Mari language
As a professor at Indiana University, Sebeok studied both human and non-human systems of signaling and communication, as well as the philosophy of mind. He was among the founders of biosemiotics, and coined the term "zoosemiotics" in 1963 to describe the development of signals and signs by non-human animal species. He also continued his work as a linguist, publishing several articles and books analyzing aspects of the Mari language (referring to it by the name "Cheremis"). His transdisciplinary work and professional collaborations spanned the fields of anthropology, biology, folklore studies, linguistics, psychology, and semiotics.