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He studied Biology at the New Asia College of the Chinese University of Hong Kong and was awarded a B.Sc. (3rd Class Honours) and a M.Phil. in 1972 and 1974, respectively. Upon the recommendation of his mentor at the CUHK, he continued his graduate education in the United States and received his Ph.D. from the University of Pittsburgh in 1979. He became Postdoctoral Investigator and Postdoctoral Fellow in 1979 at Oak Ridge National Laboratory, Tennessee, then joined the Department of Genetics of the Hospital for Sick Children in Toronto in 1981.