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At one time, the fossil species "Rimasuchus lloydi" was thought to be the ancestor of the Nile crocodile, but more recent research has indicated that "Rimasuchus", despite its very large size (about 20–30% bigger than a Nile crocodile with a skull length estimated up to ), is more closely related to the dwarf crocodile ("Osteolaemus tetraspis") among living species. Two other fossil species from Africa retained in the genus "Crocodylus" appear to be closely related to the Nile crocodile: "C. anthropophagus" from Plio-Pleistocene Tanzania and "C. thorbjarnarsoni" from Plio-Pleistocene Kenya. "C. anthropophagus" and "C. thorbjarnarsoni" were both somewhat larger, with projected total lengths up to . As well as being larger, "C. anthropophagus" and "C. thorbjarnarsoni", as well as "Rimasuchus" spp., were all relatively broad-snouted, indicating a specialization at hunting sizeable prey, such as large mammals and freshwater turtles, the latter much larger than any in present-day Africa. Studies have since shown these other African crocodiles to be only more distantly related to the Nile crocodile.