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In fishes, gill lamellae are used to increase the surface area between the surface area in contact with the environment to maximize gas exchange (both to attain oxygen and to expel carbon dioxide) between the water and the blood. In fish gills there are two types of lamellae, primary and secondary. The primary gill lamellae (also called gill filament) extends from the gill arch, and the secondary gill lamellae extends from the primary gill lamellae. Gas exchange primarily occurs at the secondary gill lamellae, where the tissue is notably only one cell layer thick. Furthermore, countercurrent gas exchange at the secondary gill lamellae further maximizes oxygen uptake and carbon dioxide release.