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Matriarchy may also refer to non-human animal species in which females hold higher status and hierarchical positions, such as among spotted hyenas, and bonobos. The social structure of European bison herds has also been described by specialists as a matriarchy – the cows of the group lead it as the entire herd follows them to grazing areas. Though heavier and larger than the females, the older and more powerful males of the European bison usually fulfill the role of satellites that hang around the edges of the herd. Apart from the mating season when they begin to compete with each other, European bison bulls serve a more active role in the herd only once a danger to the group's safety appears.