his life's work. in 1992 he was awarded the Gairdner Foundation International Award.