중동호흡기증후군
Tylonycteris bat coronavirus HKU4 (Bat-CoV HKU4) is an enveloped, positive-sense single-stranded RNA virus mammalian Group 2 "Betacoronavirus" that has been found to be genetically related to the "Middle East respiratory syndrome-related coronavirus" ("MERS-CoV") that is responsible for the 2012 Middle East respiratory syndrome coronavirus outbreak in Saudi Arabia, Jordan, United Arab Emirates, the United Kingdom, France, and Italy.